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Thursday, July 18, 2024

Image editing

From Wikipedia, the free encyclopedia
https://en.wikipedia.org/wiki/Image_editing
A colorized version of originally black and white photo, colorized using GIMP
Original black and white photo: Migrant Mother, showing Florence Owens Thompson, taken by Dorothea Lange in 1936.
This is a photo that has been edited as a Bokeh effect, using a Gaussian blur.

Image editing encompasses the processes of altering images, whether they are digital photographs, traditional photo-chemical photographs, or illustrations. Traditional analog image editing is known as photo retouching, using tools such as an airbrush to modify photographs or editing illustrations with any traditional art medium. Graphic software programs, which can be broadly grouped into vector graphics editors, raster graphics editors, and 3D modelers, are the primary tools with which a user may manipulate, enhance, and transform images. Many image editing programs are also used to render or create computer art from scratch. The term "image editing" usually refers only to the editing of 2D images, not 3D ones.

Basics of image editing

Raster images are stored in a computer in the form of a grid of picture elements, or pixels. These pixels contain the image's color and brightness information. Image editors can change the pixels to enhance the image in many ways. The pixels can be changed as a group, or individually, by the sophisticated algorithms within the image editors. This article mostly refers to bitmap graphics editors, which are often used to alter photographs and other raster graphics. However, vector graphics software, such as Adobe Illustrator, CorelDRAW, Xara Designer Pro or Inkscape, are used to create and modify vector images, which are stored as descriptions of lines, Bézier curves, and text instead of pixels. It is easier to rasterize a vector image than to vectorize a raster image; how to go about vectorizing a raster image is the focus of much research in the field of computer vision. Vector images can be modified more easily because they contain descriptions of the shapes for easy rearrangement. They are also scalable, being rasterizable at any resolution.

Automatic image enhancement

Camera or computer image editing programs often offer basic automatic image enhancement features that correct color hue and brightness imbalances as well as other image editing features, such as red eye removal, sharpness adjustments, zoom features and automatic cropping. These are called automatic because generally they happen without user interaction or are offered with one click of a button or mouse button or by selecting an option from a menu. Additionally, some automatic editing features offer a combination of editing actions with little or no user interaction.

Super-resolution imaging

There is promising research on using deep convolutional networks to perform super-resolution. In particular work has been demonstrated showing the transformation of a 20x microscope image of pollen grains into a 1500x scanning electron microscope image using it. While this technique can increase the information content of an image, there is no guarantee that the upscaled features exist in the original image and deep convolutional upscalers should not be used in analytical applications with ambiguous inputs. These methods can hallucinate image features, which can make them unsafe for medical use.

Digital data compression

Many image file formats use data compression to reduce file size and save storage space. Digital compression of images may take place in the camera, or can be done in the computer with the image editor. When images are stored in JPEG format, compression has already taken place. Both cameras and computer programs allow the user to set the level of compression.

Some compression algorithms, such as those used in PNG file format, are lossless, which means no information is lost when the file is saved. By contrast, the more popular JPEG file format uses a lossy compression algorithm (based on discrete cosine transform coding) by which the greater the compression, the more information is lost, ultimately reducing image quality or detail that can not be restored. JPEG uses knowledge of the way the human brain and eyes perceive color to make this loss of detail less noticeable.

Image editor features

Listed below are some of the most used capabilities of the better graphics manipulation programs. The list is by no means all-inclusive. There are a myriad of choices associated with the application of most of these features.

Selection

One of the prerequisites for many of the applications mentioned below is a method of selecting part(s) of an image, thus applying a change selectively without affecting the entire picture. Most graphics programs have several means of accomplishing this, such as:

  • a marquee tool for selecting rectangular or other regular polygon-shaped regions,
  • a lasso tool for freehand selection of a region,
  • a magic wand tool that selects objects or regions in the image defined by proximity of color or luminance,
  • vector-based pen tools,

as well as more advanced facilities such as edge detection, masking, alpha compositing, and color and channel-based extraction. The border of a selected area in an image is often animated with the marching ants effect to help the user to distinguish the selection border from the image background.

Layers

Leonardo da Vinci's Vitruvian Man overlaid with Goethe's Color Wheel using a screen layer in Adobe Photoshop. Screen layers can be helpful in graphic design and in creating multiple exposures in photography.
Leonardo da Vinci's Vitruvian Man overlaid a soft light layer Moses Harris's Color Wheel and a soft light layer of Ignaz Schiffermüller's Color Wheel. Soft light layers have a darker, more translucent look than screen layers.

Another feature common to many graphics applications is that of Layers, which are analogous to sheets of transparent acetate (each containing separate elements that make up a combined picture), stacked on top of each other, each capable of being individually positioned, altered and blended with the layers below, without affecting any of the elements on the other layers. This is a fundamental workflow which has become the norm for the majority of programs on the market today, and enables maximum flexibility for the user while maintaining non-destructive editing principles and ease of use.

Image size alteration

Image editors can resize images in a process often called image scaling, making them larger, or smaller. High image resolution cameras can produce large images which are often reduced in size for Internet use. Image editor programs use a mathematical process called resampling to calculate new pixel values whose spacing is larger or smaller than the original pixel values. Images for Internet use are kept small, say 640 x 480 pixels which would equal 0.3 megapixels.

Cropping an image

Digital editors are used to crop images. Cropping creates a new image by selecting a desired rectangular portion from the image being cropped. The unwanted part of the image is discarded. Image cropping does not reduce the resolution of the area cropped. Best results are obtained when the original image has a high resolution. A primary reason for cropping is to improve the image composition in the new image.

Uncropped image from camera
Lily cropped from larger image

Cutting out a part of an image from the background

Using a selection tool, the outline of the figure or element in the picture is traced/selected, and then the background is removed. Depending on how intricate the "edge" is this may be more or less difficult to do cleanly. For example, individual hairs can require a lot of work. Hence the use of the "green screen" technique (chroma key) which allows one to easily remove the background.

Histogram

Image editors have provisions to create an image histogram of the image being edited. The histogram plots the number of pixels in the image (vertical axis) with a particular brightness value (horizontal axis). Algorithms in the digital editor allow the user to visually adjust the brightness value of each pixel and to dynamically display the results as adjustments are made. Improvements in picture brightness and contrast can thus be obtained.

Sunflower image
Histogram of Sunflower image

Noise reduction

Image editors may feature a number of algorithms which can add or remove noise in an image. Some JPEG artifacts can be removed; dust and scratches can be removed and an image can be de-speckled. Noise reduction merely estimates the state of the scene without the noise and is not a substitute for obtaining a "cleaner" image. Excessive noise reduction leads to a loss of detail, and its application is hence subject to a trade-off between the undesirability of the noise itself and that of the reduction artifacts.

Noise tends to invade images when pictures are taken in low light settings. A new picture can be given an 'antiqued' effect by adding uniform monochrome noise.

Removal of unwanted elements

Most image editors can be used to remove unwanted branches, etc., using a "clone" tool. Removing these distracting elements draws focus to the subject, improving overall composition.

Notice the branch in the original image
The eye is drawn to the center of the globe.

Selective color change

Some image editors have color swapping abilities to selectively change the color of specific items in an image, given that the selected items are within a specific color range.

Selective color change

Image orientation

Image orientation (from left to right): original, 30° CCW rotation, and flipped.

Image editors are capable of altering an image to be rotated in any direction and to any degree. Mirror images can be created and images can be horizontally flipped or vertically flopped. A small rotation of several degrees is often enough to level the horizon, correct verticality (of a building, for example), or both. Rotated images usually require cropping afterwards, in order to remove the resulting gaps at the image edges.

Perspective control and distortion

Perspective control: original (left), perspective distortion removed (right).

Some image editors allow the user to distort (or "transform") the shape of an image. While this might also be useful for special effects, it is the preferred method of correcting the typical perspective distortion which results from photographs being taken at an oblique angle to a rectilinear subject. Care is needed while performing this task, as the image is reprocessed using interpolation of adjacent pixels, which may reduce overall image definition. The effect mimics the use of a perspective control lens, which achieves a similar correction in-camera without loss of definition.

Lens correction

Photo manipulation packages have functions to correct images for various lens distortions including pincushion, fisheye and barrel distortions. The corrections are in most cases subtle, but can improve the appearance of some photographs.

Enhancing images

In computer graphics, the enhancement of an image is the process of improving the quality of a digitally stored image by manipulating the image with software. It is quite easy, for example, to make an image lighter or darker, or to increase or decrease contrast. Advanced photo enhancement software also supports many filters for altering images in various ways. Programs specialized for image enhancement are sometimes called image editors.

Sharpening and softening images

Graphics programs can be used to both sharpen and blur images in a number of ways, such as unsharp masking or deconvolution. Portraits often appear more pleasing when selectively softened (particularly the skin and the background) to better make the subject stand out.  This can be achieved with a camera by using a large aperture, or in the image editor by making a selection and then blurring it. Edge enhancement is an extremely common technique used to make images appear sharper, although purists frown on the result as appearing unnatural.

Image sharpening: original (top), image sharpened (bottom).

Another form of image sharpening involves a form of contrast. This is done by finding the average color of the pixels around each pixel in a specified radius, and then contrasting that pixel from that average color. This effect makes the image seem clearer, seemingly adding details. An example of this effect can be seen to the right. It is widely used in the printing and photographic industries for increasing the local contrasts and sharpening the images.

Selecting and merging of images

Photomontage of 16 photos which have been digitally manipulated in Photoshop to give the impression that it is a real landscape

Many graphics applications are capable of merging one or more individual images into a single file. The orientation and placement of each image can be controlled.

When selecting a raster image that is not rectangular, it requires separating the edges from the background, also known as silhouetting. This is the digital-analog of cutting out the image from a physical picture. Clipping paths may be used to add silhouetted images to vector graphics or page layout files that retain vector data. Alpha compositing, allows for soft translucent edges when selecting images. There are a number of ways to silhouette an image with soft edges, including selecting the image or its background by sampling similar colors, selecting the edges by raster tracing, or converting a clipping path to a raster selection. Once the image is selected, it may be copied and pasted into another section of the same file, or into a separate file. The selection may also be saved in what is known as an alpha channel.

A popular way to create a composite image is to use transparent layers. The background image is used as the bottom layer, and the image with parts to be added are placed in a layer above that. Using an image layer mask, all but the parts to be merged are hidden from the layer, giving the impression that these parts have been added to the background layer. Performing a merge in this manner preserves all of the pixel data on both layers to more easily enable future changes in the new merged image.

Slicing of images

A more recent tool in digital image editing software is the image slicer. Parts of images for graphical user interfaces or web pages are easily sliced, labeled and saved separately from whole images so the parts can be handled individually by the display medium. This is useful to allow dynamic swapping via interactivity or animating parts of an image in the final presentation.

Special effects

An example of some special effects that can be added to a picture

Image editors usually have a list of special effects that can create unusual results. Images may be skewed and distorted in various ways. Scores of special effects can be applied to an image which include various forms of distortion, artistic effects, geometric transforms and texture effects, or combinations thereof.

A complex effect in the first image from the right


Using custom Curves settings in Image editors such as Photoshop, one can mimic the "pseudo-solarisation" effect, better known in photographic circles as the Sabattier-effect.

A pseudo-solarised color image

Stamp Clone Tool

The Clone Stamp tool selects and samples an area of your picture and then uses these pixels to paint over any marks. The Clone Stamp tool acts like a brush so you can change the size, allowing cloning from just one pixel wide to hundreds. You can change the opacity to produce a subtle clone effect. Also, there is a choice between Clone align or Clone non-align the sample area. In Photoshop this tool is called Clone Stamp, but it may also be called a Rubber Stamp tool.



Image after stamp tool processed

Change color depth

An example of converting an image from color to grayscale

It is possible, using the software, to change the color depth of images. Common color depths are 2, 4, 16, 256, 65,536 and 16.7 million colors. The JPEG and PNG image formats are capable of storing 16.7 million colors (equal to 256 luminance values per color channel). In addition, grayscale images of 8 bits or less can be created, usually via conversion and down-sampling from a full-color image. Grayscale conversion is useful for reducing the file size dramatically when the original photographic print was monochrome, but a color tint has been introduced due to aging effects.

Contrast change and brightening

An example of contrast correction. Left side of the image is untouched.

Image editors have provisions to simultaneously change the contrast of images and brighten or darken the image. Underexposed images can often be improved by using this feature. Recent advances have allowed more intelligent exposure correction whereby only pixels below a particular luminosity threshold are brightened, thereby brightening underexposed shadows without affecting the rest of the image. The exact transformation that is applied to each color channel can vary from editor to editor. GIMP applies the following formula:

if (brightness < 0.0)  value = value * ( 1.0 + brightness);
                  else value = value + ((1 - value) * brightness);
value = (value - 0.5) * (tan ((contrast + 1) * PI/4) ) + 0.5;

where value is the input color value in the 0..1 range and brightness and contrast are in the −1..1 range.

Gamma correction

In addition to the capability of changing the images' brightness and/or contrast in a non-linear fashion, most current image editors provide an opportunity to manipulate the images' gamma value.

Gamma correction is particularly useful for bringing details that would be hard to see on most computer monitors out of shadows. In some image editing software, this is called "curves", usually, a tool found in the color menu, and no reference to "gamma" is used anywhere in the program or the program documentation. Strictly speaking, the curves tool usually does more than simple gamma correction, since one can construct complex curves with multiple inflection points, but when no dedicated gamma correction tool is provided, it can achieve the same effect.

Color adjustments

An example of color adjustment using raster graphics editor

The color of images can be altered in a variety of ways. Colors can be faded in and out, and tones can be changed using curves or other tools. The color balance can be improved, which is important if the picture was shot indoors with daylight film, or shot on a camera with the white balance incorrectly set. Special effects, like sepia tone and grayscale, can be added to an image. In addition, more complicated procedures such as the mixing of color channels are possible using more advanced graphics editors.

The red-eye effect, which occurs when flash photos are taken when the pupil is too widely open (so that light from the flash that passes into the eye through the pupil reflects off the fundus at the back of the eyeball), can also be eliminated at this stage.

Dynamic blending

Before and After example of Advanced Dynamic Blending Technique created by Elia Locardi

Advanced Dynamic Blending is a concept introduced by photographer Elia Locardi in his blog Blame The Monkey to describe the photographic process of capturing multiple bracketed exposures of a land or cityscape over a specific span of time in a changing natural or artificial lighting environment. Once captured, the exposure brackets are manually blended together into a single High Dynamic Range image using post-processing software. Dynamic Blending images serve to display a consolidated moment. This means that while the final image may be a blend of a span of time, it visually appears to represent a single instant.

Printing

Control printed image by changing pixels-per-inch

Controlling the print size and quality of digital images requires an understanding of the pixels-per-inch (ppi) variable that is stored in the image file and sometimes used to control the size of the printed image. Within Adobe Photoshop's Image Size dialog, the image editor allows the user to manipulate both pixel dimensions and the size of the image on the printed document. These parameters work together to produce a printed image of the desired size and quality. Pixels per inch of the image, pixel per inch of the computer monitor, and dots per inch on the printed document are related, but in use are very different. The Image Size dialog can be used as an image calculator of sorts. For example, a 1600 × 1200 image with a resolution of 200 ppi will produce a printed image of 8 × 6 inches. The same image with 400 ppi will produce a printed image of 4 × 3 inches. Change the resolution to 800 ppi, and the same image now prints out at 2 × 1.5 inches. All three printed images contain the same data (1600 × 1200 pixels), but the pixels are closer together on the smaller prints, so the smaller images will potentially look sharp when the larger ones do not. The quality of the image will also depend on the capability of the printer.

California mixed evergreen forest

From Wikipedia, the free encyclopedia
California mixed evergreen forest in the Santa Cruz Mountains.

California mixed evergreen forest is a plant community found in the mountain ranges of California and southwestern Oregon.

The Mixed evergreen forest plant community is native to the Northern and Southern California Coast Ranges and Sierra Nevada of central and northern California; the Transverse Ranges and Peninsular Ranges of southern California; and the southwestern Oregon Coast Ranges.

California mixed evergreen forests occur in ecoregions of the California Floristic Province, including in areas of the California chaparral and woodlands and its sub-ecoregions, Klamath-Siskiyou Mountains forests, Northern California coastal forests, and Sierra Nevada lower montane forest. The mixed evergreen forests of each ecoregion have slightly different species composition.

Klamath-Siskiyou mixed evergreen forest

The mixed evergreen forests of the Klamath Mountains-Siskiyou Mountains occur above 300 meters (1000 ft) elevation, and are of four main types. Douglas-fir forests are found on gentle slopes, north-facing slopes, ridges with deep soil, and river terraces with deep sediments, usually underlain with sedimentary rocks. Coast Douglas-fir (Pseudotsuga menziesii ssp. menziesii) is the predominant tree, occupying up to 70% of the forest cover. Broadleaf evergreen trees are relatively few. Tree species of secondary importance are:

On granite soils, a similar species composition predominates, but with more broadleaf evergreens, chiefly:

On steep, well drained slopes, canyon live oak is the dominant species, with coast Douglas-fir in a minor role.

On serpentine soils, mixed evergreen forests are made up of:

moister areas
drier areas

California Coast Ranges mixed evergreen forest

Mixed evergreen forests in the California Coast Ranges vary in species related to coastal (Outer Ranges) or inland (Inner Ranges) climate influences. It usually happens in the elevation from 0 meter to 500 meters high. Mountains in the San Francisco Bay Area frequently support mixed evergreen forests. Generally the moister the forest habitat the greater the understory density. Coast mixed evergreen is drier than coast redwood and it is a four-layer forest structure. The conifers are usually emergent as canopy layer and broad-leaf evergreen trees emerge as sub-canopy layer. The shrubs occur as the understory species.

Those closer to the coast receiving adequate fog moisture and high rainfall generally will feature the following species:
Further inland, on north facing slopes or protected canyons, these forests are drier and generally lack tanoak, Douglas fir, and coast redwood. Species typical of the drier neighboring oak woodlands join the mix:

Common wildlife include Steller's jay, chestnut-backed chickadee, western gray squirrel, raccoon and many others. These forests are more diverse in animal life than those having only conifers. The predominance of broadleaf trees provide a greater abundance of food sources.

Southern Coast, Transverse, and Peninsular Ranges mixed evergreen forest

Also called southern mixed evergreen forests, the mixed evergreen forests of the southern California Coast Ranges, the Transverse Ranges, and the northern Peninsular Ranges, are restricted to cooler north-facing slopes at higher elevations.

Predominant tree species include:

Northern California coastal forests

Northern California coastal forests
Ecology
RealmNearctic
BiomeTemperate coniferous forests
Borders
Bird species232
Mammal species77
Geography
Area13,300 km2 (5,100 sq mi)
CountryUnited States
States
Climate typeMediterranean (Csb)
Conservation
Conservation statusCritical/Endangered
Habitat loss4.6828%
Protected15.76%

The Northern California coastal forests are a temperate coniferous forests ecoregion of coastal Northern California and southwestern Oregon.

Setting

The ecoregion covers 13,300 square kilometres (5,100 sq mi), extending from just north of the California-Oregon border south, to southern Monterey County. The ecoregion rarely extends more than 65 km inland from the coast, narrower in the southernmost parts of the ecoregion.

The ecoregion is a sub-ecoregion of the Pacific temperate rain forests ecoregion, which extends up the Pacific Coast to Kodiak Island in Alaska. The ecoregion lies close to the Pacific Ocean, and is kept moist by Pacific Ocean storms during the winter months, and by coastal fogs in the summer months. These factors keep the ecoregion cooler in the summer and warmer in the winter, as compared to ecoregions further inland. The ecoregion is also defined by the distribution of the Coast Redwood (Sequoia sempervirens), with isolated groves located in protected canyons as far south as Redwood Gulch, in southern Monterey County. The greatest concentration of remaining Old-growth forest are in the northernmost portion of the ecoregion, primarily within Humboldt and Del Norte counties.

Major urban centers located within this ecoregion include the montane portions of various cities of the San Francisco Peninsula, Fort Bragg, Eureka, and Brookings.

Habitats

Redwood forests are interspersed with several other plant communities throughout this ecoregion.

Coastal redwood forests

Notholithocarpus densiflorus, with Coast Douglas-firs Pseudotsuga menziesii subsp. menziesii and Coast Redwood behind in Sunset Trail, Big Basin Redwoods State Park, Santa Cruz Mountains, California.

The dominant forest type in this ecoregion is the coastal redwood forest. These are the tallest forests on Earth, with individual redwood (Sequoia sempervirens) trees reaching heights of 100 metres (330 ft). These forests are generally found in areas exposed to coastal fog. In the north, they occur on upland slopes, in riparian zones, and on riverine terraces. In the south, where annual precipitation is lower, they are constrained to coves and ravines. Coast Douglas-firs (Pseudotsuga menziesii var. menziesii) are nearly always associated with redwoods, but in the north the forests can also include Sitka spruce (Picea sitchensis), western hemlock (Tsuga heterophylla) and western red cedar (Thuja plicata). Like coast Douglas-fir, tanoak (Notholithocarpus densiflorus) is often present. Other hardwoods include California bay laurel (Umbellularia californica), red alder (Alnus rubra), madrone (Arbutus menziesii), and bigleaf maple (Acer macrophyllum). The deep shade cast by redwoods often results in a sparse understory, but shade-tolerant species include thimbleberry (Rubus parviflorus), redwood sorrel (Oxalis oregana), elk clover (Aralia californica), dwarf Oregon grape (Mahonia nervosa), salal (Gaultheria shallon), and many ferns, such as deer fern (Blechnum spicant), sword fern (Polystichum munitum), and leathery polypody (Polypodium scouleri). 

Mixed evergreen forests

Mixed evergreen forests are found just inland of the redwood forests, on Franciscan Assemblage soils that receive moderate to high rainfall. The trees are a variety of needle-leaved and broad-leaved evergreen species. Characteristic trees include coast Douglas-fir (Pseudotsuga menziesii var. menziesii), canyon live oak (Quercus chrysolepis), tanoak (Notholithocarpus densiflorus), madrone (Arbutus menziesii), California bay laurel (Umbellularia californica), and golden chinquapin (Chrysolepis chrysophylla). The shrub understory is dense and diverse; beaked hazel (Corylus cornuta), evergreen huckleberry (Vaccinium ovatum), Pacific rhododendron (Rhododendron macrophyllum), salal (Gaultheria shallon), Sadler's oak (Quercus sadleriana), dwarf Oregon-grape (Mahonia nervosa), and poison oak (Toxicodendron diversilobum) are typically found.

Closed-cone conifer forests and woodlands

Pinus muricata forest growing at Point Reyes, California

Closed-cone conifer forests are found in small, scattered patches throughout the ecoregion, typically adjacent to maritime chaparral. Common pines are lodgepole pine (Pinus contorta), bishop pine (Pinus muricata), Monterey pine (Pinus radiata), and knobcone pine (Pinus attenuata). These forests can also be home to several endemic cypresses, including Monterey cypress (Cupressus macrocarpa), Gowen cypress (Cupressus goveniana), and Santa Cruz cypress (Cupressus abramsiana). Shrub species include glossyleaf manzanita (Arctostaphylos nummularia), bog Labrador tea (Rhododendron groenlandicum), evergreen huckleberry (Vaccinium ovatum), salal (Gaultheria shallon), Pacific rhododendron (Rhododendron macrophyllum), and California bayberry (Myrica californica). Soil conditions sometimes cause these forests to take on a pygmy form. Lichens and mosses are diverse and can be abundant.

Maritime chaparral

Maritime chaparral is composed of a variety of shrubs that grow in the fog belt. Endemic species of manzanita (Arctostaphylos) and Ceanothus are locally common. Manzanita species include woolyleaf manzanita (Arctostaphylos tomentosa), glossyleaf manzanita (Arctostaphylos nummularia), Hooker's manzanita (Arctostaphylos hookeri), pajaro manzanita (Arctostaphylos pajaroensis), Montara manzanita (Arctostaphylos montaraensis), and others. Gasquet manzanita (Arctostaphylos hispidula) occurs in southern Oregon. Among Ceanothus, hairy ceanothus (Ceanothus oliganthus) is common, while Mason's ceanothus (Ceanothus masonii), Carmel ceanothus (Ceanothus griseus), and wart-stem ceanothus (Ceanothus verrucosus) are local endemics. Other widespread shrubs and trees include chamise (Adenostoma fasciculatum), California buckwheat (Eriogonum fasciculatum), black sage (Salvia mellifera), coffeeberry (Rhamnus californica), buckthorn (Rhamnus crocea), and coast live oak (Quercus agrifolia). This habitat is often found near closed-cone conifer forests and woodlands.

Coastal grassland

Iris douglasiana patch at Point Reyes

Northern coastal grasslands, or coastal prairies, are generally found below 1,000 feet (300 m) on coastal terraces or mountain balds. In areas where fire has been suppressed, coastal scrub plants invade. Common grasses include bentgrass (Agrostis spp.), California brome (Bromus carinatus), Nootka reedgrass (Calamagrostis nutkaensis), California oatgrass (Danthonia californica), red fescue (Festuca rubra), Idaho fescue (Festuca idahoensis), tufted hair-grass (Deschampsia caespitosa), prairie Junegrass (Koeleria macrantha), tall trisetuem (Trisetum canescens). Common forbs include Douglas iris (Iris douglasiana), western blue-eyed grass (Sisyrinchium bellum), hairy gumplant (Grindelia hirsutula), and footsteps of spring (Sanicula arctopoides).

Coastal scrub

Northern coastal scrub consists of shrublands found at elevations below 1,500 feet (460 m) on bluffs, terraces, dunes, and hills near the coast. This habitat is often subject to wind and maritime fog. The shrubs are mostly evergreen, small-leaved, and sclerophyllous. Characteristic species include coyote brush (Baccharis pilularis), yellow bush lupine (Lupinus arboreus), blueblossom (Ceanothus thyrsiflorus), seaside woolly sunflower (Eriophyllum stoechadifolium), sticky monkey-flower (Mimulus aurantiacus), poison oak (Toxicodendron diversilobum), California blackberry (Rubus ursinus), thimbleberry (Rubus parviflorus), salmonberry (Rubus spectabilis), coffeeberry (Rhamnus californica), oceanspray (Holodiscus discolor), salal (Gaultheria shallon), cow parsnip (Heracleum maximum), and western sword fern (Polystichum munitum). Coastal grassland succeeds to coastal scrub in the absence of fire, and coastal scrub succeeds to mixed evergreen forest under further absence of fire.

Riparian woodlands and shrublands

Riparian woodlands and shrublands are a mosaic of tree-dominated plant communities and open shrublands found along rivers. Species composition varies with elevation, slope, floodplain width, and flooding history. Nevertheless, common trees include white alder (Alnus rhombifolia), red alder (Alnus rubra), box elder (Acer negundo), Fremont cottonwood (Populus fremontii), red willow (Salix laevigata), coast Douglas-fir (Pseudotsuga menziesii var. menziesii), California sycamore (Platanus racemosa), coast live oak (Quercus agrifolia), and bigleaf maple (Acer macrophyllum). Common shrubs include sandbar willow (Salix exigua) and arroyo willow (Salix lasiolepis).

Live oak woodlands and savannas

Live oak woodlands and savannas are dominated by coast live oak (Quercus agrifolia). Canopy cover varies from dense forest to open savannas. In forests, California blackberry (Rubus ursinus), creeping snowberry (Symphoricarpos mollis), toyon (Heteromeles arbutifolia), and poison oak (Toxicodendron diversilobum) are common in the understory.

Ponderosa pine forests

Some of the rarest forests that occurs in this coastal region are the Maritime Coast Range Ponderosa Pine forests, an example of which occurs in the Carbonera Creek watershed of Santa Cruz County, California. These forest are dominated by ponderosa pine (Pinus ponderosa).

Sequoia sempervirens

From Wikipedia, the free encyclopedia
Sequoia sempervirens
Sequoia sempervirens along US 199




















Binomial name
Sequoia sempervirens
Natural range of California subfamily Sequoioideae
green - Sequoia sempervirens
Trunk in sectional view
Redwood cone scales begin to open mid November, with seeds dispersing by the wind.

Sequoia sempervirens (/səˈkwɔɪ.ə ˌsɛmpərˈvrənz/) is the sole living species of the genus Sequoia in the cypress family Cupressaceae (formerly treated in Taxodiaceae). Common names include coast redwood, coastal redwood and California redwood. It is an evergreen, long-lived, monoecious tree living 1,200–2,200 years or more. This species includes the tallest living trees on Earth, reaching up to 115.9 m (380.1 ft) in height (without the roots) and up to 8.9 m (29 ft) in diameter at breast height. These trees are also among the longest-living trees on Earth. Before commercial logging and clearing began by the 1850s, this massive tree occurred naturally in an estimated 810,000 ha (2,000,000 acres) along much of coastal California (excluding southern California where rainfall is not sufficient) and the southwestern corner of coastal Oregon within the United States.

The name sequoia sometimes refers to the subfamily Sequoioideae, which includes S. sempervirens along with Sequoiadendron (giant sequoia) and Metasequoia (dawn redwood). Here, the term redwood on its own refers to the species covered in this article but not to the other two species.

Description

The coast redwood normally reaches a height of 60 to 100 m (200 to 330 ft), but will be more than 110 m (360 ft) in extraordinary circumstances, with a trunk diameter of 9 m (30 ft). It has a conical crown, with horizontal to slightly drooping branches. The trunk is remarkably straight. The bark can be very thick, up to 35 cm (1.15 ft), and quite soft and fibrous, with a bright red-brown color when freshly exposed (hence the name redwood), weathering darker. The root system is composed of shallow, wide-spreading lateral roots.

The leaves are variable, being 15–25 mm (58–1 in) long and flat on young trees and shaded lower branches in older trees. The leaves are scalelike, 5–10 mm (1438 in) long on shoots in full sun in the upper crown of older trees, with a full range of transition between the two extremes. They are dark green above and have two blue-white stomatal bands below. Leaf arrangement is spiral, but the larger shade leaves are twisted at the base to lie in a flat plane for maximum light capture.

The species is monoecious, with pollen and seed cones on the same plant. The seed cones are ovoid, 15–32 mm (9161+14 in) long, with 15–25 spirally arranged scales; pollination is in late winter with maturation about 8–9 months after. Each cone scale bears three to seven seeds, each seed 3–4 mm (18316 in) long and 0.5 mm (132 in) broad, with two wings 1 mm (116 in) wide. The seeds are released when the cone scales dry and open at maturity. The pollen cones are ovular and 4–6 mm (31614 in) long.

Its genetic makeup is unusual among conifers, being a hexaploid (6n) and possibly allopolyploid (AAAABB). Both the mitochondrial and chloroplast genomes of the redwood are paternally inherited.

Taxonomy

Scottish botanist David Don described the redwood as Taxodium sempervirens, the "evergreen Taxodium", in his colleague Aylmer Bourke Lambert's 1824 work A description of the genus Pinus. Austrian botanist Stephan Endlicher erected the genus Sequoia in his 1847 work Synopsis coniferarum, giving the redwood its current binomial name of Sequoia sempervirens. It is unknown how Endlicher derived the name Sequoia. See Sequoia Etymology.

The redwood is one of three living species, each in its own genus, in the subfamily Sequoioideae. Molecular studies have shown that the three are each other's closest relatives, generally with the redwood and giant sequoia (Sequoiadendron giganteum) as each other's closest relatives.

However, Yang and colleagues in 2010 queried the polyploid state of the redwood and speculate that it may have arisen as an ancient hybrid between ancestors of the giant sequoia and dawn redwood (Metasequoia). Using two different single copy nuclear genes, LFY and NLY, to generate phylogenetic trees, they found that Sequoia was clustered with Metasequoia in the tree generated using the LFY gene, but with Sequoiadendron in the tree generated with the NLY gene. Further analysis strongly supported the hypothesis that Sequoia was the result of a hybridization event involving Metasequoia and Sequoiadendron. Thus, Yang and colleagues hypothesize that the inconsistent relationships among Metasequoia, Sequoia, and Sequoiadendron could be a sign of reticulate evolution (in which two species hybridize and give rise to a third) among the three genera. However, the long evolutionary history of the three genera (the earliest fossil remains being from the Jurassic) make resolving the specifics of when and how Sequoia originated once and for all a difficult matter—especially since it in part depends on an incomplete fossil record.

Names

The species name "sempervirens" means "evergreen", thought to be because of its previous placement in the same genus as Taxodium distichum (baldcypress) of the southeastern USA. Unlike coast redwood, baldcypress loses its leaves in winter. The common name "redwood", applied to both the coast redwood and the giant redwood, is a reference to the red heartwood of the trees. Common names that refer to Sequoia sempervirens alone include "California redwood", "coastal redwood", "coastal sequoia", and "coast redwood".

Distribution and habitat

Coast redwoods occupy a narrow strip of land approximately 750 km (470 mi) in length and 8–75 km (5–47 mi) in width along the Pacific coast of North America; the most southerly grove is in Monterey County, California, and the most northerly groves are in extreme southwestern Oregon. The prevailing elevation range is 30–750 m (100–2,460 ft) above sea level, occasionally down to 0 and up to about 900 m (3,000 ft). They usually grow in the mountains where precipitation from the incoming moisture off the ocean is greater. The tallest and oldest trees are found in deep valleys and gullies, where year-round streams can flow, and fog drip is regular. The terrain also made it harder for loggers to get to the trees and to get them out after felling. The trees above the fog layer, above about 700 m (2,300 ft), are shorter and smaller due to the drier, windier, and colder conditions. In addition, Douglas-fir, pine, and tanoak often crowd out redwoods at these elevations. Few redwoods grow close to the ocean, due to intense salt spray, sand, and wind. Coalescence of coastal fog accounts for a considerable part of the trees' water needs. Fog in the 21st century is, however, reduced from what it was in the prior century, which is a problem that may be compounded by climate change.

Painting of a hillside of redwoods with the edge sun seen behind a hill
Albert Bierstadt, California Redwoods, 1872, Oil on paper mounted on canvas

The northern boundary of its range is marked by two groves on mountain slopes along the north side of the Chetco River, which is on the western fringe of the Klamath Mountains, near the California–Oregon border. The northernmost grove is located within Alfred A. Loeb State Park and Siskiyou National Forest at the approximate coordinates 42°07'36"N 124°12'17"W. The southern boundary of its range is the Los Padres National Forest's Silver Peak Wilderness in the Santa Lucia Mountains of the Big Sur area of Monterey County, California. The southernmost grove is in the Southern Redwood Botanical Area, just north of the national forest's Salmon Creek trailhead and near the San Luis Obispo County line.

The largest and tallest populations are in California's Redwood National and State Parks (Del Norte and Humboldt counties) and Humboldt Redwoods State Park, with the overall majority located in the large Humboldt County.

The ancient range of the genus is considerably greater, with relatives of the coast redwood living in Europe and Asia prior to the Quaternary geologic period. In recent geologic time there have been considerable shifts in the coast redwood's range in North America. Coast redwood bark has been found in the La Brea Tar Pits, showing that 25,000–40,000 years before the present redwood trees grew as far south as the Los Angeles during the last ice age. The authors of a 2022 paper suggest, "Were it not for the remarkable ability to sprout after fire, many southern forests may have lost their Sequoia component long ago." As to previous redwood range to the north, an upright fossil stump of a coast redwood on a beach in central Oregon was documented 257 km north of the current range.

Assisted migration

Coast redwood (2016 photo) planted in Seabeck, WA in 1949.

The ability of Coast Redwood to live for more than a thousand years, along with its unusual capacity to resprout from its root crown when felled by natural or human causes, have earned this species the label of "carbon-sequestration champion." Its potential to contribute toward climate change mitigation, as well as its demonstrated ability to thrive in coastal regions of the Pacific Northwest, led to the formation of a citizen group in Seattle, Washington undertaking assisted migration of this species hundreds of miles north of its native range.

In contrast to cautionary statements made by forestry professionals assessing other tree species for assisted migration, the citizens involved with the group known as PropagationNation had met with little controversy until in 2023 a national news outlet published a lengthy article that cast a favorable light on their efforts. The New York Times Magazine wrote:

Not wanting to cause ecological problems by planting the trees across the Pacific Northwest, [Philip] Stielstra would eventually contact one of the foremost experts on the coast redwood, a botanist and forest ecologist named Stephen Sillett, at Cal Poly Humboldt, and ask if moving redwoods north was safe. Sillett thought planting redwoods around Seattle was a fantastic idea. ("It's not like it's going to escape and become a nuisance species," Sillett told me, before adding, "it just has so many benefits.") Another factor encouraged Stielstra too: Millions of years ago, redwoods — or their close relatives — grew across the Pacific Northwest. By moving them, Stielstra reasoned, he was helping the magnificent trees regain lost territory.

In December 2023, the Associated Press exclusively reported criticism from professionals in the region and nationally: While beginning to favor experiments in assisted population migration of more southerly genetics of the main native timber tree, Douglas-fir, professionals were united against large-scale plantings of California redwoods into the Pacific Northwest. The next month, January 2024, carried a regional news article that, once again, showed strong support as well as bold statements by the group's founder.

Even before the controversy developed in Washington state, professionals in Canada were documenting horticultural plantings of the California species already in place in southwestern British Columbia. In 2022 a Canadian Forestry Service publication used northward horticultural plantings, along with a review of research detailing redwood's paleobiogeography and current range conditions, as grounds for proposing that Canada's Vancouver Island already offered "narrow strips of optimal habitat" for extending the range of coast redwood. The authors point to a topographical "bottleneck" north of the California border that could have impeded northward migration during the Holocene. The bottleneck entails a lack of lowland passages through the Oregon Coast Range north of the Chetco River, coupled with the absence of coastal landscapes beyond storm salt-spray and tsunami inundation — for which this conifer species is highly intolerant.

Ecology

Fog and flood adaptations

Fog is of major importance in coast redwood ecology. Redwood National Park.
Cross section of a Sequoia sempervirens showing tree rings

The native area provides a unique environment with heavy seasonal rains up to 2,500 mm (100 in) annually. Cool coastal air and fog drip keep the forest consistently damp year round. Several factors, including the heavy rainfall, create a soil with fewer nutrients than the trees need, causing them to depend heavily on the entire biotic community of the forest, and making efficient recycling of dead trees especially important. This forest community includes coast Douglas-fir, Pacific madrone, tanoak, western hemlock, and other trees, along with a wide variety of ferns, mosses, mushrooms, and redwood sorrel. Redwood forests provide habitat for a variety of amphibians, birds, mammals, and reptiles. Old-growth redwood stands provide habitat for the federally threatened spotted owl and the California-endangered marbled murrelet.

The height of S. sempervirens is closely tied to fog availability; taller trees become less frequent as fog becomes less frequent. As S. sempervirens' height increases, transporting water via water potential to the leaves becomes increasingly difficult due to gravity. Despite the high rainfall that the region receives (up to 100 cm), the leaves in the upper canopy are perpetually stressed for water. This water stress is exacerbated by long droughts in the summer. Water stress is believed to cause the morphological changes in the leaves, stimulating reduced leaf length and increased leaf succulence. To supplement their water needs, redwoods use frequent summer fog events. Fog water is absorbed through multiple pathways. Leaves directly take in fog from the surrounding air through the epidermal tissue, bypassing the xylem. Coast redwoods also absorb water directly through their bark. The uptake of water through leaves and bark repairs and reduces the severity of xylem embolisms, which occur when cavitations form in the xylem preventing the transport of water and nutrients. Fog may also collect on redwood leaves, drip to the forest floor, and be absorbed by the tree's roots. This fog drip may form 30% of the total water used by a tree in a year.

Redwoods often grow in flood-prone areas. Sediment deposits can form impermeable barriers that suffocate tree roots, and unstable soil in flooded areas often causes trees to lean to one side, increasing the risk of the wind toppling them. Immediately after a flood, redwoods grow their existing roots upwards into recently deposited sediment layers. A second root system then develops from adventitious buds on the newly buried trunk and the old root system dies. To counter lean, redwoods increase wood production on the vulnerable side, creating a supporting buttress. These adaptations create forests of almost exclusively redwood trees in flood-prone regions.

Pest and pathogen resistance

A recently fallen canopy branch reveals thin bark and tunnels made by bark beetle larvae.

Coast redwoods are resistant to insect attack, fungal infection, and rot. These properties are conferred by concentrations of terpenoids and tannic acid in redwood leaves, roots, bark, and wood. Despite these chemical defenses, redwoods are still subject to insect infestations; none, however, are capable of killing a healthy tree. Bark is so thick on the bole that bark beetles cannot enter there. However, the canopy branches have thin bark (see photo at right) that native bark beetles are able to bore into for egg-laying and larval growth via tunnels.

Redwoods also face herbivory from mammals: black bears are reported to consume the inner bark of small redwoods, and black-tailed deer are known to eat redwood sprouts.

The oldest known coast redwood is about 2,200 years old; many others in the wild exceed 600 years. The numerous claims of older redwoods are incorrect. Because of their seemingly timeless lifespans, coast redwoods were deemed the "everlasting redwood" at the turn of the century; in Latin, sempervirens means "ever green" or "everlasting". Redwoods must endure various environmental disturbances to attain such great ages.

Fire adaptations

In response to forest fires, the trees have developed various adaptations. The thick, fibrous bark of coast redwoods is extremely fire-resistant; it grows to at least a foot thick and protects mature trees from fire damage. In addition, the redwoods contain little flammable pitch or resin. Fires, moreover, appear to actually benefit redwoods by causing substantial mortality in competing species, while having only minor effects on redwood. Burned areas are favorable to the successful germination of redwood seeds. A study published in 2010, the first to compare post-wildfire survival and regeneration of redwood and associated species, concluded that fires of all severity increase the relative abundance of redwood, and higher-severity fires provide the greatest benefit.

Two years after the 2020 CZU fire that burned as high as the canopy in Big Basin Redwoods State Park. In addition to the customary sprouting of massive numbers of vertical shoots growing at ground-level around the charred trunks, thick epicormic greenery appears high up on some of the fire-pruned redwoods. (Photos were taken along the old growth loop trail, September 2022.)

Self-pruning of its lower limbs as height is gained is a crucial adaptation to prevent ground fires from rising into the canopy, where branch bark is thin and leaves are vulnerable. In the millions of years that predated human evolution, this level of protection worked well against natural fires originating from lightning strikes.

When the first humans arrived in North America, redwoods thrived in regions where ground fires were intentionally set by indigenous populations on a seasonal basis. However, when peoples arrived from other continents in the past few centuries, indigenous fire practices were disallowed — even in the few places where the first peoples were permitted to continue living. Flammable brush and young trees thus accumulated.

Clearcut logging further hampered a return to a fire-resistant tall canopy. Governmental policies aimed at suppressing all natural and human-caused fires, even in parks and wilderness areas, amplified the accumulation of dense undergrowth and woody debris. Thus, even a naturally occurring ground fire could threaten to spread upwards and become a canopy fire spreading uncontrollably over a widening area.

Reproduction

Lignotuber actively budding.
Lignotuber, Roosevelt Grove.
Basal regrowth after logging.
View upward from the center of an old, logged stump (Freshwater Creek).

Coast redwood reproduces both sexually by seed and asexually by sprouting of buds, layering, or lignotubers. Seed production begins at 10–15 years of age. Cones develop in the winter and mature by fall. In the early stages, the cones look like flowers, and are commonly called "flowers" by professional foresters, although this is not strictly correct. Coast redwoods produce many cones, with redwoods in new forests producing thousands per year. The cones themselves hold 90–150 seeds, but viability of seed is low, typically well below 15% with one estimate of average rates being 3 to 10 percent.The viability does increase with age, trees under 20 years old have a viability of about 1%, and do not generally reach the highest levels of viability until age 250. The rates decrease as the tree starts to get very old, with trees over 1,200 not reaching viability rates over 3%. The low viability may discourage seed predators, which do not want to waste time sorting chaff (empty seeds) from edible seeds. Successful germination often requires a fire or flood, reducing competition for seedlings. The winged seeds are small and light, weighing 3.3–5.0 mg (200–300 seeds/g; 5,600–8,500/ounce). The wings are not effective for wide dispersal, and seeds are dispersed by wind an average of only 60–120 m (200–390 ft) from the parent tree. Seedlings are susceptible to fungal infection and predation by banana slugs, brush rabbits, and nematodes. Most seedlings do not survive their first three years. However, those that become established grow rapidly, with young trees known to reach 20 m (66 ft) tall in 20 years. When canopy space is not available, small trees can remain suppressed for up to 400 years before accelerating their growth rate.

Coast redwoods can also reproduce asexually by layering or sprouting from the root crown, stump, or even fallen branches; if a tree falls over, it generates a row of new trees along the trunk, so many trees naturally grow in a straight line. Sprouts originate from dormant or adventitious buds at or under the surface of the bark. The dormant sprouts are stimulated when the main adult stem gets damaged or starts to die. Many sprouts spontaneously erupt and develop around the circumference of the tree trunk. Within a short period after sprouting, each sprout develops its own root system, with the dominant sprouts forming a ring of trees around the parent root crown or stump. This ring of trees is called a "fairy ring". Sprouts can achieve heights of 2.3 m (7 ft 7 in) in a single growing season.

Redwoods may also reproduce using burls. A burl is a woody lignotuber that commonly appears on a redwood tree below the soil line, though usually within 3 m (10 ft) in depth from the soil surface. Coast redwoods develop burls as seedlings from the axils of their cotyledon, a trait that is extremely rare in conifers. When provoked by damage, dormant buds in the burls sprout new shoots and roots. Burls are also capable of sprouting into new trees when detached from the parent tree, though exactly how this happens is yet to be studied. Shoot clones commonly sprout from burls and are often turned into decorative hedges when found in suburbia.

Cultivation and uses

An example of a bonsai redwood, from the Pacific Bonsai Museum

Coast redwood is one of the most valuable timber species in the lumbering industry. In California, 3,640 km2 (899,000 acres) of redwood forest are logged, virtually all of it second growth. Though many entities have existed in the cutting and management of redwoods, perhaps none has had a more storied role than the Pacific Lumber Company (1863–2008) of Humboldt County, California, where it owned and managed over 810 km2 (200,000 acres) of forests, primarily redwood. Coast redwood lumber is highly valued for its beauty, light weight, and resistance to decay. Its lack of resin makes it absorb water and resist fire.

P. H. Shaughnessy, Chief Engineer of the San Francisco Fire Department wrote:

In the recent great fire of San Francisco, that began April 18th, 1906, we succeeded in finally stopping it in nearly all directions where the unburned buildings were almost entirely of frame construction, and if the exterior finish of these buildings had not been of redwood lumber, I am satisfied that the area of the burned district would have been greatly extended.

Because of its impressive resistance to decay, redwood was extensively used for railroad ties and trestles throughout California. Many of the old ties have been recycled for use in gardens as borders, steps, house beams, etc. Redwood burls are used in the production of table tops, veneers, and turned goods.

The Skyline-to-the-Sea Trail passing through a cut-out in a fallen California redwood tree

The Yurok people, who occupied the region before European settlement, regularly burned ground cover in redwood forests to bolster tanoak populations from which they harvested acorns, to maintain forest openings, and to boost populations of useful plant species such as those for medicine or basketmaking.

Extensive logging of redwoods began in the early nineteenth century. The trees were felled by ax and saw onto beds of tree limbs and shrubs to cushion their fall. Stripped of their bark, the logs were transported to mills or waterways by oxen or horse. Loggers then burned the accumulated tree limbs, shrubs, and bark. The repeated fires favored secondary forests of primarily redwoods as redwood seedlings sprout readily in burned areas. The introduction of steam engines let crews drag logs through long skid trails to nearby railroads, furthering the reach of loggers beyond the land near rivers previously used to transport trees. This method of harvesting, however, disturbed large amounts of soil, producing secondary-growth forests of species other than redwood such as Douglas-fir, grand fir, and western hemlock. After World War II, trucks and tractors gradually replaced steam engines, giving rise to two harvesting approaches: clearcutting and selection harvesting. Clearcutting involved felling all the trees in a particular area. It was encouraged by tax laws that exempted all standing timber from taxation if 70% of trees in the area were harvested.[52] Selection logging, by contrast, called for the removal 25% to 50% of mature trees in the hopes that the remaining trees would allow for future growth and reseeding. This method, however, encouraged growth of other tree species, converting redwood forests into mixed forests of redwood, grand fir, Sitka spruce, and western hemlock. Moreover, the trees left standing were often felled by windthrow; that is, they were often blown over by the wind.

The coast redwood is naturalized in New Zealand, notably at Whakarewarewa Forest, Rotorua. Redwood has been grown in New Zealand plantations for more than 100 years, and those planted in New Zealand have higher growth rates than those in California, mainly because of even rainfall distribution through the year.

Other areas of successful cultivation outside of the native range include Great Britain, Italy, France, Haida Gwaii, middle elevations of Hawaii, Hogsback in South Africa, the Knysna Afromontane forests in the Western Cape, Grootvadersbosch Forest Reserve near Swellendam, South Africa and the Tokai Arboretum on the slopes of Table Mountain above Cape Town, a small area in central Mexico (Jilotepec), and the southeastern United States from eastern Texas to Maryland. It also does well in the Pacific Northwest (Oregon, Washington, and British Columbia), far north of its northernmost native range in southwestern Oregon. Coast redwood trees were used in a display at Rockefeller Center and then given to Longhouse Reserve in East Hampton, Long Island, New York, and these have now been living there for over twenty years and have survived at 2 °F (−17 °C).

This fast-growing tree can be grown as an ornamental specimen in those large parks and gardens that can accommodate its massive size. It has gained the Royal Horticultural Society's Award of Garden Merit.

Statistics

Dried resin
The foliage of an "albino" exhibiting lack of chlorophyll
"Albinos" lack chlorophyll

Fairly solid evidence indicates that coast redwoods were the world's largest trees before logging, with numerous historical specimens reportedly over 122 m (400 ft). The theoretical maximum potential height of coast redwoods is thought to be limited to between 122 and 130 m (400 and 427 ft), as evapotranspiration is insufficient to transport water to leaves beyond this range. Further studies have indicated that this maximum requires fog, which is prevalent in these trees' natural environment.

A tree reportedly 114.3 m (375 ft) in length was felled in Sonoma County by the Murphy Brothers saw mill in the 1870s, another claimed to be 115.8 m (380 ft) and 7.9 m (26 ft) in diameter was cut down near Eureka in 1914, and the Lindsey Creek tree was documented to have a height of 120 m (390 ft) when it was uprooted and felled by a storm in 1905. A tree reportedly 129.2 m (424 ft) tall was felled in November 1886 by the Elk River Mill and Lumber Company in Humboldt County, yielding 79,736 marketable board feet from 21 cuts. In 1893, a Redwood cut at the Eel River, near Scotia, reportedly measured 130.1 m (427 ft) in length, and 23.5 m (77 ft) in girth. However, limited evidence corroborates these historical measurements.

Today, trees over 60 m (200 ft) are common, and many are over 90 m (300 ft). The current tallest tree is the Hyperion tree, measuring 115.61 m (379.3 ft). The tree was discovered in Redwood National Park during mid-2006 by Chris Atkins and Michael Taylor, and is thought to be the world's tallest living organism. The previous record holder was the Stratosphere Giant in Humboldt Redwoods State Park at 112.84 m (370.2 ft) (as measured in 2004). Until it fell in March 1991, the "Dyerville Giant" was the record holder. It, too, stood in Humboldt Redwoods State Park and was 113.4 m (372 ft) high and estimated to be 1,600 years old. This fallen giant has been preserved in the park.

The largest known living coast redwood is Grogan's Fault, discovered in 2014 by Chris Atkins and Mario Vaden in Redwood National Park, with a main trunk volume of at least 1,084.5 cubic meters (38,299 cu ft) Other high-volume coast redwoods include Iluvatar, with a main trunk volume of 1,033 m3 (36,470 cu ft),[71]: 160  and the Lost Monarch, with a main trunk volume of 988.7 m3 (34,914 cu ft).

Albino redwoods are mutants that cannot manufacture chlorophyll. About 230 examples (including growths and sprouts) are known to exist, reaching heights of up to 20 m (66 ft). These trees survive like parasites, obtaining food from green parent trees. While similar mutations occur sporadically in other conifers, no cases are known of such individuals surviving to maturity in any other conifer species. Recent research news reports that albino redwoods can store higher concentrations of toxic metals, going so far as comparing them to organs or "waste dumps".

List of tallest trees

Heights of the tallest coast redwoods are measured yearly by experts. Even with recent discoveries of tall coast redwoods above 100 m (330 ft), it is likely that no taller trees will be discovered.

Ten tallest Sequoia sempervirens
Rank Name Height Diameter Location
Meters Feet Meters Feet
1 Hyperion 115.85 380.1 4.84 15.9 Redwood National Park
2 Helios 114.58 375.9 4.96 16.3 Redwood National Park
3 Icarus 113.14 371.2 3.78 12.4 Redwood National Park
4 Stratosphere Giant 113.05 370.9 5.18 17.0 Humboldt Redwoods State Park
5 National Geographic 112.71 369.8 4.39 14.4 Redwood National Park
6 Orion 112.63 369.5 4.33 14.2 Redwood National Park
7 Federation Giant 112.62 369.5 4.54 14.9 Humboldt Redwoods State Park
8 Paradox 112.51 369.1 3.90 12.8 Humboldt Redwoods State Park
9 Mendocino 112.32 368.5 4.19 13.7 Montgomery Woods State Natural Reserve
10 Millennium 111.92 367.2 2.71 8.9 Humboldt Redwoods State Park
The Del Norte Titan

Diameter is measured at 1.4 m (4 ft 7 in) above average ground level (at breast height). Details of the precise locations for most of the tallest trees were not announced to the general public for fear of causing damage to the trees and the surrounding habitat. The tallest coast redwood easily accessible to the public is the National Geographic Tree, immediately trailside in the Tall Trees Grove of Redwood National Park.

List of largest trees

The following list shows the largest S. sempervirens by volume known as of 2001.

Five largest Sequoia sempervirens Rank Name Height Diameter Trunk volume Location Meters Feet Meters Feet Cubic meters Cubic feet 1 Del Norte Titan 93.6 307 7.23 23.7 1,045 36,900 Jedediah Smith Redwoods State Park 2 Iluvatar 91.4 300 6.14 20.1 1,033 36,500 Prairie Creek Redwoods State Park 3 Lost Monarch 97.8 321 7.68 25.2 989 34,900 Jedediah Smith Redwoods State Park 4 Howland Hill Giant 100.3 329 6.02 19.8 951 33,600 Jedediah Smith Redwoods State Park 5 Sir Isaac Newton 94.8 311 7.01 23.0 940 33,000 Prairie Creek Redwoods State Park

Calculating the volume of a standing tree is the practical equivalent of calculating the volume of an irregular cone, and is subject to error for various reasons. This is partly due to technical difficulties in measurement, and variations in the shape of trees and their trunks. Measurements of trunk circumference are taken at only a few predetermined heights up the trunk, and assume that the trunk is circular in cross-section, and that taper between measurement points is even. Also, only the volume of the trunk (including the restored volume of basal fire scars) is taken into account, and not the volume of wood in the branches or roots. The volume measurements also do not take cavities into account. Most coast redwoods with volumes greater than 850 m3 (30,000 cu ft) represent ancient fusions of two or more separate trees, which makes determining whether a coast redwood has a single stem or multiple stems difficult. Starting in 2014, more record-breaking coast redwood trees were discovered. The largest disclosed was a massive redwood called Grogan's Fault/Spartan, which has been measured to have a volume of 38,300 cubic feet. In 2021 during a meeting presentation titled Redwoods 101 run by Henry Cowell Redwoods State Park, an even larger redwood was revealed, allegedly surpassed only by 3 giant sequoias in size. This tree is popularly known as 'Hail Storm', and has a volume of 44,750 cubic feet.

Details of the precise locations for most of the tallest trees were not announced to the general public for fear of causing damage to the trees and the surrounding habitat. The largest coast redwood easily accessible to the public is Iluvatar, which stands prominently about 5 meters (16 ft) to the southeast of the Foothill Trail of Prairie Creek Redwoods State Park.

Canopy Layers

Clouded Salamander, Aneides ferreus

Redwood canopy soil forms from leaf and organic material litter shedding from upper portions of the tree, accumulating and decomposing on larger branches. These clusters of soil require a lot of hydration, but they have an incredible amount of retention once saturated. Redwoods can send roots into these wet soils, providing a water source removed from the forest floor. This creates a unique ecosystem within old growth trees full of fungi, vascular plants, and small creatures. An example of a creature that lives there are the Clouded Salamanders that has been discovered up to 40 meters high. Evidence shows they breed and are born in the canopy soil of Redwood trees. Due to the sheer mass height of these trees and the canopy layer, it was almost never explored for the last century. Due to the mass of these trees and the amount of trees in the surrounding area different molds of moss form on these canopies that are called epiphytes. These epiphytes have different characteristics but all of said species are very adaptable to the tough treetop weather and characteristics. After hundreds of years these trees have been shaped into making it possible for these epiphytes to survive through the winter rain and the fall fog.

Other notable examples

  • The Blossom Rock Navigation Trees were two especially tall sequoias located in the Berkeley Hills used as a navigational aid by sailors to avoid the treacherous Blossom Rock near Yerba Buena Island.
  • The Crannell Creek Giant was documented to have a trunk volume of at least 1,744 m3 (61,573 cu ft) – about 32% larger than Grogan's Fault and 17% larger than General Sherman, the current largest tree. It was felled around 1945.
  • The Lindsey Creek tree was documented to have a height of 120 m (390 ft) and a trunk volume of at least 2,500 cubic meters (90,000 cu ft) when it was uprooted and felled by a storm in 1905. If these measurements are to be believed, the Lindsey Creek tree was about 3 m (10 ft) taller than Hyperion, the current tallest tree, 213% larger than Grogan's Fault, and 171% larger than General Sherman.
  • Old Survivor, also known as the Grandfather, is the last remaining old-growth coastal redwood of the redwood forest that populated the Oakland Hills. The tree was seeded sometime between 1549 and 1554.
  • One of the largest redwood stumps ever found, measuring 9.4 m (31 ft) in diameter, is in Oakland, California, the Berkeley Hills, in the Roberts Regional Recreation Area section of Redwood Regional Park.
  • Animism

    From Wikipedia, the free encyclopedia https://en.wikipedia.org/wiki/Animism ...