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Friday, June 19, 2020

Apex predator

From Wikipedia, the free encyclopedia

The lion is one of Africa's apex land predators.
 
The saltwater crocodile is the largest living reptile and the dominant predator throughout its range.
 
The great white shark (bottom) was originally considered the apex predator of the ocean; however, the killer whale (top) has proved to be a predator of the shark

An apex predator, also known as an alpha predator or top predator, is a predator at the top of a food chain, without natural predators.

Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic levels. Food chains are often far shorter on land, usually limited to being secondary consumers – for example, wolves prey mostly upon large herbivores (primary consumers), which eat plants (primary producers). The apex predator concept is applied in wildlife management, conservation and ecotourism.

Apex predators have a long evolutionary history, dating at least to the Cambrian period when animals such as Anomalocaris dominated the seas.

Humans have for many centuries interacted with apex predators including the wolf, birds of prey and cormorants to hunt game animals, birds, and fish respectively. More recently, humans have started interacting with apex predators in new ways. These include interactions via ecotourism, such as with the tiger shark, and through rewilding efforts, such as the proposed reintroduction of the lynx.

Ecological roles

Effects on community

The great skua is an aerial apex predator, both preying on other seabirds and bullying them for their catches.
 
Apex predators affect prey species' population dynamics and populations of other predators, both in aquatic and in terrestrial ecosystems. Non-native predatory fish, for instance, have sometimes devastated formerly dominant predators. A lake manipulation study found that when the non-native smallmouth bass was removed, lake trout, the suppressed native apex predator, diversified its prey selection and increased its trophic level. As a terrestrial example, the badger, an apex predator, predates on and also competes with the hedgehog, a mesopredator, for food such as insects, small mammals, reptiles, amphibians and ground-nesting bird's eggs. Removal of badgers (in a trial investigating bovine tuberculosis) caused hedgehog densities to more than double. Predators that exert a top-down control on organisms in their community are often considered keystone species. Humans are not considered apex predators because their diets are typically diverse, although human trophic levels increase with consumption of meat.

Effects on ecosystem

Apex predators can have profound effects on ecosystems, as the consequences of both controlling prey density and restricting smaller predators, and may be capable of self-regulation. They are central to the functioning of ecosystems, the regulation of disease, and the maintenance of biodiversity. When introduced to subarctic islands, for example, Arctic foxes' predation of seabirds has been shown to turn grassland into tundra. Such wide-ranging effects on lower levels of an ecosystem are termed trophic cascades. The removal of top-level predators, often through human agency, can cause or disrupt trophic cascades. For example, reduction in the population of sperm whales, apex predators with a fractional trophic level of 4.7, by hunting has caused an increase in the population of large squid, trophic level over 4 (carnivores that eat other carnivores). This effect, called mesopredator release, occurs in terrestrial and marine ecosystems; for instance, in North America, the ranges of all apex carnivores have contracted whereas those of 60% of mesopredators have grown in the past two centuries.

Conservation

The gray wolf is both an apex predator and a keystone species, affecting its prey's behaviour and the wider ecosystem.
 
Because apex predators have powerful effects on other predators, on herbivores, and on plants, they can be important in nature conservation. Humans have hunted many apex predators close to extinction, but in some parts of the world these predators are now returning. They are increasingly threatened by climate change. For example, the polar bear requires extensive areas of sea ice to hunt its prey, typically seals, but climate change is shrinking the sea ice of the Arctic, forcing polar bears to fast on land for increasingly long periods.

Dramatic changes in the Greater Yellowstone Ecosystem were recorded after the gray wolf, both an apex predator and a keystone species (one with a large effect on its ecosystem), was reintroduced to Yellowstone National Park in 1995 as a conservation measure. Elk, the wolves' primary prey, became less abundant and changed their behavior, freeing riparian zones from constant grazing and allowing willows, aspens and cottonwoods to flourish, creating habitats for beaver, moose and scores of other species. In addition to their effect on prey species, the wolves' presence also affected one of the park's vulnerable species, the grizzly bear: emerging from hibernation, having fasted for months, the bears chose to scavenge wolf kills, especially during the autumn as they prepared to hibernate once again. The grizzly bear gives birth during hibernation, so the increased food supply is expected to produce an increase in the numbers of cubs observed. Dozens of other species, including eagles, ravens, magpies, coyotes and black bears have also been documented as scavenging from wolf kills within the park.

Human trophic level

Humans sometimes live by hunting other animals for food and materials such as fur, sinew, and bone, as in this walrus hunt in the Arctic, but their status as apex predators is debated.

Ecologists have debated whether humans are apex predators. For instance, Sylvain Bonhommeau and colleagues argued in 2013 that across the global food web, a fractional human trophic level (HTL) can be calculated as the mean trophic level of every species in the human diet, weighted by the proportion which that species forms in the diet. This analysis gives an average HTL of 2.21, varying between 2.04 (for Burundi, with a 96.7% plant-based diet) and 2.57 (for Iceland, with 50% meat and fish, 50% plants). These values are comparable to those of non-apex predators such as anchovy or pig.

However, Peter D. Roopnarine criticised Bonhommeau's approach in 2014, arguing that humans are apex predators, and that the HTL was based on terrestrial farming where indeed humans have a low trophic level, mainly eating producers (crop plants at level 1) or primary consumers (herbivores at level 2), which as expected places humans at a level slightly above 2. Roopnarine instead calculated the position of humans in two marine ecosystems, a Caribbean coral reef and the Benguela system near South Africa. In these systems, humans mainly eat predatory fish and have a fractional trophic level of 4.65 and 4.5 respectively, which in Roopnarine's view makes those humans apex predators.

Evolutionary history

Anomalocaris was an apex predator in the Cambrian seas.

Apex predators are thought to have existed since at least the Cambrian period, around 500 million years ago. Extinct species cannot be directly determined to be apex predators as their behaviour cannot be observed, and clues to ecological relationships, such as bite marks on bones or shells, do not form a complete picture. However, indirect evidence such as the absence of any discernible predator in an environment is suggestive. Anomalocaris was an aquatic apex predator, in the Cambrian. Its mouthparts are clearly predatory, and there were no larger animals in the seas at that time.

Carnivorous theropod dinosaurs including Allosaurus and Tyrannosaurus have been described as apex predators, based on their size, morphology, and dietary needs.

A Permian shark, Triodus sessilis, was discovered containing two amphibians (Archegosaurus decheni and Cheliderpeton latirostre), one of which had consumed a fish, Acanthodes bronni, showing that the shark had lived at a trophic level of at least 4.

Among more recent fossils, the sabre-tooth cats, like Smilodon, are considered to have been apex predators in the Cenozoic.

Interactions with humans

Dogs have been used in hunting for many centuries, as in this 14th century French depiction of a boar hunt.

In hunting

Humans hunted with wolves and in turn with domestic dogs for some 40,000 years; this collaboration may have helped modern humans to outcompete the Neanderthals. Humans still hunt with dogs, which have often been bred as gun dogs to point to, flush out, or retrieve prey. The Portuguese Water Dog was used to drive fish into nets. Several breeds of dog have been used to chase large prey such as deer and wolves.

Eagles and falcons, which are apex predators, are used in falconry, hunting birds or mammals. Tethered cormorants, also top predators, have been used to catch fish.

In ecotourism

Tiger sharks are popular ecotourism subjects, but their ecosystems may be affected by the food provided to attract them.

Ecotourism sometimes relies on apex predators to attract business. Tour operators may in consequence decide to intervene in ecosystems, for example by providing food to attract predators to areas that can conveniently be visited. This in turn can have effects on predator population and therefore on the wider ecosystem. As a result, provisioning of species such as the tiger shark is controversial, but its effects are not well established by empirical evidence. Other affected apex predators include big cats and crocodiles.

Rewilding

The reintroduction of predators like the lynx is attractive to conservationists, but alarming to farmers.

In some densely populated areas like the British Isles, all the large native predators like the wolf, bear, wolverine and lynx have become locally extinct, allowing herbivores such as deer to multiply unchecked except by hunting. In 2015, plans were made to reintroduce lynx to the counties of Norfolk, Cumbria, and Northumberland in England, and Aberdeenshire in Scotland as part of the rewilding movement. The reintroduction of large predators is controversial, in part because of concern among farmers for their livestock. Conservationists such as Paul Lister propose instead to allow wolves and bears to hunt their prey in a "managed environment" on large fenced reserves.

Alpha (ethology)

From Wikipedia, the free encyclopedia

Male "silverback" gorillas are usually the alpha animal and may receive preferential treatment such as being groomed by other group members

In studies of social animals, the highest ranking individual is sometimes designated as the alpha. Males, females, or both, can be alphas, depending on the species. Where one male and one female fulfill this role together, they are sometimes referred to as the alpha pair. Other animals in the same social group may exhibit deference or other species-specific subordinate behavior towards the alpha or alphas. 

Alpha animals usually gain preferential access to food and other desirable items or activities, though the extent of this varies widely between species. Male or female alphas may gain preferential access to sex or mates; in some species, only alphas or an alpha pair reproduce.

Alphas may achieve their status by superior physical strength and aggression, or through social efforts and building alliances within the group, or more often, simply by breeding and being the parent of all in their pack.

The individual with alpha status sometimes changes, often through a fight between the dominant and a subordinate animal. These fights are often to the death, depending on the animal.

Beta and omega

Social animals in a hierarchical community are sometimes assigned ranks in ethology studies.

Beta animals often act as second-in-command to the reigning alpha or alphas and will act as new alpha animals if an alpha dies or is otherwise no longer considered an alpha. In some species of birds, males pair up in twos when courting, the beta male aiding the alpha male. It has been found that the social context of the animals has a significant impact on courtship behavior and the overall reproductive success of that animal.

Omega has a conflicted meaning. As an antonym it is used to refer to the lowest caste of a hierarchical society. Following the theories that omega animals are subordinate to all others in the community, and are expected by others in the group to remain submissive to everyone. Omega animals may also be used as communal scapegoats or outlets for frustration, or given the lowest priority when distributing food. These same researchers also compared the omega wolf to a "court jester," being first to initiate play, and while submissive, well liked and not outcast. Interestingly, in the pack under study, the omega named Lakota was in fact the largest and strongest of the wolves.

Alternately, and in popular use, is the omega as the romanticized "Lone Wolf" or the omega wolf as discussed by David Mech. Per this pro-omega type, the omega is not the submissive “second place” as a beta is. Rather, they are equally dominant yet in an independent way. Instead of leading a group like an alpha or collaborating like a beta, this omega type is self-reliant, and less collaborative — but also less focused on leading or commanding groups.

Examples

Primates

Common chimpanzees use strength, intelligence, and political alliances to establish and maintain alpha position. Alpha males who solely use intimidation and aggression to keep their position often provoke dissent. Coalitions will eventually form, which at some point will topple the alpha male. There have been rare cases where a group has killed the alpha male. Common chimpanzees show deference to the alpha of the community by ritualized postures and gestures such as presenting their back, crouching, bowing, or bobbing. Chimpanzees lower in rank than the alpha male will offer their hand while grunting to the alpha male as a sign of submission. Bonobo society on the other hand is governed by alpha females. Males will associate with females for rank acquisition because females dominate the social environment. If a male is to achieve alpha status in a bonobo group, he must be accepted by the alpha female. Female bonobos use homosexual sex to increase social status. High-ranking females rarely interact sexually with other females, but low-ranking females interact sexually with all females.

Gorillas use intimidation to establish and maintain alpha position. A study conducted regarding the reproductive behavior of male mountain gorillas (Gorilla beringei beringei) found further evidence that dominant males are favored to father offspring, even when there is a greater number of males in a notably enlarged group size. The study also concluded that mating access dropped off less steeply with status; alpha, beta, and gamma showing more similar mating success, compared to what had been previously thought.

A study on the association of alpha males and females during the non-breeding season in wild Capuchin monkeys (Cebus apella nigritus) examined whether alpha males are the preferred mate for females and, secondly, whether female-alpha status and relationship to the alpha-male can be explained through the individual characteristics and or social network of the female. The results indicated that alpha male Capuchin are the preferred mate for adult females. However, only the alpha females had strong interactions with the alpha males by virtue of a dominance hierarchy among the females in which only the most dominant and strong females were able to interact with the alpha male.

Researcher M.W. Foster investigated primates and found that the leaders were more likely to be those who did more for those around them instead of being determined by strength.

Canines

In the past, the prevailing view on gray wolf packs was that they consisted of individuals vying with each other for dominance, with dominant gray wolves being referred to as the "alpha" male and female, and the subordinates as "beta" and "omega" wolves. This terminology was first used in 1947 by Rudolf Schenkel of the University of Basel, who based his findings on researching the behavior of captive gray wolves. This view on gray wolf pack dynamics was later popularized by the researcher L. David Mech in his 1970 book The Wolf. He later found additional evidence that the concept of an Alpha male may have been an interpretation of incomplete data and formally disavowed this terminology in 1999. He explained that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves. Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and their offspring of the previous 1–3 years. In the article, Mech wrote that the use of the term "alpha" to describe the breeding pair adds no additional information, and is "no more appropriate than referring to a human parent or a doe deer as an alpha." He further notes the terminology falsely implies a "force-based dominance hierarchy." In 13 years of summer observations of wild wolves, he witnessed no dominance contests between them.

In some other wild canids, the alpha male may not have exclusive access to the alpha female. Other pack members as in the African painted dog (Lycaon pictus) may guard the maternity den used by the alpha female.

Elite

From Wikipedia, the free encyclopedia

Political cartoon from October 1884, showing wealthy plutocrats feasting at a table while a poor family begs beneath

In political and sociological theory, the elite (French élite, from Latin eligere, to select or to sort out) are a small group of powerful people who hold a disproportionate amount of wealth, privilege, political power, or skill in a society. Defined by the Cambridge Dictionary, the "elite" are "those people or organizations that are considered the best or most powerful compared to others of a similar type."

American sociologist C. Wright Mills states that the power elite members recognize other members' mutual exalted position in society. "As a rule, 'they accept one another, understand one another, marry one another, tend to work, and to think, if not together at least alike'." "It is a well-regulated existence where education plays a critical role. Youthful upper-class members attend prominent preparatory schools, which not only open doors to such elite universities as Harvard, Columbia, Dartmouth College, Yale, and Princeton, but also to the universities' highly exclusive clubs. These memberships in turn pave the way to the prominent social clubs located in all major cities and serving as sites for important business contacts".

Elitist privilege

According to Mills, men receive the education necessary for elitist privilege to obtain their background and contacts, allowing them to enter three branches of the power elite, which are;
  • The Political Leadership: Mills contended that since the end of World War II, corporate leaders had become more prominent in the political process, with a decline in central decision-making for professional politicians.
  • The Military Circle: In Mills' time a heightened concern about warfare existed, making top military leaders and such issues as defense funding and personnel recruitment very important. Most prominent corporate leaders and politicians were strong proponents of military spending.
  • The Corporate Elite: According to Mills, in the 1950s when the military emphasis was pronounced, it was corporate leaders working with prominent military officers who dominated the development of policies. These two groups tended to be mutually supportive.
According to Mills, the governing elite in the United States primarily draws its members from political leaders, including the president, and a handful of key cabinet members, as well as close advisers, major corporate owners and directors, and high-ranking military officers. These groups overlap and elites tend to circulate from one sector to another, consolidating power in the process.
Unlike the ruling class, a social formation based on heritage and social ties, the power elite is characterized by the organizational structures through which its wealth is acquired. According to Mills, the power elite rose from "the managerial reorganization of the propertied classes into the more or less unified stratum of the corporate rich". Domhoff further clarified the differences in the two terms: "The upper class as a whole does not do the ruling. Instead, class rule is manifested through the activities of a wide variety of organizations and institutions...Leaders within the upper class join with high-level employees in the organizations they control to make up what will be called the power elite".

The Marxist theoretician Nikolai Bukharin anticipated the elite theory in his 1929 work, Imperialism and World Economy: "present-day state power is nothing but an entrepreneurs' company of tremendous power, headed even by the same persons that occupy the leading positions in the banking and syndicate offices".

Power elite

The power elite is a term used by Mills to describe a relatively small, loosely connected group of individuals who dominate American policymaking. This group includes bureaucratic, corporate, intellectual, military, media, and government elites who control the principal institutions in the United States and whose opinions and actions influence the decisions of the policymakers. The basis for membership of a power elite is institutional power, namely an influential position within a prominent private or public organization. A study of the French corporate elite has shown that social class continues to hold sway in determining who joins this elite group, with those from the upper-middle class tending to dominate. Another study (published in 2002) of power elites in the United States under President George W. Bush (in office 2001-2009) identified 7,314 institutional positions of power encompassing 5,778 individuals. A later study of U.S. society noted demographic characteristics of this elite group as follows:
Age
Corporate leaders aged about 60; heads of foundations, law, education, and civic organizations aged around 62; government employees aged about 56.
Gender
Men contribute roughly 80% in the political realm whereas women contribute roughly only 20% in the political realm. In the economic denomination, as of October 2017, only 32 (6.4%) of the Fortune 500 CEOs are women.
Ethnicity
White Anglo-Saxons dominate in the power elite, with Protestants representing about 80% of the top business leaders, and about 73% of members of Congress. As of October 2017, only 4 (0.8%) of the Fortune 500 CEOs are African American. In similarly low proportions, as of October 2017, 10 (2%) of the Fortune 500 CEOs are Latino, and 10 (2%) are Asian.
Education
Nearly all the leaders have a college education, with almost half graduating with advanced degrees. About 54% of the big-business leaders, and 42% of the government elite graduated from just 12 prestigious universities with large endowments.
Social clubs
Most holders of top positions in the power elite possess exclusive membership to one or more social clubs. About a third belong to a small number of especially prestigious clubs in major cities like London, New York City, Chicago, Boston, and Washington, D.C.

Impacts on economy

In the 1970s an organized set of policies promoted reduced taxes, especially for the wealthy, and a steady erosion of the welfare safety net. Starting with legislation in the 1980s, the wealthy banking community successfully lobbied for reduced regulation. The wide range of financial and social capital accessible to the power elite gives their members heavy influence in economic and political decision making, allowing them to move toward attaining desired outcomes. Sociologist Christopher Doob gives a hypothetical alternative, stating that these elite individuals would consider themselves the overseers of the national economy. Also appreciating that it is not only a moral, but a practical necessity to focus beyond their group interests. Doing so would hopefully alleviate various destructive conditions affecting large numbers of less affluent citizens.

Global politics and hegemony

Mills determined that there is an "inner core" of the power elite involving individuals that are able to move from one seat of institutional power to another. They, therefore, have a wide range of knowledge and interests in many influential organizations, and are, as Mills describes, "professional go-betweens of economic, political, and military affairs". Relentless expansion of capitalism and the globalizing of economic and military power, binds leaders of the power elite into complex relationships with nation states that generate global-scale class divisions. Sociologist Manuel Castells writes in The Rise of the Network Society that contemporary globalization does not mean that "everything in the global economy is global". So, a global economy becomes characterized by fundamental social inequalities with respect to the "level of integration, competitive potential and share of the benefits from economic growth". Castells cites a kind of "double movement" where on one hand, "valuable segments of territories and people" become "linked in the global networks of value making and wealth appropriation", while, on the other, "everything and everyone" that is not valued by established networks gets "switched off...and ultimately discarded". The wide-ranging effects of global capitalism ultimately affect everyone on the planet, as economies around the world come to depend on the functioning of global financial markets, technologies, trade and labor.

Political party

From Wikipedia, the free encyclopedia

A political party is an organized group of people who have the same ideology, or who otherwise have the same political positions, and who field candidates for elections, in an attempt to get them elected and thereby implement the party's agenda. They are a defining element of representative democracy.

While there is some international commonality in the way political parties are recognized and in how they operate, there are often many differences, and some are significant. Most of political parties have an ideological core, but some do not, and many represent ideologies very different from their ideology at the time the party was founded. Many countries, such as Germany and India, have several significant political parties, and some nations have one-party systems, such as China and Cuba. The United States is in practice a two-party system but with many smaller parties also participating.

Historical development

The idea of people forming large groups or factions to advocate for their shared interests is ancient. Plato mentions the political factions of Classical Athens in the Republic, and Aristotle discusses the tendency of different types of government to produce factions in the Politics. Certain ancient disputes were also factional, like the Nika riots between two chariot racing factions at the Hippodrome of Constantinople. However, modern political parties are considered to have emerged around the end of the 18th or early 19th centuries, appearing first in Europe and the United States. What distinguishes political parties from factions and interest groups is that political parties use an explicit label to identify their members as having shared electoral and legislative goals. The transformation from loose factions into organised modern political parties is considered to have first occurred in either the United Kingdom or the United States, with the United Kingdom's Conservative Party and the Democratic Party of the United States both frequently called the world's "oldest continuous political party".

Emergence in Britain

The party system that emerged in early modern Britain is considered to be one of the world's first, with origins in the factions that emerged from the Exclusion Crisis and Glorious Revolution of the late 17th century. The Whig faction originally organised itself around support for Protestant constitutional monarchy as opposed to absolute rule, whereas the conservative Tory faction (originally the Royalist or Cavalier faction of the English Civil War) supported a strong monarchy. These two groups structured disputes in the politics of the United Kingdom throughout the 18th century. Throughout the next several centuries, these loose factions began to adopt more coherent political tendencies and ideologies: the liberal political ideas of John Locke and the notion of universal rights espoused by theorists like Algernon Sidney and later John Stuart Mill were major influences on the Whigs, whereas the Tories eventually came to be identified with conservative philosophers like Edmund Burke.

The period between the advent of factionalism, around the Glorious Revolution, and the accession of George III in 1760 was characterised by Whig supremacy, during which the Whigs remained the most powerful bloc and consistently championed constitutional monarchy with strict limits on the monarch's power, opposed the accession of a Catholic king, and believed in extending toleration to nonconformist Protestants and dissenters. Although the Tories were out of office for half a century, they largely remained a united opposition to the Whigs.

When they lost power, the old Whig leadership dissolved into a decade of factional chaos with distinct Grenvillite, Bedfordite, Rockinghamite, and Chathamite factions successively in power, and all referring to themselves as "Whigs". The first distinctive political parties emerged from this chaos. The first such party was the Rockingham Whigs under the leadership of Charles Watson-Wentworth and the intellectual guidance of the political philosopher Edmund Burke. Burke laid out a philosophy that described the basic framework of the political party as "a body of men united for promoting by their joint endeavours the national interest, upon some particular principle in which they are all agreed". As opposed to the instability of the earlier factions, which were often tied to a particular leader and could disintegrate if removed from power, the party was centred around a set of core principles and remained out of power as a united opposition to government.

In A Block for the Wigs (1783), James Gillray caricatured Fox's return to power in a coalition with North. George III is the blockhead in the centre.

A coalition including the Rockingham Whigs, led by the Earl of Shelburne, took power in 1782, only to collapse after Rockingham's death. The new government, led by the radical politician Charles James Fox in coalition with Lord North, was soon brought down and replaced by William Pitt the Younger in 1783. It was now that a genuine two-party system began to emerge, with Pitt leading the new Tories against a reconstituted "Whig" party led by Fox. The modern Conservative Party was created out of these Pittite Tories. In 1859 under Lord Palmerston, the Whigs, heavily influenced by the classical liberal ideas of Adam Smith, joined together with the free trade Tory followers of Robert Peel and the independent Radicals to form the Liberal Party.

Emergence in the United States

Although the framers of the 1787 United States Constitution did not anticipate that American political disputes would be primarily organised around political parties, political controversies in the early 1790s over the extent of federal government powers saw the emergence of two proto-political parties: the Federalist Party and the Democratic-Republican Party, which were championed by Alexander Hamilton and Thomas Jefferson, respectively. However, a consensus reached on these issues ended party politics in 1816 for nearly a decade, a period commonly known as the Era of Good Feelings.

The splintering of the Democratic-Republican Party in the aftermath of the contentious 1824 presidential election led to the re-emergence of political parties. Two major parties would dominate the political landscape for the next quarter-century: the Democratic Party, led by Andrew Jackson, and the Whig Party, established by Henry Clay from the National Republicans and from other Anti-Jackson groups. When the Whig Party fell apart in the mid-1850s, its position as a major U.S. political party was filled by the Republican Party.

Worldwide spread


Another candidate for the first modern party system to emerge is that of Sweden. Throughout the second half of the 19th century, the party model of politics was adopted across Europe. In Germany, France, Austria and elsewhere, the 1848 Revolutions sparked a wave of liberal sentiment and the formation of representative bodies and political parties. The end of the century saw the formation of large socialist parties in Europe, some conforming to the philosophy of Karl Marx, others adapting social democracy through the use of reformist and gradualist methods.

At the same time, the Home Rule League Party, campaigning for Home Rule for Ireland in the British Parliament, was fundamentally changed by the Irish political leader Charles Stewart Parnell in the 1880s. In 1882, he changed his party's name to the Irish Parliamentary Party and created a well-organized grassroots structure, introducing membership to replace ad hoc informal groupings. He created a new selection procedure to ensure the professional selection of party candidates committed to taking their seats, and in 1884 he imposed a firm 'party pledge' which obliged MPs to vote as a bloc in parliament on all occasions. The creation of a strict party whip and a formal party structure was unique at the time, preceded only by the Social Democratic Party of Germany (1875), even though the latter was persecuted by Otto von Bismarck from 1878 to 1890. These parties' efficient structure and control contrasted with the loose rules and flexible informality found in the main British parties, and represented the development of new forms of party organisation, which constituted a "model" in the 20th-century.

Origin of political parties

Political parties are a nearly ubiquitous feature of modern countries. Nearly all democratic countries have strong political parties, and many political scientists consider countries with fewer than two parties to necessarily be autocratic. However, these sources allow that a country with multiple competitive parties is not necessarily democratic, and the politics of many autocratic countries are organised around one dominant political party. There are many explanations for how and why political parties are such a crucial part of modern states.

Social cleavages

One of the core explanations for why political parties exist is that they arise from existing divisions among people. Building on Harold Hotelling's work on the aggregation of preferences and Duncan Black's development of social choice theory, Anthony Downs showed how an underlying distribution of preferences in an electorate can produce regular results in the aggregate, such as the median voter theorem. This abstract model shows that parties can arise from variations within an electorate, and can adjust themselves to the patterns in the electorate. However, Downs assumed that some distribution of preferences exists, rather than attributing any meaning to that distribution.

Seymour Martin Lipset and Stein Rokkan made the idea of differences within an electorate more concrete by arguing that several major party systems of the 1960s were the result of social cleavages that had already existed in the 1920s. They identify four lasting cleavages in the countries they examine: a Center-Periphery cleavage regarding religion and language, a State-Church cleavage centered on control of mass education, a Land-Industry cleavage regarding freedom of industry and agricultural policies, and an Owner-Worker cleavage which includes a conflict between nationalism and internationalism. Subsequent authors have expanded on or modified these cleavages, particularly when examining parties in other parts of the world.

The argument that parties are produced by social cleavages has drawn several criticisms. Some authors have challenged the theory on empirical grounds, either finding no evidence for the claim that parties emerge from existing cleavages or arguing that this claim is not empirically testable. Others note that while social cleavages might cause political parties to exist, this obscures the opposite effect: that political parties also cause changes in the underlying social cleavages. A further objection is that, if the explanation for where parties come from is that they emerge from existing social cleavages, then the theory has not identified what causes parties unless it also explains where social cleavages come from; one response to this objection, along the lines of Charles Tilly's bellicist theory of state-building, is that social cleavages are formed by historical conflicts.

Individual and group incentives

An alternative explanation for why parties are ubiquitous across the world is that the formation of parties provides compatible incentives for candidates and legislators. One explanation for the existence of parties, advanced by John Aldrich, is that the existence of political parties means that a candidate in one electoral district has an incentive to assist a candidate in a different district, when those two candidates have a similar ideology.

One reason that this incentive exists is that parties can solve certain legislative challenges that a legislature of unaffiliated members might face. Gary W. Cox and Mathew D. McCubbins argue that the development of many institutions can be explained by their power to constrain the incentives of individuals; a powerful institution can prohibit individuals from acting in ways that harm the community. This suggests that political parties might be mechanisms for preventing candidates with similar ideologies from acting to each other's detriment. One specific advantage that candidates might obtain from helping similar candidates in other districts is that the existence of a party apparatus can help coalitions of electors to agree on ideal policy choices, which is in general not possible. This could be true even in contexts where it is only slightly beneficial to be part of a party; models of how individuals coordinate on joining a group or participating in an event show how even a weak preference to be part of a group can provoke mass participation.

Parties as heuristics

Parties may be necessary for many individuals to participate in politics, because they provide a massively simplifying heuristic which allows people to make informed choices with a much lower cognitive cost. Without political parties, electors would have to evaluate every individual candidate in every single election they are eligible to vote in. Instead, parties enable electors to make judgments about a few groups instead of a much larger number of individuals. Angus Campbell, Philip Converse, Warren Miller, and Donald E. Stokes argued in The American Voter that identification with a political party is a crucial determinant of whether and how an individual will vote. Because it is much easier to become informed about a few parties' platforms than about many candidates' personal positions, parties reduce the cognitive burden for people to cast informed votes. However, evidence suggests that over the last several decades the strength of party identification has been weakening, so this may be a less important function for parties to provide than it was in the past.

Structure

A political party is typically led by a party leader (the most powerful member and spokesperson representing the party), a party secretary (who maintains the daily work and records of party meetings), party treasurer (who is responsible for membership dues) and party chair (who forms strategies for recruiting and retaining party members, and also chairs party meetings). Most of the above positions are also members of the party executive, the leading organization which sets policy for the entire party at the national level. The structure is far more decentralized in the United States because of the separation of powers, federalism and the multiplicity of economic interests and religious sects. Even state parties are decentralized as county and other local committees are largely independent of state central committees. The national party leader in the U.S. will be the president, if the party holds that office, or a prominent member of Congress in opposition (although a big-state governor may aspire to that role). Officially, each party has a chairman for its national committee who is a prominent spokesman, organizer and fund-raiser, but without the status of prominent elected office holders. 

In parliamentary democracies, on a regular, periodic basis, party conferences are held to elect party officers, although snap leadership elections can be called if enough members opt for such. Party conferences are also held in order to affirm party values for members in the coming year. American parties also meet regularly and, again, are more subordinate to elected political leaders.

Depending on the demographic spread of the party membership, party members form local or regional party committees in order to help candidates run for local or regional offices in government. These local party branches reflect the officer positions at the national level.

It is also customary for political party members to form wings for current or prospective party members, most of which fall into the following two categories:
  • identity-based: including youth wings and/or armed wings
  • position-based: including wings for candidates, mayors, governors, professionals, students, etc. The formation of these wings may have become routine but their existence is more of an indication of differences of opinion, intra-party rivalry, the influence of interest groups, or attempts to wield influence for one's state or region.
These are useful for party outreach, training and employment. Many young aspiring politicians seek these roles and jobs as stepping stones to their political careers in legislative or executive offices.
The internal structure of political parties has to be democratic in some countries. In Germany Art. 21 Abs. 1 Satz 3 GG establishes a command of inner-party democracy.

Parliamentary parties

When the party is represented by members in the lower house of parliament, the party leader simultaneously serves as the leader of the parliamentary group of that full party representation; depending on a minimum number of seats held, Westminster-based parties typically allow for leaders to form frontbench teams of senior fellow members of the parliamentary group to serve as critics of aspects of government policy. When a party becomes the largest party not part of the Government, the party's parliamentary group forms the Official Opposition, with Official Opposition frontbench team members often forming the Official Opposition Shadow cabinet. When a party achieves enough seats in an election to form a majority, the party's frontbench becomes the Cabinet of government ministers. They are all elected members. There are members who attend party without promotion.

Regulation

The freedom to form, declare membership in, or campaign for candidates from a political party is considered a measurement of a state's adherence to liberal democracy as a political value. Regulation of parties may run from a crackdown on or repression of all opposition parties, a norm for authoritarian governments, to the repression of certain parties which hold or promote ideals which run counter to the general ideology of the state's incumbents (or possess membership by-laws which are legally unenforceable).

Furthermore, in the case of far-right, far-left and regionalism parties in the national parliaments of much of the European Union, mainstream political parties may form an informal cordon sanitaire which applies a policy of non-cooperation towards those "Outsider Parties" present in the legislature which are viewed as 'anti-system' or otherwise unacceptable for government. Cordons sanitaire, however, have been increasingly abandoned over the past two decades in multi-party democracies as the pressure to construct broad coalitions in order to win elections – along with the increased willingness of outsider parties themselves to participate in government – has led to many such parties entering electoral and government coalitions.

Starting in the second half of the 20th century, modern democracies have introduced rules for the flow of funds through party coffers, e.g. the Canada Election Act 1976, the PPRA in the U.K. or the FECA in the U.S. Such political finance regimes stipulate a variety of regulations for the transparency of fundraising and expenditure, limit or ban specific kinds of activity and provide public subsidies for party activity, including campaigning.

Partisan style

Partisan style varies according to each jurisdiction, depending on how many parties there are, and how much influence each individual party has.

Nonpartisan systems

In a nonpartisan system, no official political parties exist, sometimes reflecting legal restrictions on political parties. In nonpartisan elections, each candidate is eligible for office on his or her own merits. In nonpartisan legislatures, there are no typically formal party alignments within the legislature. The administration of George Washington and the first few sessions of the United States Congress were nonpartisan. Washington also warned against political parties during his Farewell Address. In the United States, the unicameral legislature of Nebraska is nonpartisan but is elected and often votes on informal party lines. In Canada, the territorial legislatures of the Northwest Territories and Nunavut are nonpartisan. In New Zealand, Tokelau has a nonpartisan parliament. Many city and county governments in the United States and Canada are nonpartisan. Nonpartisan elections and modes of governance are common outside of state institutions. Unless there are legal prohibitions against political parties, factions within nonpartisan systems often evolve into political parties.

Uni-party systems

In one-party systems, one political party is legally allowed to hold effective power. Although minor parties may sometimes be allowed, they are legally required to accept the leadership of the dominant party. This party may not always be identical to the government, although sometimes positions within the party may in fact be more important than positions within the government. North Korea and China are examples; others can be found in Fascist states, such as Nazi Germany between 1934 and 1945. The one-party system is thus often equated with dictatorships and tyranny.

In dominant-party systems, opposition parties are allowed, and there may be even a deeply established democratic tradition, but other parties are widely considered to have no real chance of gaining power. Sometimes, political, social and economic circumstances, and public opinion are the reason for others parties' failure. Sometimes, typically in countries with less of an established democratic tradition, it is possible the dominant party will remain in power by using patronage and sometimes by voting fraud. In the latter case, the definition between dominant and one-party system becomes rather blurred. Examples of dominant party systems include the People's Action Party in Singapore, the African National Congress in South Africa, the Cambodian People's Party in Cambodia, the Liberal Democratic Party in Japan, and the National Liberation Front in Algeria. One-party dominant system also existed in Mexico with the Institutional Revolutionary Party until the 1990s, in the southern United States with the Democratic Party from the late 19th century until the 1970s, in Indonesia with the Golkar from the early 1970s until 1998.

Bi-party systems

Two-party systems are states such as Honduras, Jamaica, Malta, Ghana and the United States in which there are two political parties dominant to such an extent that electoral success under the banner of any other party is almost impossible. One right wing coalition party and one left wing coalition party is the most common ideological breakdown in such a system but in two-party states political parties are traditionally catch all parties which are ideologically broad and inclusive.

The United States has gone through several party systems, each of which has been essentially two-party in nature. The divide has typically been between a conservative and liberal party; presently, the Republican Party and Democratic Party serve these roles. Third parties have seen extremely little electoral success, and successful third party runs typically lead to vote splitting due to the first-past-the-post, winner-takes-all systems used in most US elections. There have been several examples of third parties siphoning votes from major parties, such as Theodore Roosevelt in 1912 and George Wallace in 1968, resulting in the victory of the opposing major party. In presidential elections, the Electoral College system has prevented third party candidates from being competitive, even when they have significant support (such as in 1992). More generally, parties with a broad base of support across regions or among economic and other interest groups have a greater chance of winning the necessary plurality in the U.S.'s largely single-member district, winner-take-all elections.

The UK political system, while technically a multi-party system, has functioned generally as a two-party (sometimes called a "two-and-a-half party") system; since the 1920s the two largest political parties have been the Conservative Party and the Labour Party. Before the Labour Party rose in British politics the Liberal Party was the other major political party along with the Conservatives. Though coalition and minority governments have been an occasional feature of parliamentary politics, the first-past-the-post electoral system used for general elections tends to maintain the dominance of these two parties, though each has in the past century relied upon a third party to deliver a working majority in Parliament. (A plurality voting system usually leads to a two-party system, a relationship described by Maurice Duverger and known as Duverger's Law.) There are also numerous other parties that hold or have held a number of seats in Parliament.

Multi-party systems

A poster for the European Parliament election 2004 in Italy, showing party lists

Multi-party systems are systems in which more than two parties are represented and elected to public office.

Australia, Canada, Nepal, Pakistan, India, Ireland, United Kingdom and Norway are examples of countries with two strong parties and additional smaller parties that have also obtained representation. The smaller or "third" parties may hold the balance of power in a parliamentary system, and thus may be invited to form a part of a coalition government together with one of the larger parties, or may provide a supply and confidence agreement to the government; or may instead act independently from the dominant parties. 

More commonly, in cases where there are three or more parties, no one party is likely to gain power alone, and parties have to work with each other to form coalition governments. This is almost always the case in Germany on national and state level, and in most constituencies at the communal level. Furthermore, since the forming of the Republic of Iceland there has never been a government not led by a coalition, usually involving the Independence Party or the Progressive Party. A similar situation exists in the Republic of Ireland, where no one party has held power on its own since 1989. Since then, numerous coalition governments have been formed. These coalitions have been led exclusively by either Fianna Fáil or Fine Gael.

Political change is often easier with a coalition government than in one-party or two-party dominant systems. If factions in a two-party system are in fundamental disagreement on policy goals, or even principles, they can be slow to make policy changes, which appears to be the case now in the U.S. with power split between Democrats and Republicans. Still coalition governments struggle, sometimes for years, to change policy and often fail altogether, post World War II France and Italy being prime examples. When one party in a two-party system controls all elective branches, however, policy changes can be both swift and significant. Democrats Woodrow Wilson, Franklin Roosevelt and Lyndon Johnson were beneficiaries of such fortuitous circumstances, as were Republicans as far removed in time as Abraham Lincoln and Ronald Reagan. Barack Obama briefly had such an advantage between 2009 and 2011.

Funding

Political parties are funded by contributions from
Political parties, still called factions by some, especially those in the governmental apparatus, are lobbied vigorously by organizations, businesses and special interest groups such as trade unions. Money and gifts-in-kind to a party, or its leading members, may be offered as incentives. Such donations are the traditional source of funding for all right-of-centre cadre parties. Starting in the late 19th century these parties were opposed by the newly founded left-of-centre workers' parties. They started a new party type, the mass membership party, and a new source of political fundraising, membership dues.

From the second half of the 20th century on parties which continued to rely on donations or membership subscriptions ran into mounting problems. Along with the increased scrutiny of donations there has been a long-term decline in party memberships in most western democracies which itself places more strains on funding. For example, in the United Kingdom and Australia membership of the two main parties in 2006 is less than an 1/8 of what it was in 1950, despite significant increases in population over that period.

In some parties, such as the post-communist parties of France and Italy or the Sinn Féin party and the Socialist Party, elected representatives (i.e. incumbents) take only the average industrial wage from their salary as a representative, while the rest goes into party coffers. Although these examples may be rare nowadays, "rent-seeking" continues to be a feature of many political parties around the world.

In the United Kingdom, it has been alleged that peerages have been awarded to contributors to party funds, the benefactors becoming members of the House of Lords and thus being in a position to participate in legislating. Famously, Lloyd George was found to have been selling peerages. To prevent such corruption in the future, Parliament passed the Honours (Prevention of Abuses) Act 1925 into law. Thus the outright sale of peerages and similar honours became a criminal act. However, some benefactors are alleged to have attempted to circumvent this by cloaking their contributions as loans, giving rise to the 'Cash for Peerages' scandal.

Such activities as well as assumed "influence peddling" have given rise to demands that the scale of donations should be capped. As the costs of electioneering escalate, so the demands made on party funds increase. In the UK some politicians are advocating that parties should be funded by the state; a proposition that promises to give rise to interesting debate in a country that was the first to regulate campaign expenses (in 1883).

In many other democracies such subsidies for party activity (in general or just for campaign purposes) have been introduced decades ago. Public financing for parties and/ or candidates (during election times and beyond) has several permutations and is increasingly common. Germany, Sweden, Israel, Canada, Australia, Austria and Spain are cases in point. More recently among others France, Japan, Mexico, the Netherlands and Poland have followed suit.

There are two broad categories of public funding, direct, which entails a monetary transfer to a party, and indirect, which includes broadcasting time on state media, use of the mail service or supplies. According to the Comparative Data from the ACE Electoral Knowledge Network, out of a sample of over 180 nations, 25% of nations provide no direct or indirect public funding, 58% provide direct public funding and 60% of nations provide indirect public funding. Some countries provide both direct and indirect public funding to political parties. Funding may be equal for all parties or depend on the results of previous elections or the number of candidates participating in an election. Frequently parties rely on a mix of private and public funding and are required to disclose their finances to the Election management body.

In fledgling democracies funding can also be provided by foreign aid. International donors provide financing to political parties in developing countries as a means to promote democracy and good governance. Support can be purely financial or otherwise. Frequently it is provided as capacity development activities including the development of party manifestos, party constitutions and campaigning skills. Developing links between ideologically linked parties is another common feature of international support for a party. Sometimes this can be perceived as directly supporting the political aims of a political party, such as the support of the US government to the Georgian party behind the Rose Revolution. Other donors work on a more neutral basis, where multiple donors provide grants in countries accessible by all parties for various aims defined by the recipients. There have been calls by leading development think-tanks, such as the Overseas Development Institute, to increase support to political parties as part of developing the capacity to deal with the demands of interest-driven donors to improve governance.

Colors and emblems

Generally speaking, over the world, political parties associate themselves with colors, primarily for identification, especially for voter recognition during elections.
Color associations are useful when it is not desirable to make rigorous links to parties, particularly when coalitions and alliances are formed between political parties and other organizations, for example: "Purple" (Red-Blue) alliances, Red-green alliances, Blue-green alliances, Traffic light coalitions, Pan-green coalitions, and Pan-blue coalitions.




Political color schemes in the United States diverge from international norms. Since 2000, red has become associated with the right-wing Republican Party and blue with the left-wing Democratic Party. However, unlike political color schemes of other countries, the parties did not choose those colors; they were used in news coverage of the 2000 election results and ensuing legal battle and caught on in popular usage. Prior to the 2000 election the media typically alternated which color represented which party each presidential election cycle. The color scheme happened to get inordinate attention that year, so the cycle was stopped lest it cause confusion the following election.

Emblems

The emblem of socialist parties is often a red rose held in a fist. Communist parties often use a hammer to represent the worker, a sickle to represent the farmer, or both a hammer and a sickle to refer to both at the same time.

The emblem of Nazism, the swastika or "hakenkreuz", has been adopted as a near-universal symbol for almost any organised white supremacist group, even though it dates from more ancient times.

International organization

During the 19th and 20th century, many national political parties organized themselves into international organizations along similar policy lines. Notable examples are The Universal Party, International Workingmen's Association (also called the First International), the Socialist International (also called the Second International), the Communist International (also called the Third International), and the Fourth International, as organizations of working class parties, or the Liberal International (yellow), Hizb ut-Tahrir, Christian Democratic International and the International Democrat Union (blue). Organized in Italy in 1945, the International Communist Party, since 1974 headquartered in Florence has sections in six countries. Worldwide green parties have recently established the Global Greens. The Universal Party, The Socialist International, the Liberal International, and the International Democrat Union are all based in London. Some administrations (e.g. Hong Kong) outlaw formal linkages between local and foreign political organizations, effectively outlawing international political parties.

Types

Klaus von Beyme categorised European parties into nine families, which described most parties. He was able to arrange seven of them from left to right: Communist, Socialist, Green, Liberal, Christian democratic, Conservative and Libertarian. The position of two other types, Agrarian and Regional/Ethnic parties varied.

Political scientists have distinguished between different types of political parties that have evolved throughout history. These include cadre parties, mass parties, catch-all parties and cartel parties. Cadre parties were political elites that were concerned with contesting elections and restricted the influence of outsiders, who were only required to assist in election campaigns. Mass parties tried to recruit new members who were a source of party income and were often expected to spread party ideology as well as assist in elections. In the United States, where both major parties were cadre parties, the introduction of primaries and other reforms has transformed them so that power is held by activists who compete over influence and nomination of candidates.

Cadre party

A cadre party, or elite party, is a type of political party that was dominant in the nineteenth century before the introduction of universal suffrage and that was made up of a collection of individuals or political elites. The French political scientist Marcel Duverger first distinguished between “cadre” and “mass” parties, founding his distinction on the differences within the organisational structures of these two types. Cadre parties are characterised by minimal and loose organisation, and are financed by fewer larger monetary contributions typically originating from outside the party. Cadre parties give little priority to expanding the party’s membership base, and its leaders are its only members. The earliest parties, such as the early American political parties, the Democratic-Republicans and the Federalists, are classified as cadre parties.

Mass party

A mass party is a type of political party that developed around cleavages in society and mobilised the ordinary citizens or 'masses' in the political process. In Europe, the introduction of universal suffrage resulted in the creation of worker’s parties that later evolved into mass parties; an example is the German Social Democratic Party. These parties represented large groups of citizens who had previously not been represented in political processes, articulating the interests of different groups in society. In contrast to cadre parties, mass parties are funded by their members, and rely on and maintain a large membership base. Further, mass parties prioritise the mobilisation of voters and are more centralised than cadre parties.

Catch-all party

The catch-all party, also called the 'big tent' party, is a term developed by German-American political scientist Otto Kirchheimer used to describe the parties that developed in the 1950s and 1960s from changes within the mass parties. Kirchheimer characterised the shift from the traditional mass parties to catch-all parties as a set of developments including the “drastic reduction of the party’s ideological baggage” and the "downgrading of the role of the individual party member". By broadening their central ideologies into more open-ended ones, catch-all parties seek to secure the support of a wider section of the population. Further, the role of members is reduced as catch-all parties are financed in part by the state or by donations. In Europe, the shift of Christian Democratic parties that were organised around religion into broader centre-right parties epitomises this type.

Cartel party

Cartel parties are a type of political party that emerged post-1970s and are characterised by heavy state financing and the diminished role of ideology as an organising principle. The cartel party thesis was developed by Richard Katz and Peter Mair who wrote that political parties have turned into "semi-state agencies", acting on behalf of the state rather than groups in society. The term 'cartel' refers to the way in which prominent parties in government make it difficult for new parties to enter, as such forming a cartel of established parties. As with catch-all parties, the role of members in cartel parties is largely insignificant as parties use the resources of the state to maintain their position within the political system.

Niche party

Niche parties are a type of political party that developed on the basis of the emergence of new cleavages and issues in politics, such as immigration and the environment. In contrast to mainstream or catch-all parties, niche parties articulate an often limited set of interests in a way that does not conform to the dominant economic left-right divide in politics, emphasising issues that do not attain prominence within the other parties. Further, niche parties do not respond to changes in public opinion to the extent that mainstream parties do. Examples of niche parties include Green parties and extreme nationalist parties, such as the Front National in France. However, over time these parties may lose some of their niche qualities, instead adopting those of mainstream parties, for example after entering government.

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