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Sunday, August 4, 2019

Sierra Nevada (U.S.)

From Wikipedia, the free encyclopedia

Sierra Nevada
Sierra Nevada aerial.jpg
The Sierra's Mills Creek cirque (center) is on the west side of the Sierra Crest, south of Mono Lake (top, blue).
Highest point
PeakMount Whitney
Elevation14,505 ft (4,421 m) 
Coordinates36°34′43″N 118°17′31″W
Dimensions
Length400 mi (640 km) north-south from Fredonyer Pass to Tehachapi Pass
Width65 mi (105 km) 
Area24,370 sq mi (63,100 km2) 
Naming
Etymology1777: Spanish for "snowy mountain range"
Nicknamethe Sierra, the High Sierra, Range of Light (1894, John Muir)
Geography
Sierra Nevada map.png
Position of Sierra Nevada inside California
CountryUnited States
StatesCalifornia and Nevada
Range coordinates37°43′51″N 119°34′22″WCoordinates: 37°43′51″N 119°34′22″W
Geology
Age of rockMesozoic
Type of rockbatholith and igneous

The Sierra Nevada (/siˌɛrə nɪˈvædə, -ˈvɑːdə/, Spanish: [ˈsjera neˈβaða], snowy range) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily in Nevada. The Sierra Nevada is part of the American Cordillera, a chain of mountain ranges that consists of an almost continuous sequence of such ranges that form the western "backbone" of North America, Central America, South America and Antarctica.

The Sierra runs 400 miles (640 km) north-to-south, and is approximately 70 miles (110 km) across east-to-west. Notable Sierra features include Lake Tahoe, the largest alpine lake in North America; Mount Whitney at 14,505 ft (4,421 m), the highest point in the contiguous United States; and Yosemite Valley, sculpted by glaciers from one-hundred-million-year-old granite. The Sierra is home to three national parks, twenty wilderness areas, and two national monuments. These areas include Yosemite, Sequoia, and Kings Canyon National Parks; and Devils Postpile National Monument.

The character of the range is shaped by its geology and ecology. More than one hundred million years ago during the Nevadan orogeny, granite formed deep underground. The range started to uplift four million years ago, and erosion by glaciers exposed the granite and formed the light-colored mountains and cliffs that make up the range. The uplift caused a wide range of elevations and climates in the Sierra Nevada, which are reflected by the presence of five life zones (areas with similar plant and animal communities). Uplift continues due to faulting caused by tectonic forces, creating spectacular fault block escarpments along the eastern edge of the southern Sierra.

The Sierra Nevada has a significant history. The California Gold Rush occurred in the western foothills from 1848 through 1855. Due to inaccessibility, the range was not fully explored until 1912.

Geography

The Sierra Nevada lies in Central and Eastern California, with a very small but historically important spur extending into Nevada. West-to-east, the Sierra Nevada's elevation increases gradually from 1,000 feet (300 m) in the Central Valley to heights of about 14,000 feet (4,300 m) at its crest 50–75 miles (80–121 km) to the east. The east slope forms the steep Sierra Escarpment. Unlike its surroundings, the range receives a substantial amount of snowfall and precipitation due to orographic lift.

Setting

The Sierra Nevada's irregular northern boundary stretches from the Susan River and Fredonyer Pass to the North Fork Feather River. It represents where the granitic bedrock of the Sierra Nevada dives below the southern extent of Cenozoic igneous surface rock from the Cascade Range. It is bounded on the west by California's Central Valley and on the east by the Basin and Range Province. The southern boundary is at Tehachapi Pass.

Physiographically, the Sierra is a section of the Cascade–Sierra Mountains province, which in turn is part of the larger Pacific Mountain System physiographic division. The California Geological Survey states that "the northern Sierra boundary is marked where bedrock disappears under the Cenozoic volcanic cover of the Cascade Range."

Watersheds

The Sierra hosts many waterways, such as the Tuolumne River.
 
The range is drained on its western slope by the Central Valley watershed, which discharges into the Pacific Ocean at San Francisco. The northern third of the western Sierra is part of the Sacramento River watershed (including the Feather, Yuba, and American River tributaries), and the middle third is drained by the San Joaquin River (including the Mokelumne, Stanislaus, Tuolumne, and Merced River tributaries). The southern third of the range is drained by the Kings, Kaweah, Tule, and Kern rivers, which flow into the endorheic basin of Tulare Lake, which rarely overflows into the San Joaquin during wet years. 

The eastern slope watershed of the Sierra is much narrower; its rivers flow out into the endorheic Great Basin of eastern California and western Nevada. From north to south, the Susan River flows into intermittent Honey Lake, the Truckee River flows from Lake Tahoe into Pyramid Lake, the Carson River runs into Carson Sink, the Walker River into Walker Lake; Rush, Lee Vining and Mill Creeks flow into Mono Lake; and the Owens River into dry Owens Lake. Although none of the eastern rivers reach the sea, many of the streams from Mono Lake southwards are diverted into the Los Angeles Aqueduct which provides water to Southern California.

Elevation

Mount Whitney, the highest peak in the range and the contiguous United States
 
The height of the mountains in the Sierra Nevada increases gradually from north to south. Between Fredonyer Pass and Lake Tahoe, the peaks range from 5,000 feet (1,500 m) to more than 9,000 feet (2,700 m). The crest near Lake Tahoe is roughly 9,000 feet (2,700 m) high, with several peaks approaching the height of Freel Peak (10,881 ft or 3,317 m). Farther south, the highest peak in Yosemite National Park is Mount Lyell (13,120 ft or 3,999 m). The Sierra rises to almost 14,000 feet (4,300 m) with Mount Humphreys near Bishop, California. Finally, near Lone Pine, Mount Whitney is at 14,505 feet (4,421 m), the highest point in the contiguous United States

South of Mount Whitney, the elevation of the range quickly dwindles. The crest elevation is almost 10,000 feet (3,000 m) near Lake Isabella, but south of the lake, the peaks reach to only a modest 8,000 feet (2,400 m).

Notable features

There are several notable geographical features in the Sierra Nevada:

Communities

Protected areas

View of Sequoia National Park from Moro Rock
 
Much of the Sierra Nevada consists of federal lands and is either protected from development or strictly managed. The three National Parks (Yosemite, Kings Canyon, Sequoia), two national monuments (Devils Postpile, Giant Sequoia), and 26 wilderness areas lie within the Sierra. These areas protect 15.4% of the Sierra's 63,118 km2 (24,370 sq mi) from logging and grazing.

The United States Forest Service and the Bureau of Land Management currently control 52% of the land in the Sierra Nevada. Logging and grazing are generally allowed on land controlled by these agencies, under federal regulations that balance recreation and development on the land. 

The California Bighorn Sheep Zoological Area near Mount Williamson in the southern Sierra was established to protect the endangered Sierra Nevada bighorn sheep. Starting in 1981, hikers were unable to enter the Area from May 15 through December 15, in order to protect the sheep. As of 2010, the restriction has been lifted and access to the Area is open for the whole year.

Geologic history

Sevehah Cliff, near Convict Lake, shows severely deformed Devonian rock
 
Yosemite Valley in Yosemite National Park was carved by glaciers
The earliest rocks in the Sierra Nevada are metamorphic roof pendants of Paleozoic age, the oldest being metasedimentary rocks from the Cambrian in the Mount Morrison region. These dark-colored hornfels, slates, marbles, and schists are found in the western foothills (notably around Coarsegold, west of the Tehachapi Pass) and east of the Sierra Crest. The earliest granite of the Sierra started to form in the Triassic period. This granite is mostly found east of the crest and north of 37.2°N. In the Triassic and into the Jurassic, an island arc collided with the west coast of North America and raised a chain of volcanoes, in an event called the Nevadan orogeny. Nearly all subaerial Sierran Arc volcanoes have since disappeared; their remains were redeposited during the Great Valley Sequence and the subsequent Cenozoic filling of the Great Valley, which is the source of much of the sedimentary rock in California.

In the Cretaceous, a subduction zone formed at the edge of the continent. This means that an oceanic plate started to dive beneath the North American plate. Magma formed through the subduction of the ancient Farallon Plate rose in plumes (plutons) deep underground, their combined mass forming what is called the Sierra Nevada batholith. These plutons formed at various times, from 115 Ma to 87 Ma. The earlier plutons formed in the western half of the Sierra, while the later plutons formed in the eastern half of the Sierra. By 66 Ma, the proto-Sierra Nevada had been worn down to a range of rolling low mountains, a few thousand feet high.

Twenty million years ago, crustal extension associated with the Basin and Range Province caused extensive volcanism in the Sierra. About 10 Ma, the Sierra Nevada started to form when a block of crust between the Coast Range and the Basin and Range Province started to tilt to the west as heat from the Basin and Range extension thinned the eastern part of the block, making it more buoyant than the western portion of the block. Rivers started cutting deep canyons on both sides of the range. Lava filled some of these canyons, which have subsequently eroded leaving table mountains that follow the old river channels.

About 2.5 Ma, the Earth's climate cooled, and ice ages started. Glaciers carved out characteristic U-shaped canyons throughout the Sierra. The combination of river and glacier erosion exposed the uppermost portions of the plutons emplaced millions of years before, leaving only a remnant of metamorphic rock on top of some Sierra peaks. 

Uplift of the Sierra Nevada continues today, especially along its eastern side. This uplift causes large earthquakes, such as the Lone Pine earthquake of 1872.

Sierra Escarpment viewed from the east. In the foreground is Tinemaha Reservoir in the Owens Valley.

Climate and meteorology

Red Slate Mountain (elevation 13,156 ft or 4,010 m) is still covered with snow in June.
 
The climate of the Sierra Nevada is influenced by the Mediterranean climate of California. During the fall, winter and spring, precipitation in the Sierra ranges from 20 to 80 in (510 to 2,030 mm) where it occurs mostly as snow above 6,000 ft (1,800 m). Precipitation is highest on the central and northern portions of the western slope between 5,000 and 8,000 feet (1,500 and 2,400 m) elevation, due to orographic lift. Above 8,000 feet (2,400 m), precipitation diminishes on the western slope up to the crest, since most of the precipitation has been wrung out at lower elevations. Most parts of the range east of the crest are in a rain shadow, and receive less than 25 inches of precipitation per year. While most summer days are dry, afternoon thunderstorms sometimes occur, particularly during the North American Monsoon in mid and late summer. Some of these summer thunderstorms drop over an inch of rain in a short period, and the lightning can start fires. Summer high temperatures average 42–90 °F (6–32 °C). Winters are comparatively mild, and the temperature is usually only just low enough to sustain a heavy snowpack. For example, Tuolumne Meadows, at 8,600 feet (2,600 m) elevation, has winter daily highs about 40 °F (4 °C) with daily lows about 10 °F (−12 °C). The growing season lasts 20 to 230 days, strongly dependent on elevation. The highest elevations of the Sierra have an alpine climate

The Sierra Nevada snowpack is the major source of water and a significant source of electric power generation in California. Many reservoirs were constructed in the canyons of the Sierra throughout the 20th century, Several major aqueducts serving both agriculture and urban areas distribute Sierra water throughout the state. However, the Sierra casts a rain shadow, which greatly affects the climate and ecology of the central Great Basin. This rain shadow is largely responsible for Nevada being the driest state in the United States.

Precipitation varies substantially from year to year. It is not uncommon for some years to receive precipitation totals far above or below normal.

The height of the range and the steepness of the Sierra Escarpment, particularly at the southern end of the range, produces a wind phenomenon known as the "Sierra Rotor". This is a horizontal rotation of the atmosphere just east of the crest of the Sierra, set in motion as an effect of strong westerly winds.

Because of the large number of airplanes that have crashed in the Sierra Nevada, primarily due to the complex weather and atmospheric conditions such as downdrafts and microbursts caused by geography there, a portion of the area, a triangle whose vertices are Reno, Nevada; Fresno, California; and Las Vegas, Nevada, has been dubbed the "Nevada Triangle", in reference to the Bermuda Triangle. Some counts put the number of crashes in the triangle at 2,000, including millionaire and record-breaking flyer Steve Fossett. Hypotheses that the crashes are related in some way to the United States Air Force's Area 51, or to the activities of extra-terrestrial aliens, have no evidence to support them.

Ecology

Tuolumne Meadows is an example of a subalpine meadow in the Sierra.
 
The Sierra Nevada is divided into a number of biotic zones, each of which is defined by its climate and supports a number of interdependent species. Life in the higher elevation zones adapted to colder weather, and to most of the precipitation falling as snow. The rain shadow of the Sierra causes the eastern slope to be warmer and drier: each life zone is higher in the east. A list of biotic zones, and corresponding elevations, is presented below:

History

John Frémont was an early American explorer of the Sierra

Native Americans

Archaeological excavations placed Martis people of Paleo-Indians in northcentral Sierra Nevada during the period of 3,000 BCE to 500 CE. The earliest identified sustaining indigenous people in the Sierra Nevada were the Northern Paiute tribes on the east side, with the Mono tribe and Sierra Miwok tribe on the western side, and the Kawaiisu and Tübatulabal tribes in the southern Sierra. Today, some historic intertribal trade route trails over mountain passes are known artifact locations, such as Duck Pass with its obsidian arrowheads. The California and Sierra Native American tribes were predominantly peaceful, with occasional territorial disputes between the Paiute and Sierra Miwok tribes in the mountains. Washo and Maidu were also in this area prior to the era of European exploration and displacement.

Etymology

Kearsarge Lakes Basin is named after the USS Kearsarge
 
Used in 1542 by Juan Rodríguez Cabrillo to describe a Pacific Coast Range (Santa Cruz Mountains), the term "sierra nevada" was a general identification of less familiar ranges toward the interior. In 1776, Pedro Font's map applied the name to the range currently known as the Sierra Nevada.

The literal translation is "snowy mountains", from sierra "a range of hills", 1610s, from Spanish sierra "jagged mountain range", lit. "saw", from Latin serra "a saw"; and from fem. of Spanish nevado "snowy".

Initial European-American exploration

American exploration of the mountain range started in 1827. Although prior to the 1820s there were Spanish missions, pueblos (towns), presidios (forts), and ranchos along the coast of California, no Spanish explorers visited the Sierra Nevada. The first Americans to visit the mountains were amongst a group led by fur trapper Jedediah Smith, crossing north of the Yosemite area in May 1827, at Ebbetts Pass.

In 1833, a subgroup of the Bonneville Expedition led by Joseph Reddeford Walker was sent westward to find an overland route to California. Eventually the party discovered a route along the Humboldt River across present-day Nevada, ascending the Sierra Nevada, starting near present-day Bridgeport and descending between the Tuolumne and Merced River drainage. The group may have been the first non-indigenous people to see Yosemite Valley. The Walker Party probably visited either the Tuolumne or Merced Groves of Giant Sequoia, becoming the first non-indigenous people to see the giant trees, but journals relating to the Walker party were destroyed in 1839, in a print shop fire in Philadelphia.

In the winter of 1844, Lt. John C. Frémont, accompanied by Kit Carson, was the first American to see Lake Tahoe. The Frémont party camped at 8,050 ft (2,450 m).

Gold rush

Map of gold fields in the Sierra

The California Gold Rush began at Sutter's Mill, near Coloma, in the western foothills of the Sierra. On January 24, 1848, James W. Marshall, a foreman working for Sacramento pioneer John Sutter, found shiny metal in the tailrace of a lumber mill Marshall was building for Sutter on the American River. Rumors soon started to spread and were confirmed in March 1848 by San Francisco newspaper publisher and merchant Samuel Brannan. Brannan strode through the streets of San Francisco, holding aloft a vial of gold, shouting "Gold! Gold! Gold from the American River!"

On August 19, 1848, the New York Herald was the first major newspaper on the East Coast to report the discovery of gold. On December 5, 1848, President James Polk confirmed the discovery of gold in an address to Congress. Soon, waves of immigrants from around the world, later called the "forty-niners", invaded the Gold Country of California or "Mother Lode". Miners lived in tents, wood shanties, or deck cabins removed from abandoned ships. Wherever gold was discovered, hundreds of miners would collaborate to put up a camp and stake their claims. 

Because the gold in the California gravel beds was so richly concentrated, the early forty-niners simply panned for gold in California's rivers and streams. However, panning cannot take place on a large scale, and miners and groups of miners graduated to more complex placer mining. Groups of prospectors would divert the water from an entire river into a sluice alongside the river, and then dig for gold in the newly exposed river bottom.

By 1853, most of the easily accessible gold had been collected, and attention turned to extracting gold from more difficult locations. Hydraulic mining was used on ancient gold-bearing gravel beds on hillsides and bluffs in the gold fields. In hydraulic mining, a high-pressure hose directed a powerful stream or jet of water at gold-bearing gravel beds. It is estimated that by the mid-1880s, 11 million ounces (340 t) of gold (worth approximately US$15 billion at December 2010 prices) had been recovered by "hydraulicking". A byproduct of these extraction methods was that large amounts of gravel, silt, heavy metals, and other pollutants went into streams and rivers. As of 1999, many areas still bear the scars of hydraulic mining, since the resulting exposed earth and downstream gravel deposits do not support plant life.

It is estimated that by 1855, at least 300,000 gold-seekers, merchants, and other immigrants had arrived in California from around the world. The huge numbers of newcomers brought by the Gold Rush drove Native Americans out of their traditional hunting, fishing and food-gathering areas. To protect their homes and livelihood, some Native Americans responded by attacking the miners, provoking counter-attacks on native villages. The Native Americans, out-gunned, were often slaughtered.

The exploration team for the California Geological Survey, 1864

Thorough exploration

The Gold Rush populated the western foothills of the Sierra Nevada, but even by 1860, most of the Sierra was unexplored. The state legislature authorized the California Geological Survey to officially explore the Sierra (and survey the rest of the state). Josiah Whitney was appointed to head the survey. Men of the survey, including William H. Brewer, Charles F. Hoffmann and Clarence King, explored the backcountry of what would become Yosemite National Park in 1863. In 1864, they explored the area around Kings Canyon. In 1869, John Muir started his wanderings in the Sierra Nevada range, and in 1871, King was the first to climb Mount Langley, mistakenly believing he had summited Mount Whitney, the highest peak in the range. In 1873, Mount Whitney was climbed for the first time by 3 men from Lone Pine, CA on a fishing trip. From 1892–7 Theodore Solomons made the first attempt to map a route along the crest of the Sierra.

Other people finished exploring and mapping the Sierra. Bolton Coit Brown explored the Kings River watershed in 1895–1899. Joseph N. LeConte mapped the area around Yosemite National Park and what would become Kings Canyon National Park. James S. Hutchinson, a noted mountaineer, climbed the Palisades (1904) and Mount Humphreys (1905). By 1912, the USGS published a set of maps of the Sierra Nevada, and the era of exploration was over.

Conservation

The General Sherman Tree, a Giant Sequoia in Sequoia National Park, is the world's largest tree by volume.
 
The tourism potential of the Sierra Nevada was recognized early in the European history of the range. Yosemite Valley was first protected by the federal government in 1864. The Valley and Mariposa Grove were ceded to California in 1866 and turned into a state park. John Muir perceived overgrazing by sheep and logging of Giant Sequoia to be a problem in the Sierra. Muir successfully lobbied for the protection of the rest of Yosemite National Park: Congress created an Act to protect the park in 1890. The Valley and Mariposa Grove were added to the Park in 1906. In the same year, Sequoia National Park was formed to protect the Giant Sequoia: all logging of the Sequoia ceased at that time.
In 1903, the city of San Francisco proposed building a hydroelectric dam to flood Hetch Hetchy Valley. The city and the Sierra Club argued over the dam for 10 years, until the U.S. Congress passed the Raker Act in 1913 and allowed dam building to proceed. O'Shaughnessy Dam was completed in 1923.

Between 1912 and 1918, Congress debated three times to protect Lake Tahoe in a national park. None of these efforts succeeded, and after World War II, towns such as South Lake Tahoe grew around the shores of the lake. By 1980, the permanent population of the Lake Tahoe area grew to 50,000, while the summer population grew to 90,000. The development around Lake Tahoe affected the clarity of the lake water. In order to preserve the lake's clarity, construction in the Tahoe basin is currently regulated by the Tahoe Regional Planning Agency.

As the 20th century progressed, more of the Sierra became available for recreation; other forms of economic activity decreased. The John Muir Trail, a trail that followed the Sierra crest from Yosemite Valley to Mount Whitney, was funded in 1915 and finished in 1938. Kings Canyon National Park was formed in 1940 to protect the deep canyon of the Kings River

By 1964, the Wilderness Act protected portions of the Sierra as primitive areas where humans are simply temporary visitors. Gradually, 20 wilderness areas were established to protect scenic backcountry of the Sierra. These wilderness areas include the John Muir Wilderness (protecting the eastern slope of the Sierra and the area between Yosemite and Kings Canyon Parks), and wilderness within each of the National Parks. Because of the Wilderness Act and the rocky terrain in the area, plans to construct two trans-Sierra highways across this portion of the Sierra Escarpment, State Route 168 and State Route 190, were abandoned; the two highways each remain split as discontiguous segments on either side of the Sierra. 

The Sierra Nevada still faces a number of issues that threaten its conservation. Logging occurs on both private and public lands, including controversial clearcut methods and thinning logging on private and public lands. Grazing occurs on private lands as well as on National Forest lands, which include Wilderness areas. Overgrazing can alter hydrologic processes and vegetation composition, remove vegetation that serves as food and habitat for native species, and contribute to sedimentation and pollution in waterways. A recent increase in large wildfires like the Rim Fire in Yosemite National Park and the Stanislaus National Forest and the King Fire on the Eldorado National Forest, has prompted concerns. A 2015 study indicated that the increase in fire risk in California may be attributable to human-induced climate change. A study looking back over 8,000 years found that warmer climate periods experienced severe droughts and more stand-replacing fires and concluded that as climate is such a powerful influence on wildfires, trying to recreate presettlement forest structure is likely impossible in a warmer future.

Saturday, August 3, 2019

Recent African origin of modern humans

From Wikipedia, the free encyclopedia

Successive dispersals of      Homo erectus (yellow),      Homo neanderthalensis (ochre) and      Homo sapiens (red).
 
Expansion of early modern humans from Africa through the Near East
 
Map of the migration of modern humans out of Africa, based on mitochondrial DNA. Colored rings indicate thousand years before present.

In paleoanthropology, the recent African origin of modern humans, also called the "Out of Africa" theory (OOA), recent single-origin hypothesis (RSOH), replacement hypothesis, or recent African origin model (RAO), is the dominant model of the geographic origin and early migration of anatomically modern humans (Homo sapiens). It follows the early expansions of hominins out of Africa, accomplished by Homo erectus and then Homo neanderthalensis

The model proposes a "single origin" of Homo sapiens in the taxonomic sense, precluding parallel evolution of traits considered anatomically modern in other regions, but not precluding multiple admixture between H. sapiens and archaic humans in Europe and Asia. H. sapiens most likely developed in the Horn of Africa between 300,000 and 200,000 years ago. The "recent African origin" model proposes that all modern non-African populations are substantially descended from populations of H. sapiens that left Africa after that time. 

There were at least several "out-of-Africa" dispersals of modern humans, possibly beginning as early as 270,000 years ago, including 215,000 years ago to at least Greece, and certainly via northern Africa about 130,000 to 115,000 years ago. These early waves appear to have mostly died out or retreated by 80,000 years ago.

The most significant "recent" wave took place about 60,000–70,000 years ago, via the so-called "Southern Route", spreading rapidly along the coast of Asia and reaching Australia by around 65,000–50,000 years ago, while Europe was populated by an early offshoot which settled the Near East and Europe less than 55,000 years ago.

In the 2010s, studies in population genetics have uncovered evidence of interbreeding between H. sapiens and archaic humans in Eurasia and Oceania but not in Africa, which means that all non-African modern population groups, while mostly derived from early H. sapiens, are to a lesser extent also descended from regional variants of archaic humans.

Proposed waves

Layer sequence at Ksar Akil in the Levantine corridor, and discovery of two fossils of Homo sapiens, dated to 40,800 to 39,200 years BP for "Egbert",and 42,400–41,700 BP for "Ethelruda"
 
"Recent African origin," or Out of Africa II, refers to the migration of anatomically modern humans (Homo sapiens) out of Africa after their emergence at c. 300,000 to 200,000 years ago, in contrast to "Out of Africa I", which refers to the migration of archaic humans from Africa to Eurasia roughly 1.8 to 0.5 million years ago. 

Since the beginning of the 21st century, the picture of "recent single-origin" migrations has become significantly more complex, not only due to the discovery of modern-archaic admixture but also due to the increasing evidence that the "recent out-of-Africa" migration took place in a number of waves spread over a long time period. As of 2010, there were two main accepted dispersal routes for the out-of-Africa migration of early anatomically modern humans: via the "Northern Route" (via Nile Valley and Sinai) and the "Southern Route" via the Bab al Mandab strait.
  • Posth et al. (2017) suggest that early Homo sapiens, or "another species in Africa closely related to us," might have first migrated out of Africa around 270,000 years ago.
  • Finds at Misliya cave, which include a partial jawbone with eight teeth, have been dated to around 185,000 years ago. Layers dating from between 250,000 and 140,000 years ago in the same cave contained tools of the Levallois type which could put the date of the first migration even earlier if the tools can be associated with the modern human jawbone finds.
  • An Eastward Dispersal from Northeast Africa to Arabia 150,000–130,000  years ago based on the finds at Jebel Faya dated to 127,000 years ago (discovered in 2011). Possibly related to this wave are the finds from Zhirendong cave, Southern China, dated to more than 100,000 years ago. Other evidence of modern human presence in China has been dated to 80,000 years ago.
  • The most significant dispersal took place around 60–70,000 years ago via the so-called Southern Route, either before or after the Toba event, which happened between 69,000 and 77,000 years ago. This dispersal followed the southern coastline of Asia, and reached Australia around 65,000-50,000 years ago. Western Asia was "re-occupied" by a different derivation from this wave around 50,000 years ago, and Europe was populated from Western Asia beginning around 43,000 years ago.
  • Wells (2003) describes an additional wave of migration after the southern coastal route, namely a northern migration into Europe at circa 45,000 years ago. However, this possibility is ruled out by Macaulay et al. (2005) and Posth et al. (2016), who argue for a single coastal dispersal, with an early offshoot into Europe.

Northern Route dispersal

Anatomically Modern Humans known archaeological remains in Europe and Africa, directly dated, calibrated carbon dates as of 2013.
 
Beginning 135,000 years ago, tropical Africa experienced megadroughts which drove humans from the land and towards the sea shores, and forced them to cross over to other continents.

Modern humans crossed the Straits of Bab el Mandab in the southern Red Sea, and moved along the green coastlines around Arabia, and thence to the rest of Eurasia. Fossils of early Homo sapiens were found in Qafzeh cave in Israel and have been dated 80,000 to 100,000 years ago. These humans seem to have either become extinct or retreated back to Africa 70,000 to 80,000 years ago, possibly replaced by southbound Neanderthals escaping the colder regions of ice-age Europe. Hua Liu et al. analyzed autosomal microsatellite markers dating to about 56,000 years ago. They interpret the paleontological fossil as an isolated early offshoot that retracted back to Africa.

The discovery of stone tools in the United Arab Emirates in 2011 at the Faya-1 site in Mleiha, Sharjah, indicated the presence of modern humans at least 125,000 years ago, leading to a resurgence of the "long-neglected" North African route.

In Oman, a site was discovered by Bien Joven in 2011 containing more than 100 surface scatters of stone tools belonging to the late Nubian Complex, known previously only from archaeological excavations in the Sudan. Two optically stimulated luminescence age estimates placed the Arabian Nubian Complex at approximately 106,000 years old. This provides evidence for a distinct stone age technocomplex in southern Arabia, around the earlier part of the Marine Isotope Stage 5.

According to Kuhlwilm and his co-authors, Neanderthals contributed genetically to modern humans then living outside of Africa around 100,000 years ago: humans which had already split off from other modern humans around 200,000 years ago, and this early wave of modern humans outside Africa also contributed genetically to the Altai Neanderthals. They found that "the ancestors of Neanderthals from the Altai Mountains and early modern humans met and interbred, possibly in the Near East, many thousands of years earlier than previously thought". According to co-author Ilan Gronau, "This actually complements archaeological evidence of the presence of early modern humans out of Africa around and before 100,000 years ago by providing the first genetic evidence of such populations." Similar genetic admixture events have been noted in other regions as well.

In China, the Liujiang man (Chinese: 柳江人) is among the earliest modern humans found in East Asia. The date most commonly attributed to the remains is 67,000 years ago. High rates of variability yielded by various dating techniques carried out by different researchers place the most widely accepted range of dates with 67,000 BP as a minimum, but do not rule out dates as old as 159,000 BP. Liu, Martinón-Torres et al. (2015) claim that modern human teeth have been found in China dating to at least 80,000 years ago.

Southern Route dispersal

Coastal route

Red Sea crossing
 
By some 50-70,000 years ago, a subset of the bearers of mitochondrial haplogroup L3 migrated from East Africa into the Near East. It has been estimated that from a population of 2,000 to 5,000 individuals in Africa, only a small group, possibly as few as 150 to 1,000 people, crossed the Red Sea. The group that crossed the Red Sea travelled along the coastal route around Arabia and Persia to India, which appears to have been the first major settling point. Wells (2003) argued for the route along the southern coastline of Asia, across about 250 kilometres (155 mi), reaching Australia by around 50,000 years ago. 

Today at the Bab-el-Mandeb straits, the Red Sea is about 20 kilometres (12 mi) wide, but 50,000 years ago sea levels were 70 m (230 ft) lower (owing to glaciation) and the water was much narrower. Though the straits were never completely closed, they were narrow enough to have enabled crossing using simple rafts, and there may have been islands in between. Shell middens 125,000 years old have been found in Eritrea, indicating the diet of early humans included seafood obtained by beachcombing.

The dating of the Southern Dispersal is a matter of dispute. It may have happened either pre- or post-Toba, a catastrophic volcanic eruption that took place between 69,000 and 77,000 years ago at the site of present-day Lake Toba. Stone tools discovered below the layers of ash disposed in India may point to a pre-Toba dispersal but the source of the tools is disputed. An indication for post-Toba is haplo-group L3, that originated before the dispersal of humans out of Africa and can be dated to 60,000–70,000 years ago, "suggesting that humanity left Africa a few thousand years after Toba". New research showing slower than expected genetic mutations in human DNA was published in 2012, indicating a revised dating for the migration to between 90,000 and 130,000 years ago.

Western Asia

A fossil of a modern human dated to 54,700 years ago was found in Manot Cave in Israel, named Manot 1, though the dating was questioned by Groucutt et al. (2015).

South-Asia and Australia

It is thought that Australia was inhabited around 65,000–50,000 years ago. As of 2017, the earliest evidence of humans in Australia is at least 65,000 years old, while McChesney stated that
...genetic evidence suggests that a small band with the marker M168 migrated out of Africa along the coasts of the Arabian Peninsula and India, through Indonesia, and reached Australia very early, between 60,000 and 50,000 years ago. This very early migration into Australia is also supported by Rasmussen et al. (2011).
Fossils from Lake Mungo, Australia, have been dated to about 42,000 years ago. Other fossils from a site called Madjedbebe have been dated to at least 65,000 years ago.

East Asia

Tianyuan man from China has a probable date range between 38,000 and 42,000 years ago, while Liujiang man from the same region has a probable date range between 67,000 and 159,000 years ago. According to 2013 DNA tests, Tianyuan man is related "to many present-day Asians and Native Americans". Tianyuan is similar in morphology to Minatogawa Man, modern humans dated between 17,000 and 19,000 years ago and found on Okinawa Island, Japan.

Europe

According to Macaulay et al. (2005), an early offshoot from the southern dispersal with haplogroup N followed the Nile from East Africa, heading northwards and crossing into Asia through the Sinai. This group then branched, some moving into Europe and others heading east into Asia. This hypothesis is supported by the relatively late date of the arrival of modern humans in Europe as well as by archaeological and DNA evidence. Based on an analysis of 55 human mitochondrial genomes (mtDNAs) of hunter-gatherers, Posth et al. (2016) argue for a "rapid single dispersal of all non-Africans less than 55,000 years ago."

Genetic reconstruction

Mitochondrial haplogroups

Within Africa

Map of early diversification of modern humans according to mitochondrial population genetics.
 
The first lineage to branch off from Mitochondrial Eve was L0. This haplogroup is found in high proportions among the San of Southern Africa and the Sandawe of East Africa. It is also found among the Mbuti people. These groups branched off early in human history and have remained relatively genetically isolated since then. Haplogroups L1, L2 and L3 are descendants of L1–L6, and are largely confined to Africa. The macro haplogroups M and N, which are the lineages of the rest of the world outside Africa, descend from L3. L3 is about 84,000 years old, while haplogroups M and N are about 63,000 years old. The relationship between such gene trees and demographic history is still debated when applied to dispersals.

Of all the lineages present in Africa, only the female descendants of one lineage, mtDNA haplogroup L3, are found outside Africa. If there had been several migrations, one would expect descendants of more than one lineage to be found. L3's female descendants, the M and N haplogroup lineages, are found in very low frequencies in Africa (although haplogroup M1 populations are very ancient and diversified in North and North-east Africa) and appear to be more recent arrivals. A possible explanation is that these mutations occurred in East Africa shortly before the exodus and became the dominant haplogroups thereafter by means of the founder effect. Alternatively, the mutations may have arisen shortly afterwards.

Southern Route and haplogroups M and N

Results from mtDNA collected from aboriginal Malaysians called Orang Asli indicate that the hapologroups M and N share characteristics with original African groups from approximately 85,000 years ago, and share characteristics with sub-haplogroups found in coastal south-east Asian regions, such as Australasia, the Indian subcontinent and throughout continental Asia, which had dispersed and separated from their African progenitor approximately 65,000 years ago. This southern coastal dispersal would have occurred before the dispersal through the Levant approximately 45,000 years ago. This hypothesis attempts to explain why haplogroup N is predominant in Europe and why haplogroup M is absent in Europe. Evidence of the coastal migration is thought to have been destroyed by the rise in sea levels during the Holocene epoch. Alternatively, a small European founder population that had expressed haplogroup M and N at first, could have lost haplogroup M through random genetic drift resulting from a bottleneck (i.e. a founder effect).

The group that crossed the Red Sea travelled along the coastal route around Arabia and Persia until reaching India. Haplogroup M is found in high frequencies along the southern coastal regions of Pakistan and India and it has the greatest diversity in India, indicating that it is here where the mutation may have occurred. Sixty percent of the Indian population belong to Haplogroup M. The indigenous people of the Andaman Islands also belong to the M lineage. The Andamanese are thought to be offshoots of some of the earliest inhabitants in Asia because of their long isolation from the mainland. They are evidence of the coastal route of early settlers that extends from India to Thailand and Indonesia all the way to Papua New Guinea. Since M is found in high frequencies in highlanders from New Guinea and the Andamanese and New Guineans have dark skin and Afro-textured hair, some scientists think they are all part of the same wave of migrants who departed across the Red Sea ~60,000 years ago in the Great Coastal Migration. The proportion of haplogroup M increases eastwards from Arabia to India; in eastern India, M outnumbers N by a ratio of 3:1. Crossing into Southeast Asia, haplogroup N (mostly in the form of derivatives of its R subclade) reappears as the predominant lineage. M is predominant in East Asia, but amongst Indigenous Australians, N is the more common lineage. This haphazard distribution of Haplogroup N from Europe to Australia can be explained by founder effects and population bottlenecks.

Autosomal DNA

A 2002 study of African, European and Asian populations, found greater genetic diversity among Africans than among Eurasians, and that genetic diversity among Eurasians is largely a subset of that among Africans, supporting the out of Africa model. A large study by Coop et al. (2009) found evidence for natural selection in autosomal DNA outside of Africa. The study distinguishes non-African sweeps (notably KITLG variants associated with skin color), West-Eurasian sweeps (SLC24A5) and East-Asian sweeps (MC1R, relevant to skin color). Based on this evidence, the study concluded that human populations encountered novel selective pressures as they expanded out of Africa. MC1R and its relation to skin color had already been discussed by Liu, Harding et al. (2000), p. 135. According to this study, Papua New Guineans continued to be exposed to selection for dark skin color so that, although these groups are distinct from Africans in other places, the allele for dark skin color shared by contemporary Africans, Andamanese and New Guineans is an archaism. Endicott et al. (2003) suggest convergent evolution. A 2014 study by Gurdasani et al. indicate that higher genetic diversity in Africa was caused by relatively recent Eurasian migrations into Africa.

Pathogen DNA

Another promising route towards reconstructing human genetic genealogy is via the JC virus (JCV), a type of human polyomavirus which is carried by 70–90 percent of humans and which is usually transmitted vertically, from parents to offspring, suggesting codivergence with human populations. For this reason, JCV has been used as a genetic marker for human evolution and migration. This method does not appear to be reliable for the migration out of Africa, in contrast to human genetics, JCV strains associated with African populations are not basal. From this Shackelton et al. (2006) conclude that either a basal African strain of JCV has become extinct or that the original infection with JCV post-dates the migration from Africa.

Admixture of archaic and modern humans

Evidence for archaic human species (descended from Homo heidelbergensis) having interbred with modern humans outside of Africa, was discovered in the 2010s. This concerns primarily Neanderthal admixture in all modern populations except for Sub-Saharan Africans but evidence has also been presented for Denisova hominin admixture in Australasia (i.e. in Melanesians, Aboriginal Australians and some Negritos).

The rate of admixture of Neanderthal admixture to European and Asian populations as of 2017 has been estimated at between about 2–3%.

Archaic admixture in some Sub-Saharan African populations hunter-gatherer groups (Biaka Pygmies and San), derived from archaic hominins that broke away from the modern human lineage around 700,000 years, was discovered in 2011. The rate of admixture was estimated at around 2%. Admixture from archaic hominins of still earlier divergence times, estimated at 1.2 to 1.3 million years ago, was found in Pygmies, Hadza and five Sandawe in 2012. From an analysis of Mucin 7, a highly divergent haplotype that has an estimated coalescence time with other variants around 4.5 million years BP and is specific to African populations is inferred to have been derived from interbreeding between African modern and archaic humans.

Stone tools

In addition to genetic analysis, Petraglia et al. also examines the small stone tools (microlithic materials) from the Indian subcontinent and explains the expansion of population based on the reconstruction of paleoenvironment. He proposed that the stone tools could be dated to 35 ka in South Asia, and the new technology might be influenced by environmental change and population pressure.

History of the theory

Classical paleoanthropology

The frontispiece to Huxley's Evidence as to Man's Place in Nature (1863): the image compares the skeleton of a human to other apes.
 
The cladistic relationship of humans with the African apes was suggested by Charles Darwin after studying the behaviour of African apes, one of which was displayed at the London Zoo. The anatomist Thomas Huxley had also supported the hypothesis and suggested that African apes have a close evolutionary relationship with humans. These views were opposed by the German biologist Ernst Haeckel, who was a proponent of the Out of Asia theory. Haeckel argued that humans were more closely related to the primates of South-east Asia and rejected Darwin's African hypothesis.

In the Descent of Man, Darwin speculated that humans had descended from apes, which still had small brains but walked upright, freeing their hands for uses which favoured intelligence; he thought such apes were African:
In each great region of the world the living mammals are closely related to the extinct species of the same region. It is, therefore, probable that Africa was formerly inhabited by extinct apes closely allied to the gorilla and chimpanzee; and as these two species are now man's nearest allies, it is somewhat more probable that our early progenitors lived on the African continent than elsewhere. But it is useless to speculate on this subject, for an ape nearly as large as a man, namely the Dryopithecus of Lartet, which was closely allied to the anthropomorphous Hylobates, existed in Europe during the Upper Miocene period; and since so remote a period the earth has certainly undergone many great revolutions, and there has been ample time for migration on the largest scale.
— Charles Darwin, Descent of Man
In 1871 there were hardly any human fossils of ancient hominins available. Almost fifty years later, Darwin's speculation was supported when anthropologists began finding fossils of ancient small-brained hominins in several areas of Africa (list of hominina fossils). The hypothesis of recent (as opposed to archaic) African origin developed in the 20th century. The "Recent African origin" of modern humans means "single origin" (monogenism) and has been used in various contexts as an antonym to polygenism. The debate in anthropology had swung in favour of monogenism by the mid-20th century. Isolated proponents of polygenism held forth in the mid-20th century, such as Carleton Coon, who thought as late as 1962 that H. sapiens arose five times from H. erectus in five places.

Multiregional origin hypothesis

The historical alternative to the recent origin model is the multiregional origin of modern humans, initially proposed by Milford Wolpoff in the 1980s. This view proposes that the derivation of anatomically modern human populations from H. erectus at the beginning of the Pleistocene 1.8 million years BP, has taken place within a continuous world population. The hypothesis necessarily rejects the assumption of an infertility barrier between ancient Eurasian and African populations of Homo. The hypothesis was controversially debated during the late 1980s and the 1990s. The now-current terminology of "recent-origin" and "Out of Africa" became current in the context of this debate in the 1990s. Originally seen as an antithetical alternative to the recent origin model, the multiregional hypothesis in its original "strong" form is obsolete, while its various modified weaker variants have become variants of a view of "recent origin" combined with archaic admixture. Stringer (2014) distinguishes the original or "classic" Multiregional model as having existed from 1984 (its formulation) until 2003, to a "weak" post-2003 variant that has "shifted close to that of the Assimilation Model".

Genetics

In the 1980s, Allan Wilson together with Rebecca L. Cann and Mark Stoneking worked on genetic dating of the matrilineal most recent common ancestor of modern human populations (dubbed "Mitochondrial Eve"). To identify informative genetic markers for tracking human evolutionary history, Wilson concentrated on mitochondrial DNA (mtDNA), passed from mother to child. This DNA material mutates quickly, making it easy to plot changes over relatively short times. With his discovery that human mtDNA is genetically much less diverse than chimpanzee mtDNA, Wilson concluded that modern human populations had diverged recently from a single population while older human species such as Neanderthals and Homo erectus had become extinct. With the advent of archaeogenetics in the 1990s, the dating of mitochondrial and Y-chromosomal haplogroups became possible with some confidence. By 1999, estimates ranged around 150,000 years for the mt-MRCA and 60,000 to 70,000 years for the migration out of Africa.

From 2000–2003, there was controversy about the mitochondrial DNA of "Mungo Man 3" (LM3) and its possible bearing on the multiregional hypothesis. LM3 was found to have more than the expected number of sequence differences when compared to modern human DNA (CRS). Comparison of the mitochondrial DNA with that of ancient and modern aborigines, led to the conclusion that Mungo Man fell outside the range of genetic variation seen in Aboriginal Australians and was used to support the multiregional origin hypothesis. A reanalysis on LM3 and other ancient specimens from the area published in 2016, showed it to be akin to modern Aboriginal Australian sequences, inconsistent with the results of the earlier study.

Y-chromosomal Adam

From Wikipedia, the free encyclopedia

In human genetics, the Y-chromosomal most recent common ancestor (Y-MRCA, informally known as Y-chromosomal Adam) is the most recent common ancestor (MRCA) from whom all currently living men are descended patrilineally. The term Y-MRCA reflects the fact that the Y chromosomes of all currently living human males are directly derived from the Y chromosome of this remote ancestor. The analogous concept of the matrilineal most recent common ancestor is known as "Mitochondrial Eve" (mt-MRCA, named for the matrilineal transmission of mtDNA), the most recent woman from whom all living humans are descended matrilineally. As with "Mitochondrial Eve", the title of "Y-chromosomal Adam" is not permanently fixed to a single individual, but can advance over the course of human history as paternal lineages become extinct.

Estimates of the time when Y-MRCA lived have also shifted as modern knowledge of human ancestry changes. In 2013, the discovery of a previously unknown Y-chromosomal haplogroup was announced, which resulted in a slight adjustment of the estimated age of the human Y-MRCA.

By definition, it is not necessary that the Y-MRCA and the mt-MRCA should have lived at the same time. While estimates as of 2014 suggested the possibility that the two individuals may well have been roughly contemporaneous (albeit with uncertainties ranging in the tens of thousands of years), the discovery of archaic Y-haplogroup has pushed back the estimated age of the Y-MRCA beyond the most likely age of the mt-MRCA. As of 2015, estimates of the age of the Y-MRCA range around 200,000 to 300,000 years ago, roughly consistent with the emergence of anatomically modern humans.

Y-chromosomal data taken from a Neanderthal from El Sidrón, Spain, produced a Y-T-MRCA of 588,000 years ago for neanderthal and Homo sapiens patrilineages, dubbed ante Adam and 275,000 years ago for Y-MRCA.

Definition

The Y-chromosomal most recent common ancestor is the most recent common ancestor of the Y-chromosomes found in currently living human males. 

Due to the definition via the "currently living" population, the identity of a MRCA, and by extension of the human Y-MRCA, is time-dependent (it depends on the moment in time intended by the term "currently"). The MRCA of a population may move forward in time as archaic lineages within the population go extinct: once a lineage has died out, it is irretrievably lost. This mechanism can thus only shift the title of Y-MRCA forward in time. Such an event could be due to the total extinction of several basal haplogroups. The same holds for the concepts of matrilineal and patrilineal MRCAs: it follows from the definition of Y-MRCA that he had at least two sons who both have unbroken lineages that have survived to the present day. If the lineages of all but one of those sons die out, then the title of Y-MRCA shifts forward from the remaining son through his patrilineal descendants, until the first descendant is reached who had at least two sons who both have living, patrilineal descendants. The title of Y-MRCA is not permanently fixed to a single individual, and the Y-MRCA for any given population would himself have been part of a population which had its own, more remote, Y-MRCA.

Although the informal name "Y-chromosomal Adam" is a reference to the biblical Adam, this should not be misconstrued as implying that the bearer of the chromosome was the only human male alive during his time. His other male contemporaries may also have descendants alive today, but not, by definition, through solely patrilineal descent; in other words, none of them have an unbroken male line of descendants (son's son's son's … son) connecting them to currently living people.

By the nature of the concept of most recent common ancestors, these estimates can only represent a terminus ante quem ("limit before which"), until the genome of the entire population has been examined (in this case, the genome of all living humans).

Age estimate

Estimates on the age of the Y-MRCA crucially depend on the most archaic known haplogroup extant in contemporary populations. As of 2018, this is haplogroup A00 (discovered in 2013). Age estimates based on this published during 2014–2015 range between 160,000 and 300,000 years, compatible with the time of emergence and early dispersal of Homo sapiens.

Method

In addition to the tendency of the title of Y-MRCA to shift forward in time, the estimate of the Y-MRCA's DNA sequence, his position in the family tree, the time when he lived, and his place of origin, are all subject to future revisions. 

The following events would change the estimate of who the individual designated as Y-MRCA was:
  • Further sampling of Y chromosomes could uncover previously unknown divergent lineages. If this happens, Y-chromosome lineages would converge on an individual who lived further back in time.
  • The discovery of additional deep rooting mutations in known lineages could lead to a rearrangement of the family tree.
  • Revision of the Y-chromosome mutation rate (see below) can change the estimate of the time when he lived.
The time when Y-MRCA lived is determined by applying a molecular clock to human Y-chromosomes. In contrast to mitochondrial DNA (mtDNA), which has a short sequence of 16,000 base pairs, and mutates frequently, the Y chromosome is significantly longer at 60 million base pairs, and has a lower mutation rate. These features of the Y chromosome have slowed down the identification of its polymorphisms; as a consequence, they have reduced the accuracy of Y-chromosome mutation rate estimates.

Methods of estimating the age of the Y-MRCA for a population of human males whose Y-chromosomes have been sequenced are based on applying the theories of molecular evolution to the Y chromosome. Unlike the autosomes, the human Y-chromosome does not recombine often with the X chromosome during meiosis, but is usually transferred intact from father to son; however, it can recombine with the X chromosome in the pseudoautosomal regions at the ends of the Y chromosome. Mutations occur periodically within the Y chromosome, and these mutations are passed on to males in subsequent generations. 

These mutations can be used as markers to identify shared patrilineal relationships. Y chromosomes that share a specific mutation are referred to as haplogroups. Y chromosomes within a specific haplogroup are assumed to share a common patrilineal ancestor who was the first to carry the defining mutation. (This assumption could be mistaken, as it is possible for the same mutation to occur more than once.) A family tree of Y chromosomes can be constructed, with the mutations serving as branching points along lineages. The Y-MRCA is positioned at the root of the family tree, as the Y chromosomes of all living males are descended from his Y chromosome.

Researchers can reconstruct ancestral Y chromosome DNA sequences by reversing mutated DNA segments to their original condition. The most likely original or ancestral state of a DNA sequence is determined by comparing human DNA sequences with those of a closely related species, usually non-human primates such as chimpanzees and gorillas. By reversing known mutations in a Y-chromosome lineage, a hypothetical ancestral sequence for the MRCA, Y-chromosomal Adam, can be inferred. 

Determining the Y-MRCA's DNA sequence, and the time when he lived, involves identifying the human Y-chromosome lineages that are most divergent from each other—the lineages that share the fewest mutations with each other when compared to a non-human primate sequence in a phylogenetic tree. The common ancestor of the most divergent lineages is therefore the common ancestor of all lineages.

History of estimates

Early estimates of the age for the Y-MRCA published during the 1990s ranged between roughly 200 and 300 kya, Such estimates were later substantially revised downward, as in Thomson et al. 2000, which proposed an age of about 59,000. This date suggested that the Y-MRCA lived about 84,000 years after his female counterpart mt-MRCA (the matrilineal most recent common ancestor), who lived 150,000–200,000 years ago. This date also meant that Y-chromosomal Adam lived at a time very close to, and possibly after, the migration from Africa which is believed to have taken place 50,000–80,000 years ago. One explanation given for this discrepancy in the time depths of patrilineal vs. matrilineal lineages was that females have a better chance of reproducing than males due to the practice of polygyny. When a male individual has several wives, he has effectively prevented other males in the community from reproducing and passing on their Y chromosomes to subsequent generations. On the other hand, polygyny does not prevent most females in a community from passing on their mitochondrial DNA to subsequent generations. This differential reproductive success of males and females can lead to fewer male lineages relative to female lineages persisting into the future. These fewer male lineages are more sensitive to drift and would most likely coalesce on a more recent common ancestor. This would potentially explain the more recent dates associated with the Y-MRCA.

The "hyper-recent" estimate of significantly below 100 kya was again corrected upward in studies of the early 2010s, which ranged at about 120 kya to 160 kya. This revision was due to the discovery of additional mutations and the rearrangement of the backbone of the Y-chromosome phylogeny following the resequencing of Haplogroup A lineages. In 2013, Francalacci et al. reported the sequencing of male-specific single-nucleotide Y-chromosome polymorphisms (MSY-SNPs) from 1204 Sardinian men, which indicated an estimate of 180,000 to 200,000 years for the common origin of all humans through paternal lineage. or again as high as 180 to 200 kya. Also in 2013, Poznik et al. reported the Y-MRCA to have lived between 120,000 and 156,000 years ago, based on genome sequencing of 69 men from 9 different populations. In addition, the same study estimated the age of Mitochondrial Eve to about 99,000 and 148,000 years. As these ranges overlap for a time-range of 28,000 years (148 to 120 kya), the results of this study have been cast in terms of the possibility that "Genetic Adam and Eve may have walked on Earth at the same time" in the popular press.

The announcement of yet another discovery of a previously unknown lineage, haplogroup A00, in 2013, resulted in another shift in the estimate for the age of Y-chromosomal. Elhaik et al. (2014) dated it to between 163,900 and 260,200 years ago (95% CI). Karmin et al. (2015) dated it to between 192,000 and 307,000 years ago (95% CI). The same study reports that non-African populations converge to a cluster of Y-MRCAs in a window close to 50kya (out-of-Africa migration), and an additional bottleneck for non-African populations at about 10kya, interpreted as reflecting cultural changes increasing the variance in male reproductive success (i.e. increased social stratification) in the Neolithic.

Family tree

 
The revised root of the y-chromosome family tree by Cruciani et al. 2011 compared with the family tree from Karafet et al. 2008. This has been further expanded by the discoveries published by Mendez et al. in 2013.
 
Initial sequencing (Karafet et al., 2008) of the human Y chromosome suggested that two most basal Y-chromosome lineages were Haplogroup A and Haplogroup BT. Haplogroup A is found at low frequencies in parts of Africa, but is common among certain hunter-gatherer groups. Haplogroup BT lineages represent the majority of African Y-chromosome lineages and virtually all non-African lineages. Y-chromosomal Adam was represented as the root of these two lineages. Haplogroup A and Haplogroup BT represented the lineages of the two male descendants of Y-chromosomal Adam. 

Cruciani et al. 2011, determined that the deepest split in the Y-chromosome tree was found between two previously reported subclades of Haplogroup A, rather than between Haplogroup A and Haplogroup BT. Subclades A1b and A1a-T are now believed to descend directly from the root of the tree and now represent the lineages of Y-chromosomal Adam's two sons. The rearrangement of the Y-chromosome family tree implies that lineages classified as Haplogroup A do not necessarily form a monophyletic clade. Haplogroup A therefore refers to a collection of lineages that do not possess the markers that define Haplogroup BT, though Haplogroup A includes the most distantly related Y chromosomes. 

The M91 and P97 mutations distinguish Haplogroup A from Haplogroup BT. Within Haplogroup A chromosomes, the M91 marker consists of a stretch of 8 T nucleobase units. In Haplogroup BT and chimpanzee chromosomes, this marker consists of 9 T nucleobase units. This pattern suggested that the 9T stretch of Haplogroup BT was the ancestral version and that Haplogroup A was formed by the deletion of one nucleobase. Haplogroups A1b and A1a were considered subclades of Haplogroup A as they both possessed the M91 with 8Ts.

But according to Cruciani et al. 2011, the region surrounding the M91 marker is a mutational hotspot prone to recurrent mutations. It is therefore possible that the 8T stretch of Haplogroup A may be the ancestral state of M91 and the 9T of Haplogroup BT may be the derived state that arose by an insertion of 1T. This would explain why subclades A1b and A1a-T, the deepest branches of Haplogroup A, both possess the same version of M91 with 8Ts. Furthermore, Cruciani et al. 2011 determined that the P97 marker, which is also used to identify Haplogroup A, possessed the ancestral state in Haplogroup A but the derived state in Haplogroup BT.

Likely geographic origin

As current estimates on TMRCA converge with estimates for the age of anatomically modern humans and well predate the Out of Africa migration, geographical origin hypotheses continue to be limited to the African continent

According to Cruciani et al. 2011, the most basal lineages have been detected in West, Northwest and Central Africa, suggesting plausibility for the Y-MRCA living in the general region of "Central-Northwest Africa".

Scozzari et al. (2012) agreed with a plausible placement in "the north-western quadrant of the African continent" for the emergence of the A1b haplogroup. The 2013 report of haplogroup A00 found among the Mbo people of western present-day Cameroon is also compatible with this picture.

The revision of Y-chromosomal phylogeny since 2011 has affected estimates for the likely geographical origin of Y-MRCA as well as estimates on time depth. By the same reasoning, future discovery of presently-unknown archaic haplogroups in living people would again lead to such revisions. In particular, the possible presence of between 1% and 4% Neanderthal-derived DNA in Eurasian genomes implies that the (unlikely) event of a discovery of a single living Eurasian male exhibiting a Neanderthal patrilineal line would immediately push back T-MRCA ("time to MRCA") to at least twice its current estimate. However, the discovery of a neanderthal Y-chromosome by Mendez et al. suggests the extinction of neanderthal patrilineages, as the lineage inferred from the neanderthal sequence is outside of the range of contemporary human genetic variation. Questions of geographical origin would become part of the debate on Neanderthal evolution from Homo erectus.

Butane

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