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Thursday, August 14, 2014

Computer network

Computer network

From Wikipedia, the free encyclopedia
   
A computer network or data network is a telecommunications network that allows computers to exchange data. In computer networks, networked computing devices pass data to each other along data connections. The connections (network links) between nodes are established using either cable media or wireless media. The best-known computer network is the Internet.
Network computer devices that originate, route and terminate the data are called network nodes.[1] Nodes can include hosts such as personal computers, phones, servers as well as networking hardware. Two such devices are said to be networked together when one device is able to exchange information with the other device, whether or not they have a direct connection to each other.
Computer networks support applications such as access to the World Wide Web, shared use of application and storage servers, printers, and fax machines, and use of email and instant messaging applications. Computer networks differ in the physical media used to transmit their signals, the communications protocols to organize network traffic, the network's size, topology and organizational intent.

History

Today, computer networks are the core of modern communication. All modern aspects of the public switched telephone network (PSTN) are computer-controlled. Telephony increasingly runs over the Internet Protocol, although not necessarily the public Internet. The scope of communication has increased significantly in the past decade. This boom in communications would not have been possible without the progressively advancing computer network. Computer networks, and the technologies that make communication between networked computers possible, continue to drive computer hardware, software, and peripherals industries. The expansion of related industries is mirrored by growth in the numbers and types of people using networks, from the researcher to the home user.

The following is a chronology of significant computer network developments:

Properties

Computer networking may be considered a branch of electrical engineering, telecommunications, computer science, information technology or computer engineering, since it relies upon the theoretical and practical application of the related disciplines.

A computer network facilitates interpersonal communications allowing people to communicate efficiently and easily via email, instant messaging, chat rooms, telephone, video telephone calls, and video conferencing. Providing access to information on shared storage devices is an important feature of many networks. A network allows sharing of files, data, and other types of information giving authorized users the ability to access information stored on other computers on the network. A network allows sharing of network and computing resources. Users may access and use resources provided by devices on the network, such as printing a document on a shared network printer. Distributed computing uses computing resources across a network to accomplish tasks. A computer network may be used by computer Crackers to deploy computer viruses or computer worms on devices connected to the network, or to prevent these devices from accessing the network (denial of service). A complex computer network may be difficult to set up. It may be costly to set up an effective computer network in a large organization.

Network packet

Most information in computer networks is carried in packets.

A network packet is a formatted unit of data (a list of bits or bytes) carried by a packet-switched network. Computer communications links that do not support packets, such as traditional point-to-point telecommunications links, simply transmit data as a bit stream. When data is formatted into packets, the bandwidth of the communication medium can be better shared among users than if the network were circuit switched.

A packet consists of two kinds of data: control information and user data (also known as payload). The control information provides data the network needs to deliver the user data, for example: source and destination network addresses, error detection codes, and sequencing information. Typically, control information is found in packet headers and trailers, with payload data in between.

Network topology

The physical layout of a network is usually less important than the topology that connects network nodes. Most diagrams that describe a physical network are therefore topological, rather than geographic. The symbols on these diagrams usually denote network links and network nodes.

Network links

The communication media used to link devices to form a computer network include electrical cable (HomePNA, power line communication, G.hn), optical fiber (fiber-optic communication), and radio waves (wireless networking). In the OSI model, these are defined at layers 1 and 2 — the physical layer and the data link layer.

A widely adopted family of communication media used in local area network (LAN) technology is collectively known as Ethernet. The media and protocol standards that enable communication between networked devices over Ethernet are defined by IEEE 802.3. Ethernet transmit data over both copper and fiber cables. Wireless LAN standards (e.g. those defined by IEEE 802.11) use radio waves, or others use infrared signals as a transmission medium. Power line communication uses a building's power cabling to transmit data.

Wired technologies

Fiber optic cables are used to transmit light from one computer/network node to another

The orders of the following wired technologies are, roughly, from slowest to fastest transmission speed.
  • Twisted pair wire is the most widely used medium for all telecommunication. Twisted-pair cabling consist of copper wires that are twisted into pairs. Ordinary telephone wires consist of two insulated copper wires twisted into pairs. Computer network cabling (wired Ethernet as defined by IEEE 802.3) consists of 4 pairs of copper cabling that can be utilized for both voice and data transmission. The use of two wires twisted together helps to reduce crosstalk and electromagnetic induction. The transmission speed ranges from 2 million bits per second to 10 billion bits per second. Twisted pair cabling comes in two forms: unshielded twisted pair (UTP) and shielded twisted-pair (STP). Each form comes in several category ratings, designed for use in various scenarios.
  • Coaxial cable is widely used for cable television systems, office buildings, and other work-sites for local area networks. The cables consist of copper or aluminum wire surrounded by an insulating layer (typically a flexible material with a high dielectric constant), which itself is surrounded by a conductive layer. The insulation helps minimize interference and distortion. Transmission speed ranges from 200 million bits per second to more than 500 million bits per second.
  • An optical fiber is a glass fiber. It carries pulses of light that represent data. Some advantages of optical fibers over metal wires are very low transmission loss and immunity from electrical interference. Optical fibers can simultaneously carry multiple wavelengths of light, which greatly increases the rate that data can be sent, and helps enable data rates of up to trillions of bits per second. Optic fibers can be used for long runs of cable carrying very high data rates, and are used for undersea cables to interconnect continents.
  • Price is a main factor distinguishing wired- and wireless-technology options in a business. Wireless options command a price premium that can make purchasing wired computers, printers and other devices a financial benefit. Before making the decision to purchase hard-wired technology products, a review of the restrictions and limitations of the selections is necessary. Business and employee needs may override any cost considerations.[5]

Wireless technologies[edit]

Computers are very often connected to networks using wireless links
  • Terrestrial microwave – Terrestrial microwave communication uses Earth-based transmitters and receivers resembling satellite dishes. Terrestrial microwaves are in the low-gigahertz range, which limits all communications to line-of-sight. Relay stations are spaced approximately 48 km (30 mi) apart.
  • Communications satellites – Satellites communicate via microwave radio waves, which are not deflected by the Earth's atmosphere. The satellites are stationed in space, typically in geosynchronous orbit 35,400 km (22,000 mi) above the equator. These Earth-orbiting systems are capable of receiving and relaying voice, data, and TV signals.
  • Cellular and PCS systems use several radio communications technologies. The systems divide the region covered into multiple geographic areas. Each area has a low-power transmitter or radio relay antenna device to relay calls from one area to the next area.
  • Radio and spread spectrum technologies – Wireless local area networks use a high-frequency radio technology similar to digital cellular and a low-frequency radio technology. Wireless LANs use spread spectrum technology to enable communication between multiple devices in a limited area. IEEE 802.11 defines a common flavor of open-standards wireless radio-wave technology known as Wifi.

Exotic technologies

There have been various attempts at transporting data over exotic media:
  • Extending the Internet to interplanetary dimensions via radio waves.[7]
Both cases have a large round-trip delay time, which gives slow two-way communication, but doesn't prevent sending large amounts of information.

Network nodes

Apart from the physical communications media described above, networks comprise additional basic system building blocks, such as network interface controller (NICs), repeaters, hubs, bridges, switches, routers, modems, and firewalls.

Network interfaces

An ATM network interface in the form of an accessory card. A lot of network interfaces are built-in.

A network interface controller (NIC) is computer hardware that provides a computer with the ability to access the transmission media, and has the ability to process low-level network information. For example the NIC may have a connector for accepting a cable, or an aerial for wireless transmission and reception, and the associated circuitry.

The NIC responds to traffic addressed to a network address for either the NIC or the computer as a whole.

In Ethernet networks, each network interface controller has a unique Media Access Control (MAC) address—usually stored in the controller's permanent memory. To avoid address conflicts between network devices, the Institute of Electrical and Electronics Engineers (IEEE) maintains and administers MAC address uniqueness. The size of an Ethernet MAC address is six octets. The three most significant octets are reserved to identify NIC manufacturers. These manufacturers, using only their assigned prefixes, uniquely assign the three least-significant octets of every Ethernet interface they produce.

Repeaters and hubs

A repeater is an electronic device that receives a network signal, cleans it of unnecessary noise, and regenerates it. The signal is retransmitted at a higher power level, or to the other side of an obstruction, so that the signal can cover longer distances without degradation. In most twisted pair
Ethernet configurations, repeaters are required for cable that runs longer than 100 meters. With fiber optics, repeaters can be tens or even hundreds of kilometers apart.

A repeater with multiple ports is known as a hub. Repeaters work on the physical layer of the OSI model. Repeaters require a small amount of time to regenerate the signal. This can cause a propagation delay that affects network performance. As a result, many network architectures limit the number of repeaters that can be used in a row, e.g., the Ethernet 5-4-3 rule.

Hubs have been mostly obsoleted by modern switches; but repeaters are used for long distance links, notably undersea cabling.

Bridges

A network bridge connects and filters traffic between two network segments at the data link layer (layer 2) of the OSI model to form a single network. This breaks the network's collision domain but maintains a unified broadcast domain. Network segmentation breaks down a large, congested network into an aggregation of smaller, more efficient networks.

Bridges come in three basic types:
  • Local bridges: Directly connect LANs
  • Remote bridges: Can be used to create a wide area network (WAN) link between LANs. Remote bridges, where the connecting link is slower than the end networks, largely have been replaced with routers.
  • Wireless bridges: Can be used to join LANs or connect remote devices to LANs.

Switches

A network switch is a device that forwards and filters OSI layer 2 datagrams between ports based on the MAC addresses in the packets.[8] A switch is distinct from a hub in that it only forwards the frames to the physical ports involved in the communication rather than all ports connected. It can be thought of as a multi-port bridge.[9] It learns to associate physical ports to MAC addresses by examining the source addresses of received frames. If an unknown destination is targeted, the switch broadcasts to all ports but the source. Switches normally have numerous ports, facilitating a star topology for devices, and cascading additional switches.

Multi-layer switches are capable of routing based on layer 3 addressing or additional logical levels. The term switch is often used loosely to include devices such as routers and bridges, as well as devices that may distribute traffic based on load or based on application content (e.g., a Web URL identifier).

Routers

A typical home or small office router showing the ADSL telephone line and Ethernet network cable connections

A router is an internetworking device that forwards packets between networks by processing the routing information included in the packet or datagram (Internet protocol information from layer 3).
The routing information is often processed in conjunction with the routing table (or forwarding table). A router uses its routing table to determine where to forward packets. (A destination in a routing table can include a "null" interface, also known as the "black hole" interface because data can go into it, however, no further processing is done for said data.)

Modems

Modems (MOdulator-DEModulator) are used to connect network nodes via wire not originally designed for digital network traffic, or for wireless. To do this one or more frequencies are modulated by the digital signal to produce an analog signal that can be tailored to give the required properties for transmission. Modems are commonly used for telephone lines, using a Digital Subscriber Line technology.

Firewalls

A firewall is a network device for controlling network security and access rules. Firewalls are typically configured to reject access requests from unrecognized sources while allowing actions from recognized ones. The vital role firewalls play in network security grows in parallel with the constant increase in cyber attacks.

Network structure

Network topology is the layout or organizational hierarchy of interconnected nodes of a computer network. Different network topologies can affect throughput, but reliability is often more critical. With many technologies, such as bus networks, a single failure can cause the network to fail entirely. In general the more interconnections there are, the more robust the network is; but the more expensive it is to install.

Common layouts

Common network topologies

Common layouts are:
  • A bus network: all nodes are connected to a common medium along this medium. This was the layout used in the original Ethernet, called 10BASE5 and 10BASE2.
  • A star network: all nodes are connected to a special central node. This is the typical layout found in a Wireless LAN, where each wireless client connects to the central Wireless access point.
  • A ring network: each node is connected to its left and right neighbour node, such that all nodes are connected and that each node can reach each other node by traversing nodes left- or rightwards. The Fiber Distributed Data Interface (FDDI) made use of such a topology.
  • A mesh network: each node is connected to an arbitrary number of neighbours in such a way that there is at least one traversal from any node to any other.
  • A fully connected network: each node is connected to every other node in the network.
  • A tree network: nodes are arranged hierarchically.
Note that the physical layout of the nodes in a network may not necessarily reflect the network topology. As an example, with FDDI, the network topology is a ring (actually two counter-rotating rings), but the physical topology is often a star, because all neighboring connections can be routed via a central physical location.

Overlay network

A sample overlay network

An overlay network is a virtual computer network that is built on top of another network. Nodes in the overlay network are connected by virtual or logical links. Each link corresponds to a path, perhaps through many physical links, in the underlying network. The topology of the overlay network may (and often does) differ from that of the underlying one. For example, many peer-to-peer networks are overlay networks. They are organized as nodes of a virtual system of links that run on top of the Internet.[10]

Overlay networks have been around since the invention of networking when computer systems were connected over telephone lines using modems, before any data network existed.

The most striking example of an overlay network is the Internet itself. The Internet itself was initially built as an overlay on the telephone network.[10] Even today, at the network layer, each node can reach any other by a direct connection to the desired IP address, thereby creating a fully connected network. The underlying network, however, is composed of a mesh-like interconnect of sub-networks of varying topologies (and technologies). Address resolution and routing are the means that allow mapping of a fully connected IP overlay network to its underlying network.

Another example of an overlay network is a distributed hash table, which maps keys to nodes in the network. In this case, the underlying network is an IP network, and the overlay network is a table (actually a map) indexed by keys.

Overlay networks have also been proposed as a way to improve Internet routing, such as through quality of service guarantees to achieve higher-quality streaming media. Previous proposals such as IntServ, DiffServ, and IP Multicast have not seen wide acceptance largely because they require modification of all routers in the network.[citation needed] On the other hand, an overlay network can be incrementally deployed on end-hosts running the overlay protocol software, without cooperation from Internet service providers. The overlay network has no control over how packets are routed in the underlying network between two overlay nodes, but it can control, for example, the sequence of overlay nodes that a message traverses before it reaches its destination.

For example, Akamai Technologies manages an overlay network that provides reliable, efficient content delivery (a kind of multicast). Academic research includes end system multicast,[11] resilient routing and quality of service studies, among others.

Communications protocols

Protocols in relation to the Internet layering scheme.
The TCP/IP model or Internet layering scheme and its relation to common protocols often layered on top of it.

A communications protocol is a set of rules for exchanging information over network links. In a protocol stack (also see the OSI model), each protocol leverages the services of the protocol below it. An important example of a protocol stack is HTTP running over TCP over IP over IEEE 802.11. (TCP and IP are members of the Internet Protocol Suite. IEEE 802.11 is a member of the Ethernet protocol suite.) This stack is used between the wireless router and the home user's personal computer when the user is surfing the web.

Whilst the use of protocol layering is today ubiquitous across the field of computer networking, it has been historically criticized by many researchers[12] for two principle reasons. Firstly, abstracting the protocol stack in this way may cause a higher layer to duplicate functionality of a lower layer, a prime example being error recovery on both a per-link basis and an end-to-end basis.[13] Secondly, it is common that a protocol implementation at one layer may require data, state or addressing information that is only present at another layer, thus defeating the point of separating the layers in the first place. For example, TCP uses the ECN field in the IPv4 header as an indication of congestion; IP is a network layer protocol whereas TCP is a transport layer protocol.
Communication protocols have various characteristics. They may be connection-oriented or connectionless, they may use circuit mode or packet switching, and they may use hierarchical addressing or flat addressing.

There are many communication protocols, a few of which are described below.

Ethernet

Ethernet is a family of protocols used in LANs, described by a set of standards together called IEEE 802 published by the Institute of Electrical and Electronics Engineers. It has a flat addressing scheme. It operates mostly at levels 1 and 2 of the OSI model. For home users today, the most well-known member of this protocol family is IEEE 802.11, otherwise known as Wireless LAN (WLAN). The complete IEEE 802 protocol suite provides a diverse set of networking capabilities. For example, MAC bridging (IEEE 802.1D) deals with the routing of Ethernet packets using a Spanning Tree Protocol, IEEE 802.1Q describes VLANs, and IEEE 802.1X defines a port-based Network Access Control protocol, which forms the basis for the authentication mechanisms used in VLANs (but it is also found in WLANs) – it is what the home user sees when the user has to enter a "wireless access key".

Internet Protocol Suite

The Internet Protocol Suite, also called TCP/IP, is the foundation of all modern networking. It offers connection-less as well as connection-oriented services over an inherently unreliable network traversed by data-gram transmission at the Internet protocol (IP) level. At its core, the protocol suite defines the addressing, identification, and routing specifications for Internet Protocol Version 4 (IPv4) and for IPv6, the next generation of the protocol with a much enlarged addressing capability.

SONET/SDH

Synchronous optical networking (SONET) and Synchronous Digital Hierarchy (SDH) are standardized multiplexing protocols that transfer multiple digital bit streams over optical fiber using lasers. They were originally designed to transport circuit mode communications from a variety of different sources, primarily to support real-time, uncompressed, circuit-switched voice encoded in PCM (Pulse-Code Modulation) format. However, due to its protocol neutrality and transport-oriented features, SONET/SDH also was the obvious choice for transporting Asynchronous Transfer Mode (ATM) frames.

Asynchronous Transfer Mode

Asynchronous Transfer Mode (ATM) is a switching technique for telecommunication networks. It uses asynchronous time-division multiplexing and encodes data into small, fixed-sized cells. This differs from other protocols such as the Internet Protocol Suite or Ethernet that use variable sized packets or frames. ATM has similarity with both circuit and packet switched networking. This makes it a good choice for a network that must handle both traditional high-throughput data traffic, and real-time, low-latency content such as voice and video. ATM uses a connection-oriented model in which a virtual circuit must be established between two endpoints before the actual data exchange begins.
While the role of ATM is diminishing in favor of next-generation networks, it still plays a role in the last mile, which is the connection between an Internet service provider and the home user. For an interesting write-up of the technologies involved, including the deep stacking of communications protocols used, see.[14]

Geographic scale

A network can be characterized by its physical capacity or its organizational purpose. Use of the network, including user authorization and access rights, differ accordingly.
Personal area network
A personal area network (PAN) is a computer network used for communication among computer and different information technological devices close to one person. Some examples of devices that are used in a PAN are personal computers, printers, fax machines, telephones, PDAs, scanners, and even video game consoles. A PAN may include wired and wireless devices. The reach of a PAN typically extends to 10 meters.[15] A wired PAN is usually constructed with USB and FireWire connections while technologies such as Bluetooth and infrared communication typically form a wireless PAN.
Local area network
A local area network (LAN) is a network that connects computers and devices in a limited geographical area such as a home, school, office building, or closely positioned group of buildings. Each computer or device on the network is a node. Wired LANs are most likely based on Ethernet technology. Newer standards such as ITU-T G.hn also provide a way to create a wired LAN using existing wiring, such as coaxial cables, telephone lines, and power lines.[16]

A LAN is depicted in the accompanying diagram. All interconnected devices use the network layer (layer 3) to handle multiple subnets (represented by different colors). Those inside the library have 10/100 Mbit/s Ethernet connections to the user device and a Gigabit Ethernet connection to the central router. They could be called Layer 3 switches, because they only have Ethernet interfaces and support the Internet Protocol. It might be more correct to call them access routers, where the router at the top is a distribution router that connects to the Internet and to the academic networks' customer access routers.

The defining characteristics of a LAN, in contrast to a wide area network (WAN), include higher data transfer rates, limited geographic range, and lack of reliance on leased lines to provide connectivity. Current Ethernet or other IEEE 802.3 LAN technologies operate at data transfer rates up to 10 Gbit/s. The IEEE investigates the standardization of 40 and 100 Gbit/s rates.[17] A LAN can be connected to a WAN using a router.
Home area network
A home area network (HAN) is a residential LAN used for communication between digital devices typically deployed in the home, usually a small number of personal computers and accessories, such as printers and mobile computing devices. An important function is the sharing of Internet access, often a broadband service through a cable TV or digital subscriber line (DSL) provider.
Storage area network
A storage area network (SAN) is a dedicated network that provides access to consolidated, block level data storage. SANs are primarily used to make storage devices, such as disk arrays, tape libraries, and optical jukeboxes, accessible to servers so that the devices appear like locally attached devices to the operating system. A SAN typically has its own network of storage devices that are generally not accessible through the local area network by other devices. The cost and complexity of SANs dropped in the early 2000s to levels allowing wider adoption across both enterprise and small to medium sized business environments.
Campus area network
A campus area network (CAN) is made up of an interconnection of LANs within a limited geographical area. The networking equipment (switches, routers) and transmission media (optical fiber, copper plant, Cat5 cabling, etc.) are almost entirely owned by the campus tenant / owner (an enterprise, university, government, etc.).

For example, a university campus network is likely to link a variety of campus buildings to connect academic colleges or departments, the library, and student residence halls.
Backbone network
A backbone network is part of a computer network infrastructure that provides a path for the exchange of information between different LANs or sub-networks. A backbone can tie together diverse networks within the same building, across different buildings, or over a wide area.

For example, a large company might implement a backbone network to connect departments that are located around the world. The equipment that ties together the departmental networks constitutes the network backbone. When designing a network backbone, network performance and network congestion are critical factors to take into account. Normally, the backbone network's capacity is greater than that of the individual networks connected to it.

Another example of a backbone network is the Internet backbone, which is the set of wide area networks (WANs) and core routers that tie together all networks connected to the Internet.
Metropolitan area network
A Metropolitan area network (MAN) is a large computer network that usually spans a city or a large campus.
Wide area network
A wide area network (WAN) is a computer network that covers a large geographic area such as a city, country, or spans even intercontinental distances. A WAN uses a communications channel that combines many types of media such as telephone lines, cables, and air waves. A WAN often makes use of transmission facilities provided by common carriers, such as telephone companies. WAN technologies generally function at the lower three layers of the OSI reference model: the physical layer, the data link layer, and the network layer.
Enterprise private network
An enterprise private network is a network that a single organization builds to interconnect its office locations (e.g., production sites, head offices, remote offices, shops) so they can share computer resources.
Virtual private network
A virtual private network (VPN) is an overlay network in which some of the links between nodes are carried by open connections or virtual circuits in some larger network (e.g., the Internet) instead of by physical wires. The data link layer protocols of the virtual network are said to be tunneled through the larger network when this is the case. One common application is secure communications through the public Internet, but a VPN need not have explicit security features, such as authentication or content encryption. VPNs, for example, can be used to separate the traffic of different user communities over an underlying network with strong security features.

VPN may have best-effort performance, or may have a defined service level agreement (SLA) between the VPN customer and the VPN service provider. Generally, a VPN has a topology more complex than point-to-point.
Global area network
A global area network (GAN) is a network used for supporting mobile across an arbitrary number of wireless LANs, satellite coverage areas, etc. The key challenge in mobile communications is handing off user communications from one local coverage area to the next. In IEEE Project 802, this involves a succession of terrestrial wireless LANs.[18]

Organizational scope

Networks are typically managed by the organizations that own them. Private enterprise networks may use a combination of intranets and extranets. They may also provide network access to the Internet, which has no single owner and permits virtually unlimited global connectivity.

Intranets

An intranet is a set of networks that are under the control of a single administrative entity. The intranet uses the IP protocol and IP-based tools such as web browsers and file transfer applications. The administrative entity limits use of the intranet to its authorized users. Most commonly, an intranet is the internal LAN of an organization. A large intranet typically has at least one web server to provide users with organizational information. An intranet is also anything behind the router on a local area network.

Extranet

An extranet is a network that is also under the administrative control of a single organization, but supports a limited connection to a specific external network. For example, an organization may provide access to some aspects of its intranet to share data with its business partners or customers.
These other entities are not necessarily trusted from a security standpoint. Network connection to an extranet is often, but not always, implemented via WAN technology.

Internetwork

An internetwork is the connection of multiple computer networks via a common routing technology using routers.

Internet

Partial map of the Internet based on the January 15, 2005 data found on opte.org. Each line is drawn between two nodes, representing two IP addresses. The length of the lines are indicative of the delay between those two nodes. This graph represents less than 30% of the Class C networks reachable.

The Internet is the largest example of an internetwork. It is a global system of interconnected governmental, academic, corporate, public, and private computer networks. It is based on the networking technologies of the Internet Protocol Suite. It is the successor of the Advanced Research Projects Agency Network (ARPANET) developed by DARPA of the United States Department of Defense. The Internet is also the communications backbone underlying the World Wide Web (WWW).

Participants in the Internet use a diverse array of methods of several hundred documented, and often standardized, protocols compatible with the Internet Protocol Suite and an addressing system (IP addresses) administered by the Internet Assigned Numbers Authority and address registries. Service providers and large enterprises exchange information about the reachability of their address spaces through the Border Gateway Protocol (BGP), forming a redundant worldwide mesh of transmission paths.

Darknet

A Darknet is an overlay network, typically running on the internet, that is only accessible through specialized software. A darknet is an anonymizing network where connections are made only between trusted peers — sometimes called "friends" (F2F)[19] — using non-standard protocols and ports.

Darknets are distinct from other distributed peer-to-peer networks as sharing is anonymous (that is, IP addresses are not publicly shared), and therefore users can communicate with little fear of governmental or corporate interference.[20]

Routing

Routing calculates good paths through a network for information to take. For example from node 1 to node 6 the best routes are likely to be 1-8-7-6 or 1-8-10-6, as this has the thickest routes.

Routing is the process of selecting network paths to carry network traffic. Routing is performed for many kinds of networks, including circuit switching networks and packet switched networks.
In packet switched networks, routing directs packet forwarding (the transit of logically addressed network packets from their source toward their ultimate destination) through intermediate nodes. Intermediate nodes are typically network hardware devices such as routers, bridges, gateways, firewalls, or switches. General-purpose computers can also forward packets and perform routing, though they are not specialized hardware and may suffer from limited performance. The routing process usually directs forwarding on the basis of routing tables, which maintain a record of the routes to various network destinations. Thus, constructing routing tables, which are held in the router's memory, is very important for efficient routing. Most routing algorithms use only one network path at a time. Multipath routing techniques enable the use of multiple alternative paths.

There are usually multiple routes that can be taken, and to choose between them, different elements can be considered to decide which routes get installed into the routing table, such as (sorted by priority):
  1. Prefix-Length: where longer subnet masks are preferred (independent if it is within a routing protocol or over different routing protocol)
  2. Metric: where a lower metric/cost is preferred (only valid within one and the same routing protocol)
  3. Administrative distance: where a lower distance is preferred (only valid between different routing protocols)
Routing, in a more narrow sense of the term, is often contrasted with bridging in its assumption that network addresses are structured and that similar addresses imply proximity within the network. Structured addresses allow a single routing table entry to represent the route to a group of devices. In large networks, structured addressing (routing, in the narrow sense) outperforms unstructured addressing (bridging). Routing has become the dominant form of addressing on the Internet. Bridging is still widely used within localized environments.

Network service

Network services are applications hosted by servers on a computer network, to provide some functionality for members or users of the network, or to help the network itself to operate.

The World Wide Web, E-mail,[21] printing and network file sharing are examples of well-known network services. Network services such as DNS (Domain Name System) give names for IP and MAC addresses (people remember names like “nm.lan” better than numbers like “210.121.67.18”),[22] and DHCP to ensure that the equipment on the network has a valid IP address.[23]
Services are usually based on a service protocol that defines the format and sequencing of messages between clients and servers of that network service.

Network performance

Quality of service

Depending on the installation requirements, network performance is usually measured by the quality of service of a telecommunications product. The parameters that affect this typically can include throughput, jitter, bit error rate and latency.

The following list gives examples of network performance measures for a circuit-switched network and one type of packet-switched network, viz. ATM:
  • Circuit-switched networks: In circuit switched networks, network performance is synonymous with the grade of service. The number of rejected calls is a measure of how well the network is performing under heavy traffic loads.[24] Other types of performance measures can include the level of noise and echo.
There are many ways to measure the performance of a network, as each network is different in nature and design. Performance can also be modelled instead of measured. For example, state transition diagrams are often used to model queuing performance in a circuit-switched network. The network planner uses these diagrams to analyze how the network performs in each state, ensuring that the network is optimally designed.[26]

Network congestion

Network congestion occurs when a link or node is carrying so much data that its quality of service deteriorates. Typical effects include queueing delay, packet loss or the blocking of new connections. A consequence of these latter two is that incremental increases in offered load lead either only to small increase in network throughput, or to an actual reduction in network throughput.

Network protocols that use aggressive retransmissions to compensate for packet loss tend to keep systems in a state of network congestion—even after the initial load is reduced to a level that would not normally induce network congestion. Thus, networks using these protocols can exhibit two stable states under the same level of load. The stable state with low throughput is known as congestive collapse.

Modern networks use congestion control and congestion avoidance techniques to try to avoid congestion collapse. These include: exponential backoff in protocols such as 802.11's CSMA/CA and the original Ethernet, window reduction in TCP, and fair queueing in devices such as routers.
Another method to avoid the negative effects of network congestion is implementing priority schemes, so that some packets are transmitted with higher priority than others. Priority schemes do not solve network congestion by themselves, but they help to alleviate the effects of congestion for some services. An example of this is 802.1p. A third method to avoid network congestion is the explicit allocation of network resources to specific flows. One example of this is the use of Contention-Free Transmission Opportunities (CFTXOPs) in the ITU-T G.hn standard, which provides high-speed (up to 1 Gbit/s) Local area networking over existing home wires (power lines, phone lines and coaxial cables).

For the Internet RFC 2914 addresses the subject of congestion control in detail.

Network resilience

Network resilience is "the ability to provide and maintain an acceptable level of service in the face of faults and challenges to normal operation.”[27]

Security

Network security

Network security consists of provisions and policies adopted by the network administrator to prevent and monitor unauthorized access, misuse, modification, or denial of the computer network and its network-accessible resources.[28] Network security is the authorization of access to data in a network, which is controlled by the network administrator. Users are assigned an ID and password that allows them access to information and programs within their authority. Network security is used on a variety of computer networks, both public and private, to secure daily transactions and communications among businesses, government agencies and individuals.

Network surveillance

Network surveillance is the monitoring of data being transferred over computer networks such as the Internet. The monitoring is often done surreptitiously and may be done by or at the behest of governments, by corporations, criminal organizations, or individuals. It may or may not be legal and may or may not require authorization from a court or other independent agency.

Computer and network surveillance programs are widespread today, and almost all Internet traffic is or could potentially be monitored for clues to illegal activity.

Surveillance is very useful to governments and law enforcement to maintain social control, recognize and monitor threats, and prevent/investigate criminal activity. With the advent of programs such as the Total Information Awareness program, technologies such as high speed surveillance computers and biometrics software, and laws such as the Communications Assistance For Law Enforcement Act, governments now possess an unprecedented ability to monitor the activities of citizens.[29]

However, many civil rights and privacy groups—such as Reporters Without Borders, the Electronic Frontier Foundation, and the American Civil Liberties Union—have expressed concern that increasing surveillance of citizens may lead to a mass surveillance society, with limited political and personal freedoms. Fears such as this have led to numerous lawsuits such as Hepting v. AT&T.[29][30] The hacktivist group Anonymous has hacked into government websites in protest of what it considers "draconian surveillance".[31][32]

End to end encryption

End-to-end encryption (E2EE) is a digital communications paradigm of uninterrupted protection of data traveling between two communicating parties. It involves the originating party encrypting data so only the intended recipient can decrypt it, with no dependency on third parties. End-to-end encryption prevents intermediaries, such as Internet providers or application service providers, from discovering or tampering with communications. End-to-end encryption generally protects both confidentiality and integrity.

Examples of end-to-end encryption include PGP for email, OTR for instant messaging, ZRTP for telephony, and TETRA for radio.

Typical server-based communications systems do not include end-to-end encryption. These systems can only guarantee protection of communications between clients and servers, not between the communicating parties themselves. Examples of non-E2EE systems are Google Talk, Yahoo Messenger, Facebook, and Dropbox. Some such systems, for example LavaBit and SecretInk, have even described themselves as offering "end-to-end" encryption when they do not. Some systems that normally offer end-to-end encryption have turned out to contain a back door that subverts negotiation of the encryption key between the communicating parties, for example Skype.

The end-to-end encryption paradigm does not directly address risks at the communications endpoints themselves, such as the technical exploitation of clients, poor quality random number generators, or key escrow. E2EE also does not address traffic analysis, which relates to things such as the identities of the end points and the times and quantities of messages that are sent.

Views of networks

Users and network administrators typically have different views of their networks. Users can share printers and some servers from a workgroup, which usually means they are in the same geographic location and are on the same LAN, whereas a Network Administrator is responsible to keep that network up and running. A community of interest has less of a connection of being in a local area, and should be thought of as a set of arbitrarily located users who share a set of servers, and possibly also communicate via peer-to-peer technologies.

Network administrators can see networks from both physical and logical perspectives. The physical perspective involves geographic locations, physical cabling, and the network elements (e.g., routers, bridges and application layer gateways) that interconnect the physical media. Logical networks, called, in the TCP/IP architecture, subnets, map onto one or more physical media. For example, a common practice in a campus of buildings is to make a set of LAN cables in each building appear to be a common subnet, using virtual LAN (VLAN) technology.

Both users and administrators are aware, to varying extents, of the trust and scope characteristics of a network. Again using TCP/IP architectural terminology, an intranet is a community of interest under private administration usually by an enterprise, and is only accessible by authorized users (e.g. employees).[33] Intranets do not have to be connected to the Internet, but generally have a limited connection. An extranet is an extension of an intranet that allows secure communications to users outside of the intranet (e.g. business partners, customers).[33]

Unofficially, the Internet is the set of users, enterprises, and content providers that are interconnected by Internet Service Providers (ISP). From an engineering viewpoint, the Internet is the set of subnets, and aggregates of subnets, which share the registered IP address space and exchange information about the reachability of those IP addresses using the Border Gateway Protocol. Typically, the human-readable names of servers are translated to IP addresses, transparently to users, via the directory function of the Domain Name System (DNS).

Over the Internet, there can be business-to-business (B2B), business-to-consumer (B2C) and consumer-to-consumer (C2C) communications. When money or sensitive information is exchanged, the communications are apt to be protected by some form of communications security mechanism. Intranets and extranets can be securely superimposed onto the Internet, without any access by general Internet users and administrators, using secure Virtual Private Network (VPN) technology.

Human brain

Human brain

From Wikipedia, the free encyclopedia
 
Human brain
Skull and brain normal human.svg
Human brain and skull
Cerebral lobes.png
Cerebral lobes: the frontal lobe (pink), parietal lobe (green) and occipital lobe (blue)
LatinCerebrum
Gray'sp.736
SystemCentral nervous system
ArteryInternal carotid arteries, vertebral arteries
VeinInternal jugular vein, cerebral veins, external veins, basal vein, terminal vein, choroid vein, cerebellar veins
PrecursorNeural tube
Anatomical terminology

The human brain has the same general structure as the brains of other mammals, but has a more developed cortex than any other. Large animals such as whales and elephants have larger brains in absolute terms, but when measured using the encephalization quotient which compensates for body size, the human brain is almost twice as large as the brain of the bottlenose dolphin, and three times as large as the brain of a chimpanzee. Much of the expansion comes from the part of the brain called the cerebral cortex, especially the frontal lobes, which are associated with executive functions such as self-control, planning, reasoning, and abstract thought. The portion of the cerebral cortex devoted to vision is also greatly enlarged in humans.

The human cerebral cortex is a thick layer of neural tissue that covers most of the brain. This layer is folded in a way that increases the amount of surface that can fit into the volume available. The pattern of folds is similar across individuals, although there are many small variations. The cortex is divided into four "lobes", called the frontal lobe, parietal lobe, temporal lobe, and occipital lobe. (Some classification systems also include a limbic lobe and treat the insular cortex as a lobe.) Within each lobe are numerous cortical areas, each associated with a particular function such as vision, motor control, language, etc. The left and right sides of the cortex are broadly similar in shape, and most cortical areas are replicated on both sides. Some areas, though, show strong lateralization, particularly areas that are involved in language. In most people, the left hemisphere is "dominant" for language, with the right hemisphere playing only a minor role. There are other functions, such as spatiotemporal reasoning, for which the right hemisphere is usually dominant.

Despite being protected by the thick bones of the skull, suspended in cerebrospinal fluid, and isolated from the bloodstream by the blood–brain barrier, the human brain is susceptible to damage and disease. The most common forms of physical damage are closed head injuries such as a blow to the head, a stroke, or poisoning by a variety of chemicals that can act as neurotoxins. Infection of the brain, though serious, is rare due to the biological barriers which protect it. The human brain is also susceptible to degenerative disorders, such as Parkinson's disease, multiple sclerosis, and Alzheimer's disease. A number of psychiatric conditions, such as schizophrenia and depression, are thought to be associated with brain dysfunctions, although the nature of such brain anomalies is not well understood.

Scientifically, the techniques that are used to study the human brain differ in important ways from those that are used to study the brains of other mammals. On the one hand, invasive techniques such as inserting electrodes into the brain, or disabling parts of the brain in order to examine the effect on behavior, are used with non-human species, but for ethical reasons, are generally not performed with humans. On the other hand, humans are the only subjects who can respond to complex verbal instructions. Thus, it is often possible to use non-invasive techniques such as functional neuroimaging or EEG recording more productively with humans than with non-humans. Furthermore, some of the most important topics, such as language, can hardly be studied at all except in humans. In many cases, human and non-human studies form essential complements to each other. Individual brain cells (except where tissue samples are taken for biopsy for suspected brain tumors) can only be studied in non-humans; complex cognitive tasks can only be studied in humans. Combining the two sources of information to yield a complete functional understanding of the human brain is an ongoing challenge for neuroscience.

Structure

Drawing of the human brain, showing several important structures
Human brain viewed from below

The adult human brain weighs on average about 1.5 kg (3.3 lb)[1] with a volume of around 1130 cubic centimetres (cm3) in women and 1260 cm3 in men, although there is substantial individual variation.[2] Neurological differences between the sexes have not been shown to correlate in any simple way with IQ or other measures of cognitive performance.[3] The human brain is composed of neurons, glial cells, and blood vessels. The number of neurons, according to array tomography, has been shown to be about 86 billion neurons in the human brain with a roughly equal number of non-neuronal cells called glia.[4]

The cerebral hemispheres (the cerebrum) form the largest part of the human brain and are situated above other brain structures. They are covered with a cortical layer (the cerebral cortex) which has a convoluted topography.[5] Underneath the cerebrum lies the brainstem, resembling a stalk on which the cerebrum is attached. At the rear of the brain, beneath the cerebrum and behind the brainstem, is the cerebellum, a structure with a horizontally furrowed surface, the cerebellar cortex, that makes it look different from any other brain area. The same structures are present in other mammals, although they vary considerably in relative size. As a rule, the smaller the cerebrum, the less convoluted the cortex. The cortex of a rat or mouse is almost perfectly smooth. The cortex of a dolphin or whale, on the other hand, is more convoluted than the cortex of a human.

The living brain is very soft, having a consistency similar to soft gelatin or soft tofu. Despite being referred to as grey matter, the live cortex is pinkish-beige in color and slightly off-white in the interior.

General features

Human brain viewed through a mid-line incision

The human brain has many properties that are common to all vertebrate brains, including a basic division into three parts called the forebrain, midbrain, and hindbrain, each with fluid-filled ventricles at their core, and a set of generic vertebrate brain structures including the medulla oblongata, pons, cerebellum, optic tectum, thalamus, hypothalamus, basal ganglia, olfactory bulb, and many others.
As a mammalian brain, the human brain has special features that are common to all mammalian brains, most notably a six-layered cerebral cortex and a set of structures associated with it, including the hippocampus and amygdala. All vertebrates have a forebrain whose upper surface is covered with a layer of neural tissue called the pallium, but in all except mammals the pallium has a relatively simple three-layered cell structure. In mammals it has a much more complex six-layered cell structure, and is given a different name, the cerebral cortex. The hippocampus and amygdala also originate from the pallium, but are much more complex in mammals than in other vertebrates.
As a primate brain, the human brain has a much larger cerebral cortex, in proportion to body size, than most mammals, and a very highly developed visual system. The shape of the brain within the skull is also altered somewhat as a consequence of the upright position in which primates hold their heads.

As a hominid brain, the human brain is substantially enlarged even in comparison to the brain of a typical monkey. The sequence of evolution from Australopithecus (four million years ago) to Homo sapiens (modern man) was marked by a steady increase in brain size, particularly in the frontal lobes, which are associated with a variety of high-level cognitive functions.

Humans and other primates have some differences in gene sequence, and genes are differentially expressed in many brain regions. The functional differences between the human brain and the brains of other animals also arise from many gene–environment interactions.[6]

Cerebral cortex

Bisection of the head of an adult female, showing the cerebral cortex, with its extensive folding, and the underlying white matter[7]

The dominant feature of the human brain is corticalization. The cerebral cortex in humans is so large that it overshadows every other part of the brain. A few subcortical structures show alterations reflecting this trend. The cerebellum, for example, has a medial zone connected mainly to subcortical motor areas, and a lateral zone connected primarily to the cortex. In humans the lateral zone takes up a much larger fraction of the cerebellum than in most other mammalian species. Corticalization is reflected in function as well as structure. In a rat, surgical removal of the entire cerebral cortex leaves an animal that is still capable of walking around and interacting with the environment.[8] In a human, comparable cerebral cortex damage produces a permanent state of coma. The amount of association cortex, relative to the other two categories, increases dramatically as one goes from simpler mammals, such as the rat and the cat, to more complex ones, such as the chimpanzee and the human.[9]

The cerebral cortex is essentially a sheet of neural tissue, folded in a way that allows a large surface area to fit within the confines of the skull. When unfolded, each cerebral hemisphere has a total surface area of about 1.3 square feet (0.12 m2).[10] Each cortical ridge is called a gyrus, and each groove or fissure separating one gyrus from another is called a sulcus.

Cortical divisions

Four lobes

The four lobes of the cerebral cortex

The cerebral cortex is nearly symmetrical with left and right hemispheres that are approximate mirror images of each other. Each hemisphere is conventionally divided into four "lobes", the frontal lobe, parietal lobe, occipital lobe, and temporal lobe. With one exception, this division into lobes does not derive from the structure of the cortex itself, though: the lobes are named after the bones of the skull that overlie them, the frontal bone, parietal bone, temporal bone, and occipital bone. The borders between lobes lie beneath the sutures that link the skull bones together. The exception is the border between the frontal and parietal lobes, which lies behind the corresponding suture; instead it follows the anatomical boundary of the central sulcus, a deep fold in the brain's structure where the primary somatosensory cortex and primary motor cortex meet.

Because of the arbitrary way most of the borders between lobes are demarcated, they have little functional significance. With the exception of the occipital lobe, a small area that is entirely dedicated to vision, each of the lobes contains a variety of brain areas that have minimal functional relationship. The parietal lobe, for example, contains areas involved in somatosensation, hearing, language, attention, and spatial cognition. In spite of this heterogeneity, the division into lobes is convenient for reference. The main functions of the frontal lobe are to control attention, abstract thinking, behavior, problem solving tasks, and physical reactions and personality.[11] The occipital lobe is the smallest lobe; its main functions are visual reception, visual-spatial processing, movement, and color recognition.[12] The temporal lobe controls auditory and visual memories, language, and some hearing and speech.[11]

Major sulci and gyri

Major gyri and sulci on the lateral surface of the cortex
Lateral surface of the cerebral cortex
Medial surface of the cerebral cortex

Although there are enough variations in the shape and placement of gyri and sulci (cortical folds) to make every brain unique, most human brains show sufficiently consistent patterns of folding that allow them to be named. Many of the gyri and sulci are named according to the location on the lobes or other major folds on the cortex. These include:

Functional divisions

Researchers who study the functions of the cortex divide it into three functional categories of regions. One consists of the primary sensory areas, which receive signals from the sensory nerves and tracts by way of relay nuclei in the thalamus. Primary sensory areas include the visual area of the occipital lobe, the auditory area in parts of the temporal lobe and insular cortex, and the somatosensory cortex in the parietal lobe. A second category is the primary motor cortex, which sends axons down to motor neurons in the brainstem and spinal cord.[13] This area occupies the rear portion of the frontal lobe, directly in front of the somatosensory area. The third category consists of the remaining parts of the cortex, which are called the association areas. These areas receive input from the sensory areas and lower parts of the brain and are involved in the complex processes of perception, thought, and decision-making.[14]

Cytoarchitecture

Brodmann's classification of areas of the cortex

Different parts of the cerebral cortex are involved in different cognitive and behavioral functions. The differences show up in a number of ways: the effects of localized brain damage, regional activity patterns exposed when the brain is examined using functional imaging techniques, connectivity with subcortical areas, and regional differences in the cellular architecture of the cortex. Neuroscientists describe most of the cortex—the part they call the neocortex—as having six layers, but not all layers are apparent in all areas, and even when a layer is present, its thickness and cellular organization may vary. Scientists have constructed maps of cortical areas on the basis of variations in the appearance of the layers as seen with a microscope. One of the most widely used schemes came from Korbinian Brodmann, who split the cortex into 51 different areas and assigned each a number (many of these Brodmann areas have since been subdivided). For example, Brodmann area 1 is the primary somatosensory cortex, Brodmann area 17 is the primary visual cortex, and Brodmann area 25 is the anterior cingulate cortex.[15]

Topography

Topography of the primary motor cortex, showing which body part is controlled by each zone

Many of the brain areas Brodmann defined have their own complex internal structures. In a number of cases, brain areas are organized into "topographic maps", where adjoining bits of the cortex correspond to adjoining parts of the body, or of some more abstract entity. A simple example of this type of correspondence is the primary motor cortex, a strip of tissue running along the anterior edge of the central sulcus, shown in the image to the right. Motor areas innervating each part of the body arise from a distinct zone, with neighboring body parts represented by neighboring zones. Electrical stimulation of the cortex at any point causes a muscle-contraction in the represented body part. This "somatotopic" representation is not evenly distributed, however. The head, for example, is represented by a region about three times as large as the zone for the entire back and trunk. The size of any zone correlates to the precision of motor control and sensory discrimination possible.= The areas for the lips, fingers, and tongue are particularly large, considering the proportional size of their represented body parts.

In visual areas, the maps are retinotopic—that is, they reflect the topography of the retina, the layer of light-activated neurons lining the back of the eye. In this case too the representation is uneven: the fovea—the area at the center of the visual field—is greatly overrepresented compared to the periphery. The visual circuitry in the human cerebral cortex contains several dozen distinct retinotopic maps, each devoted to analyzing the visual input stream in a particular way. The primary visual cortex (Brodmann area 17), which is the main recipient of direct input from the visual part of the thalamus, contains many neurons that are most easily activated by edges with a particular orientation moving across a particular point in the visual field. Visual areas farther downstream extract features such as color, motion, and shape.

In auditory areas, the primary map is tonotopic. Sounds are parsed according to frequency (i.e., high pitch vs. low pitch) by subcortical auditory areas, and this parsing is reflected by the primary auditory zone of the cortex. As with the visual system, there are a number of tonotopic cortical maps, each devoted to analyzing sound in a particular way.

Within a topographic map there can sometimes be finer levels of spatial structure. In the primary visual cortex, for example, where the main organization is retinotopic and the main responses are to moving edges, cells that respond to different edge-orientations are spatially segregated from one another.

Cognition

Understanding the relationship between the brain and the mind is a great challenge.[16] It is very difficult to imagine how mental entities such as thoughts and emotions could be implemented by physical entities such as neurons and synapses, or by any other type of mechanism. The difficulty was expressed by Gottfried Leibniz in an analogy known as Leibniz's Mill:
One is obliged to admit that perception and what depends upon it is inexplicable on mechanical principles, that is, by figures and motions. In imagining that there is a machine whose construction would enable it to think, to sense, and to have perception, one could conceive it enlarged while retaining the same proportions, so that one could enter into it, just like into a windmill. Supposing this, one should, when visiting within it, find only parts pushing one another, and never anything by which to explain a perception.
— Leibniz, Monadology[17]
Incredulity about the possibility of a mechanistic explanation of thought drove René Descartes, and most of humankind along with him, to dualism: the belief that the mind exists independently of the brain.[18] There has always, however been a strong argument in the opposite direction. There is overwhelming evidence that physical manipulations of, or damage to, the brain (for example by drugs or diseases, respectively) can affect the mind in potent and intimate ways.[19] For example, a person suffering from Alzheimer's disease—a condition that causes physical damage to the brain—also experiences a compromised "mind". Similarly, someone who has taken a psychedelic drug may temporarily lose their sense of personal identity (ego death) or experience profound changes to their perception and thought process. In this line of thinking, a large body of empirical evidence for a close relationship between brain activity and mind activity has led most neuroscientists to be materialists or physicalists, believing that mental phenomena are ultimately reducible to physical phenomena.[20]

Lateralization

 
Routing of neural signals from the two eyes to the brain

Each hemisphere of the brain interacts primarily with one half of the body, but for reasons that are unclear, the connections are crossed: the left side of the brain interacts with the right side of the body, and vice versa.[citation needed] Motor connections from the brain to the spinal cord, and sensory connections from the spinal cord to the brain, both cross the midline at the level of the brainstem.
Visual input follows a more complex rule: the optic nerves from the two eyes come together at a point called the optic chiasm, and half of the fibers from each nerve split off to join the other. The result is that connections from the left half of the retina, in both eyes, go to the left side of the brain, whereas connections from the right half of the retina go to the right side of the brain. Because each half of the retina receives light coming from the opposite half of the visual field, the functional consequence is that visual input from the left side of the world goes to the right side of the brain, and vice versa. Thus, the right side of the brain receives somatosensory input from the left side of the body, and visual input from the left side of the visual field—an arrangement that presumably is helpful for visuomotor coordination.
The corpus callosum, a nerve bundle connecting the two cerebral hemispheres, with the lateral ventricles directly below

The two cerebral hemispheres are connected by a very large nerve bundle (the largest white matter structure in the brain) called the corpus callosum, which crosses the midline above the level of the thalamus.[21] There are also two much smaller connections, the anterior commissure and hippocampal commissure, as well as many subcortical connections that cross the midline. The corpus callosum is the main avenue of communication between the two hemispheres, though. It connects each point on the cortex to the mirror-image point in the opposite hemisphere, and also connects to functionally related points in different cortical areas.

In most respects, the left and right sides of the brain are symmetrical in terms of function. For example, the counterpart of the left-hemisphere motor area controlling the right hand is the right-hemisphere area controlling the left hand. There are, however, several very important exceptions, involving language and spatial cognition. In most people, the left hemisphere is "dominant" for language: a stroke that damages a key language area in the left hemisphere can leave the victim unable to speak or understand, whereas equivalent damage to the right hemisphere would cause only minor impairment to language skills.

A substantial part of our current understanding of the interactions between the two hemispheres has come from the study of "split-brain patients"—people who underwent surgical transection of the corpus callosum in an attempt to reduce the severity of epileptic seizures. These patients do not show unusual behavior that is immediately obvious, but in some cases can behave almost like two different people in the same body, with the right hand taking an action and then the left hand undoing it. Most of these patients, when briefly shown a picture on the right side of the point of visual fixation, are able to describe it verbally, but when the picture is shown on the left, are unable to describe it, but may be able to give an indication with the left hand of the nature of the object shown.

Development

During the first 3 weeks of gestation, the human embryo's ectoderm forms a thickened strip called the neural plate. The neural plate then folds and closes to form the neural tube. This tube flexes as it grows, forming the crescent-shaped cerebral hemispheres at the head, and the cerebellum and pons towards the tail.
Brain of human embryo at 4.5 weeks, showing interior of forebrain 
Brain interior at 5 weeks 
Brain viewed at midline at 3 months 

Evolution

 
A reconstruction of Homo habilis

In the course of evolution of the Homininae, the human brain has grown in volume from about 600 cm3 in Homo habilis to about 1500 cm3 in Homo sapiens neanderthalensis. Subsequently, there has been a shrinking over the past 28,000 years. The male brain has decreased from 1,500 cm3 to 1,350 cm3 while the female brain has shrunk by the same relative proportion.[22] For comparison, Homo erectus, a relative of humans, had a brain size of 1,100 cm3. However, the little Homo floresiensis, with a brain size of 380 cm3, a third of that of their proposed ancestor H. erectus, used fire, hunted, and made stone tools at least as sophisticated as those of H. erectus.[23] In spite of significant changes in social capacity, there has been very little change in brain size from Neanderthals to the present day.[24] "As large as you need and as small as you can" has been said to summarize the opposite evolutionary constraints on human brain size.[25][26]

Studies tend to indicate small to moderate correlations (averaging around 0.3 to 0.4) between brain volume and IQ. The most consistent associations are observed within the frontal, temporal, and parietal lobes, the hippocampi, and the cerebellum, but these only account for a relatively small amount of variance in IQ, which itself has only a partial relationship to general intelligence and real-world performance.[27][28][full citation needed] One study indicated that in humans, fertility and intelligence tend to be negatively correlated—that is to say, the more intelligent, as measured by IQ, exhibit a lower total fertility rate than the less intelligent. According to the model, the present rate of decline is predicted to be 1.34 IQ points per decade.[29]

Sources of information

 
Computed tomography of human brain, from base of the skull to top, taken with intravenous contrast medium

Neuroscientists, along with researchers from allied disciplines, study how the human brain works. Such research has expanded considerably in recent decades. The "Decade of the Brain", an initiative of the United States Government in the 1990s, is considered to have marked much of this increase in research.[30] It has been followed in 2013 by the BRAIN Initiative.

Information about the structure and function of the human brain comes from a variety of experimental methods. Most information about the cellular components of the brain and how they work comes from studies of animal subjects, using techniques described in the brain article. Some techniques, however, are used mainly in humans, and therefore are described here.

Electrophysiology

Electroencephalography

By placing electrodes on the scalp it is possible to record the summed electrical activity of the cortex, using a methodology known as electroencephalography (EEG).[31] EEG records average neuronal activity from the cerebral cortex and can detect changes in activity over large areas but with low sensitivity for sub-cortical activity. EEG recordings are sensitive enough to detect tiny electrical impulses lasting only a few milliseconds. Most EEG devices have good temporal resolution, but low spatial resolution.

Electrocorticography

Electrodes can also be placed directly on the surface of the brain (usually during surgical procedures that require removal of part of the skull). This technique, called electrocorticography (ECoG), offers finer spatial resolution than electroencephalography, but is very invasive.

Magnetoencephalography

In addition to measuring the electric field directly via electrodes placed over the skull, it is possible to measure the magnetic field that the brain generates using a method known as magnetoencephalography (MEG).[32] This technique also has good temporal resolution like EEG but with much better spatial resolution. The greatest disadvantage of MEG is that, because the magnetic fields generated by neural activity are very subtle, the neural activity must be relatively close to the surface of the brain to detect its magnetic field. MEGs can only detect the magnetic signatures of neurons located in the depths of cortical folds (sulci) that have dendrites oriented in a way that produces a field.

Structural and functional imaging

 
A scan of the brain using fMRI
fMRI scan of the brain

There are several methods for detecting brain activity changes using three-dimensional imaging of local changes in blood flow. The older methods are SPECT and PET, which depend on injection of radioactive tracers into the bloodstream. A newer method, functional magnetic resonance imaging (fMRI), has considerably better spatial resolution and involves no radioactivity.[33] Using the most powerful magnets currently available, fMRI can localize brain activity changes to regions as small as one cubic millimeter. The downside is that the temporal resolution is poor: when brain activity increases, the blood flow response is delayed by 1–5 seconds and lasts for at least 10 seconds. Thus, fMRI is a very useful tool for learning which brain regions are involved in a given behavior, but gives little information about the temporal dynamics of their responses. A major advantage for fMRI is that, because it is non-invasive, it can readily be used on human subjects.

Another new non-invasive functional imaging method is functional near-infrared spectroscopy.

Effects of brain damage

A key source of information about the function of brain regions is the effects of damage to them.[34] In humans, strokes have long provided a "natural laboratory" for studying the effects of brain damage. Most strokes result from a blood clot lodging in the brain and blocking the local blood supply, causing damage or destruction of nearby brain tissue: the range of possible blockages is very wide, leading to a great diversity of stroke symptoms. Analysis of strokes is limited by the fact that damage often crosses into multiple regions of the brain, not along clear-cut borders, making it difficult to draw firm conclusions.

Transient ischemic attacks (TIAs) are mini-strokes that can cause sudden dimming or loss of vision (including amaurosis fugax), speech impairment ranging from slurring to dysarthria or aphasia, and mental confusion. But unlike a stroke, the symptoms of a TIA can resolve within a few minutes or 24 hours. Brain injury may still occur in a TIA lasting only a few minutes.[35][36] A silent stroke or silent cerebral infarct (SCI) differs from a TIA in that there are no immediately observable symptoms. An SCI may still cause long lasting neurological dysfunction affecting such areas as mood, personality, and cognition. An SCI often occurs before or after a TIA or major stroke.[37]

Language

Location of two brain areas historically associated with research on language processing, Broca's area and Wernicke's area

The study of how language is represented, processed, and acquired by the brain is neurolinguistics, which is a large multidisciplinary field drawing from cognitive neuroscience, cognitive linguistics, and psycholinguistics. This field originated from the 19th-century discovery that damage to different parts of the brain appeared to cause different symptoms: physicians noticed that individuals with damage to a portion of the left inferior frontal gyrus now known as Broca's area had difficulty in producing language (aphasia of speech), whereas those with damage to a region in the left superior temporal gyrus, now known as Wernicke's area, had difficulty in understanding it.[38]
Since then, there has been substantial debate over what linguistic processes these and other parts of the brain subserve,[39] and even over whether or not there is a strong one-to-one relationship between brain regions and language functions that emerges during neocortical development.[40] More recently, research on language has increasingly used more modern methods including electrophysiology and functional neuroimaging, to examine how language processing occurs. In the study of natural language, a dedicated network of language development has been identified as crucially involving Broca's area.[41][42]

Emotional prosody refers to speech that conveys emotions.[43]

Clinical significance

Clinically, death is defined as an absence of brain activity as measured by EEG. Injuries to the brain tend to affect large areas of the organ, sometimes causing major deficits in intelligence, memory, personality, and movement. Head trauma caused, for example, by vehicular or industrial accidents, is a leading cause of death in youth and middle age. In many cases, more damage is caused by resultant edema than by the impact itself. Stroke, caused by the blockage or rupturing of blood vessels in the brain, is another major cause of death from brain damage.

Other problems in the brain can be more accurately classified as diseases. Neurodegenerative diseases, such as Alzheimer's disease, Parkinson's disease, Huntington's disease and motor neuron diseases are caused by the gradual death of individual neurons, leading to diminution in movement control, memory, and cognition. There are five motor neuron diseases, the most common of which is amyotrophic lateral sclerosis (ALS).

Some infectious diseases affecting the brain are caused by viruses and bacteria. Infection of the meninges, the membranes that cover the brain, can lead to meningitis. Bovine spongiform encephalopathy (also known as "mad cow disease") is deadly in cattle and humans and is linked to prions. Kuru is a similar prion-borne degenerative brain disease affecting humans, (endemic only to Papua New Guinea tribes). Both are linked to the ingestion of neural tissue, and may explain the tendency in human and some non-human species to avoid cannibalism. Viral or bacterial causes have been reported in multiple sclerosis, and are established causes of encephalopathy, and encephalomyelitis.

Mental disorders, such as clinical depression, schizophrenia, bipolar disorder and post-traumatic stress disorder may involve particular patterns of neuropsychological functioning related to various aspects of mental and somatic function. These disorders may be treated by psychotherapy, psychiatric medication, social intervention and personal recovery work or cognitive behavioural therapy; the underlying issues and associated prognoses vary significantly between individuals.

Many brain disorders are congenital, occurring during development. Tay-Sachs disease, fragile X syndrome, and Down syndrome are all linked to genetic and chromosomal errors. Many other syndromes, such as the intrinsic circadian rhythm disorders, are suspected to be congenital as well. Normal development of the brain can be altered by genetic factors, drug use, nutritional deficiencies, and infectious diseases during pregnancy.

Metabolism

A flat oval object is surrounded by blue. The object is largely green-yellow, but contains a dark red patch at one end and a number of blue patches.
PET image of the human brain showing energy consumption

The brain consumes up to twenty percent of the energy used by the human body, more than any other organ.[44] Brain metabolism normally relies upon blood glucose as an energy source, but during times of low glucose (such as fasting, exercise, or limited carbohydrate intake), the brain will use ketone bodies for fuel with a smaller need for glucose. The brain can also utilize lactate during exercise.[45]
Long-chain fatty acids cannot cross the blood–brain barrier, but the liver can break these down to produce ketones. However the medium-chain fatty acids octanoic and heptanoic acids can cross the barrier and be used by the brain.[46][47][48] The brain stores glucose in the form of glycogen, albeit in significantly smaller amounts than that found in the liver or skeletal muscle.[49]

Although the human brain represents only 2% of the body weight, it receives 15% of the cardiac output, 20% of total body oxygen consumption, and 25% of total body glucose utilization.[50] The need to limit body weight has led to selection for a reduction of brain size in some species, such as bats, who need to be able to fly.[51] The brain mostly uses glucose for energy, and deprivation of glucose, as can happen in hypoglycemia, can result in loss of consciousness. The energy consumption of the brain does not vary greatly over time, but active regions of the cortex consume somewhat more energy than inactive regions: this fact forms the basis for the functional brain imaging methods PET and fMRI.[52] These are nuclear medicine imaging techniques which produce a three-dimensional image of metabolic activity.

Energy flow (ecology)

From Wikipedia, the free encyclopedia https://en.wikipedia.org/wiki/Energy_flow_(ecology) A graphic repr...