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Saturday, February 14, 2015

Amazon rainforest



From Wikipedia, the free encyclopedia

Amazon rainforest
Forest
Amazon Manaus forest.jpg
Amazon rainforest, near Manaus, Brazil.
Countries Brazil, Peru, Colombia, Venezuela, Ecuador, Bolivia, Guyana, Suriname, France (French Guiana)
Part of South America
River Amazon River
Area 5,500,000 km2 (2,123,562 sq mi)
Map of the Amazon rainforest ecoregions as delineated by the WWF. Yellow line approximately encloses the Amazon drainage basin. National boundaries shown in black. Satellite image from NASA.

The Amazon rainforest (Portuguese: Floresta Amazônica or Amazônia; Spanish: Selva Amazónica, Amazonía or usually Amazonia; French: Forêt amazonienne; Dutch: Amazoneregenwoud), also known in English as Amazonia or the Amazon Jungle, is a moist broadleaf forest that covers most of the Amazon Basin of South America. This basin encompasses 7,000,000 square kilometres (2,700,000 sq mi), of which 5,500,000 square kilometres (2,100,000 sq mi) are covered by the rainforest. This region includes territory belonging to nine nations. The majority of the forest is contained within Brazil, with 60% of the rainforest, followed by Peru with 13%, Colombia with 10%, and with minor amounts in Venezuela, Ecuador, Bolivia, Guyana, Suriname and French Guiana. States or departments in four nations contain "Amazonas" in their names. The Amazon represents over half of the planet's remaining rainforests, and comprises the largest and most biodiverse tract of tropical rainforest in the world, with an estimated 390 billion individual trees divided into 16,000 species.[1]

Etymology

The name 'Amazon' is said to arise from a war Francisco de Orellana fought with a tribe of Tapuyas and other tribes from South America. The women of the tribe fought alongside the men, as was the custom among the entire tribe.[2] Orellana derived the name Amazonas from the mythical Amazons of Asia described by Herodotus and Diodorus in Greek legends.[2]

History

Natural


Aerial view of the Amazon rainforest, near Manaus

The rainforest likely formed during the Eocene era. It appeared following a global reduction of tropical temperatures when the Atlantic Ocean had widened sufficiently to provide a warm, moist climate to the Amazon basin. The rainforest has been in existence for at least 55 million years, and most of the region remained free of savanna-type biomes at least until the current ice age, when the climate was drier and savanna more widespread.[3][4]

Following the Cretaceous–Paleogene extinction event, the extinction of the dinosaurs and the wetter climate may have allowed the tropical rainforest to spread out across the continent. From 66–34 Mya, the rainforest extended as far south as 45°. Climate fluctuations during the last 34 million years have allowed savanna regions to expand into the tropics. During the Oligocene, for example, the rainforest spanned a relatively narrow band that lay mostly above latitude 15°N. It expanded again during the Middle Miocene, then retracted to a mostly inland formation at the last glacial maximum.[5] However, the rainforest still managed to thrive during these glacial periods, allowing for the survival and evolution of a broad diversity of species.[6]

Aerial view of the Amazon rainforest.

During the mid-Eocene, it is believed that the drainage basin of the Amazon was split along the middle of the continent by the Purus Arch. Water on the eastern side flowed toward the Atlantic, while to the west water flowed toward the Pacific across the Amazonas Basin. As the Andes Mountains rose, however, a large basin was created that enclosed a lake; now known as the Solimões Basin. Within the last 5–10 million years, this accumulating water broke through the Purus Arch, joining the easterly flow toward the Atlantic.[7][8]

There is evidence that there have been significant changes in Amazon rainforest vegetation over the last 21,000 years through the Last Glacial Maximum (LGM) and subsequent deglaciation. Analyses of sediment deposits from Amazon basin paleolakes and from the Amazon Fan indicate that rainfall in the basin during the LGM was lower than for the present, and this was almost certainly associated with reduced moist tropical vegetation cover in the basin.[9] There is debate, however, over how extensive this reduction was. Some scientists argue that the rainforest was reduced to small, isolated refugia separated by open forest and grassland;[10] other scientists argue that the rainforest remained largely intact but extended less far to the north, south, and east than is seen today.[11] This debate has proved difficult to resolve because the practical limitations of working in the rainforest mean that data sampling is biased away from the center of the Amazon basin, and both explanations are reasonably well supported by the available data.

More than half of the dust needed for fertilizing the Amazon rainforest is provided by the Bodélé depression in Sahara. Up to 50 million tonnes per year are windblown across the Atlantic Ocean.[12][13]

Human activity

Based on archaeological evidence from an excavation at Caverna da Pedra Pintada, human inhabitants first settled in the Amazon region at least 11,200 years ago.[14] Subsequent development led to late-prehistoric settlements along the periphery of the forest by AD 1250, which induced alterations in the forest cover.[15]

Geoglyphs on deforested land in the Amazon rainforest, Acre.

For a long time, it was thought that the Amazon rainforest was only ever sparsely populated, as it was impossible to sustain a large population through agriculture given the poor soil. Archeologist Betty Meggers was a prominent proponent of this idea, as described in her book Amazonia: Man and Culture in a Counterfeit Paradise. She claimed that a population density of 0.2 inhabitants per square kilometre (0.52/sq mi) is the maximum that can be sustained in the rainforest through hunting, with agriculture needed to host a larger population.[16] However, recent anthropological findings have suggested that the region was actually densely populated. Some 5 million people may have lived in the Amazon region in AD 1500, divided between dense coastal settlements, such as that at Marajó, and inland dwellers.[17] By 1900 the population had fallen to 1 million and by the early 1980s it was less than 200,000.[17]

The first European to travel the length of the Amazon River was Francisco de Orellana in 1542.[18] The BBC's Unnatural Histories presents evidence that Orellana, rather than exaggerating his claims as previously thought, was correct in his observations that a complex civilization was flourishing along the Amazon in the 1540s. It is believed that the civilization was later devastated by the spread of diseases from Europe, such as smallpox.[19] Since the 1970s, numerous geoglyphs have been discovered on deforested land dating between AD 0–1250, furthering claims about Pre-Columbian civilizations.[20][21] Ondemar Dias is accredited with first discovering the geoglyphs in 1977 and Alceu Ranzi with furthering their discovery after flying over Acre.[19][22] The BBC's Unnatural Histories presented evidence that the Amazon rainforest, rather than being a pristine wilderness, has been shaped by man for at least 11,000 years through practices such as forest gardening and terra preta.[19]

Terra preta (black earth), which is distributed over large areas in the Amazon forest, is now widely accepted as a product of indigenous soil management. The development of this fertile soil allowed agriculture and silviculture in the previously hostile environment; meaning that large portions of the Amazon rainforest are probably the result of centuries of human management, rather than naturally occurring as has previously been supposed.[23] In the region of the Xingu tribe, remains of some of these large settlements in the middle of the Amazon forest were found in 2003 by Michael Heckenberger and colleagues of the University of Florida. Among those were evidence of roads, bridges and large plazas.[24]

Biodiversity


Deforestation in the Amazon rainforest threatens many species of tree frogs, which are very sensitive to environmental changes (pictured: Giant leaf frog)

Scarlet Macaw, which is indigenous to the American tropics.

Wet tropical forests are the most species-rich biome, and tropical forests in the Americas are consistently more species rich than the wet forests in Africa and Asia.[25] As the largest tract of tropical rainforest in the Americas, the Amazonian rainforests have unparalleled biodiversity. One in ten known species in the world lives in the Amazon rainforest.[26] This constitutes the largest collection of living plants and animal species in the world.

The region is home to about 2.5 million insect species,[27] tens of thousands of plants, and some 2,000 birds and mammals. To date, at least 40,000 plant species, 2,200 fishes,[28] 1,294 birds, 427 mammals, 428 amphibians, and 378 reptiles have been scientifically classified in the region.[29] One in five of all the bird species in the world live in the rainforests of the Amazon, and one in five of the fish species live in Amazonian rivers and streams. Scientists have described between 96,660 and 128,843 invertebrate species in Brazil alone.[30]

The biodiversity of plant species is the highest on Earth with one 2001 study finding a quarter square kilometer (62 acres) of Ecuadorian rainforest supports more than 1,100 tree species.[31] A study in 1999 found one square kilometer (247 acres) of Amazon rainforest can contain about 90,790 tonnes of living plants. The average plant biomass is estimated at 356 ± 47 tonnes per hectare.[32] To date, an estimated 438,000 species of plants of economic and social interest have been registered in the region with many more remaining to be discovered or catalogued.[33] The total number of tree species in the region is estimated at 16,000.[1]

The green leaf area of plants and trees in the rainforest varies by about 25% as a result of seasonal changes. Leaves expand during the dry season when sunlight is at a maximum, then undergo abscission in the cloudy wet season. These changes provide a balance of carbon between photosynthesis and respiration.[34]

The rainforest contains several species that can pose a hazard. Among the largest predatory creatures are the black caiman, jaguar, cougar, and anaconda. In the river, electric eels can produce an electric shock that can stun or kill, while piranha are known to bite and injure humans.[35] Various species of poison dart frogs secrete lipophilic alkaloid toxins through their flesh. There are also numerous parasites and disease vectors. Vampire bats dwell in the rainforest and can spread the rabies virus.[36] Malaria, yellow fever and Dengue fever can also be contracted in the Amazon region.

Deforestation


Deforestation in the Mato Grosso state of Brazil, 2007

Deforestation is the conversion of forested areas to non-forested areas. The main sources of deforestation in the Amazon are human settlement and development of the land.[37] Prior to the early 1960s, access to the forest's interior was highly restricted, and the forest remained basically intact.[38] Farms established during the 1960s were based on crop cultivation and the slash and burn method. However, the colonists were unable to manage their fields and the crops because of the loss of soil fertility and weed invasion.[39] The soils in the Amazon are productive for just a short period of time, so farmers are constantly moving to new areas and clearing more land.[39] These farming practices led to deforestation and caused extensive environmental damage.[40] Deforestation is considerable, and areas cleared of forest are visible to the naked eye from outer space.

In the 1970s construction began on the Trans-Amazonian highway. This highway represented a major threat to the Amazon rainforest.[41] Fortunately for the rainforest, the highway has not been completed, hereby reducing the environmental damage.

Between 1991 and 2000, the total area of forest lost in the Amazon rose from 415,000 to 587,000 square kilometres (160,000 to 227,000 sq mi), with most of the lost forest becoming pasture for cattle.[42] Seventy percent of formerly forested land in the Amazon, and 91% of land deforested since 1970, is used for livestock pasture.[43][44] Currently, Brazil is the second-largest global producer of soybeans after the United States. New research however, conducted by Leydimere Oliveira et al., has shown that the more rainforest is logged in the Amazon, the less precipitation reaches the area and so the lower the yield per hectare becomes. So despite the popular perception, there has been no economical advantage for Brazil from logging rainforest zones and converting these to pastoral fields.[45]

The needs of soy farmers have been used to justify many of the controversial transportation projects that are currently developing in the Amazon. The first two highways successfully opened up the rainforest and led to increased settlement and deforestation. The mean annual deforestation rate from 2000 to 2005 (22,392 km2 or 8,646 sq mi per year) was 18% higher than in the previous five years (19,018 km2 or 7,343 sq mi per year).[46] Deforestation has declined significantly in the Brazilian Amazon since 2004.[47]

Conservation and climate change

Amazon rainforest

Environmentalists are concerned about loss of biodiversity that will result from destruction of the forest, and also about the release of the carbon contained within the vegetation, which could accelerate global warming. Amazonian evergreen forests account for about 10% of the world's terrestrial primary productivity and 10% of the carbon stores in ecosystems[48]—of the order of 1.1 × 1011 metric tonnes of carbon.[49] Amazonian forests are estimated to have accumulated 0.62 ± 0.37 tons of carbon per hectare per year between 1975 and 1996.[49]

One computer model of future climate change caused by greenhouse gas emissions shows that the Amazon rainforest could become unsustainable under conditions of severely reduced rainfall and increased temperatures, leading to an almost complete loss of rainforest cover in the basin by 2100.[50][51] However, simulations of Amazon basin climate change across many different models are not consistent in their estimation of any rainfall response, ranging from weak increases to strong decreases.[52] The result indicates that the rainforest could be threatened though the 21st century by climate change in addition to deforestation.

In 1989, environmentalist C.M. Peters and two colleagues stated there is economic as well as biological incentive to protecting the rainforest. One hectare in the Peruvian Amazon has been calculated to have a value of $6820 if intact forest is sustainably harvested for fruits, latex, and timber; $1000 if clear-cut for commercial timber (not sustainably harvested); or $148 if used as cattle pasture.[53]

As indigenous territories continue to be destroyed by deforestation and ecocide, such as in the Peruvian Amazon[54] indigenous peoples' rainforest communities continue to disappear, while others, like the Urarina continue to struggle to fight for their cultural survival and the fate of their forested territories. Meanwhile, the relationship between non-human primates in the subsistence and symbolism of indigenous lowland South American peoples has gained increased attention, as has ethno-biology and community-based conservation efforts.

From 2002 to 2006, the conserved land in the Amazon rainforest has almost tripled and deforestation rates have dropped up to 60%. About 1,000,000 square kilometres (250,000,000 acres) have been put onto some sort of conservation, which adds up to a current amount of 1,730,000 square kilometres (430,000,000 acres).[55]
A 2009 study found that a 4 °C rise in global temperatures by 2100 would kill 85% of the Amazon rainforest while a temperature rise of 3 °C would kill some 75% of the Amazon.[56]

Remote sensing


This image reveals how the forest and the atmosphere interact to create a uniform layer of "popcorn-shaped" cumulus clouds.

The use of remotely sensed data is dramatically improving conservationists' knowledge of the Amazon Basin. Given the objectivity and lowered costs of satellite-based land cover analysis, it appears likely that remote sensing technology will be an integral part of assessing the extent and damage of deforestation in the basin.[57] Furthermore, remote sensing is the best and perhaps only possible way to study the Amazon on a large-scale.[58]

The use of remote sensing for the conservation of the Amazon is also being used by the indigenous tribes of the basin to protect their tribal lands from commercial interests. Using handheld GPS devices and programs like Google Earth, members of the Trio Tribe, who live in the rainforests of southern Suriname, map out their ancestral lands to help strengthen their territorial claims.[59] Currently, most tribes in the Amazon do not have clearly defined boundaries, making it easier for commercial ventures to target their territories.

To accurately map the Amazon's biomass and subsequent carbon related emissions, the classification of tree growth stages within different parts of the forest is crucial. In 2006 Tatiana Kuplich organized the trees of the Amazon into four categories: (1) mature forest, (2) regenerating forest [less than three years], (3) regenerating forest [between three and five years of regrowth], and (4) regenerating forest [eleven to eighteen years of continued development].[60] The researcher used a combination of Synthetic aperture radar (SAR) and Thematic Mapper (TM) to accurately place the different portions of the Amazon into one of the four classifications.

Impact of early 21st century Amazon droughts

In 2005, parts of the Amazon basin experienced the worst drought in one hundred years,[61] and there were indications that 2006 could have been a second successive year of drought.[62] A July 23, 2006 article in the UK newspaper The Independent reported Woods Hole Research Center results showing that the forest in its present form could survive only three years of drought.[63][64] Scientists at the Brazilian National Institute of Amazonian Research argue in the article that this drought response, coupled with the effects of deforestation on regional climate, are pushing the rainforest towards a "tipping point" where it would irreversibly start to die. It concludes that the forest is on the brink of being turned into savanna or desert, with catastrophic consequences for the world's climate.

According to the World Wide Fund for Nature, the combination of climate change and deforestation increases the drying effect of dead trees that fuels forest fires.[65]

In 2010 the Amazon rainforest experienced another severe drought, in some ways more extreme than the 2005 drought. The affected region was approximate 1,160,000 square miles (3,000,000 km2) of rainforest, compared to 734,000 square miles (1,900,000 km2) in 2005. The 2010 drought had three epicenters where vegetation died off, whereas in 2005 the drought was focused on the southwestern part. The findings were published in the journal Science. In a typical year the Amazon absorbs 1.5 gigatons of carbon dioxide; during 2005 instead 5 gigatons were released and in 2010 8 gigatons were released.[66][67]

Phytoplankton



From Wikipedia, the free encyclopedia


Diatoms are one of the most common types of phytoplankton.

Phytoplankton /ˌftˈplæŋktən/ are the autotrophic components of the plankton community and a key factor of oceans, seas and freshwater basins ecosystems. The name comes from the Greek words φυτόν (phyton), meaning "plant", and πλαγκτός (planktos), meaning "wanderer" or "drifter".[1] Most phytoplankton are too small to be individually seen with the unaided eye. However, when present in high enough numbers, some varieties may be noticeable as a green discoloration of the water; this is due to the presence of chlorophyll within their cells, and in some species also due to the presence of accessory pigments such as phycobiliproteins, xanthophylls, etc.).

Ecology

Phytoplankton come in many shapes and sizes.
Phytoplankton are the foundation of the oceanic food chain.
When two currents collide (here the Oyashio and Kuroshio currents) they create eddies. Phytoplankton concentrates along the boundaries of the eddies, tracing the motion of the water.
Algal bloom off south England.

Phytoplankton are photosynthesizing microscopic organisms that inhabit the upper sunlit layer of almost all oceans and bodies of fresh water. They are agents for "primary production," the creation of organic compounds from carbon dioxide dissolved in the water, a process that sustains the aquatic food web.[2] Phytoplankton obtain energy through the process of photosynthesis and must therefore live in the well-lit surface layer (termed the euphotic zone) of an ocean, sea, lake, or other body of water. Phytoplankton account for half of all photosynthetic activity on Earth.[3][4] Their cumulative energy fixation in carbon compounds (primary production) is the basis for the vast majority of oceanic and also many freshwater food webs (chemosynthesis is a notable exception). The effects of anthropogenic warming on the global population of phytoplankton is an area of active research.
Changes in the vertical stratification of the water column, the rate of temperature-dependent biological reactions, and the atmospheric supply of nutrients are expected to have important effects on future phytoplankton productivity.[5][6] Additionally, changes in the mortality of phytoplankton due to rates of zooplankton grazing may be significant. As a side note, one of the more remarkable food chains in the ocean – remarkable because of the small number of links – is that of phytoplankton-feeding krill (a crustacean similar to a tiny shrimp) feeding baleen whales.

Phytoplankton are also crucially dependent on minerals. These are primarily macronutrients such as nitrate, phosphate or silicic acid, whose availability is governed by the balance between the so-called biological pump and upwelling of deep, nutrient-rich waters. However, across large regions of the World Ocean such as the Southern Ocean, phytoplankton are also limited by the lack of the micronutrient iron. This has led to some scientists advocating iron fertilization as a means to counteract the accumulation of human-produced carbon dioxide (CO2) in the atmosphere.[7] Large-scale experiments have added iron (usually as salts such as iron sulphate) to the oceans to promote phytoplankton growth and draw atmospheric CO2 into the ocean. However, controversy about manipulating the ecosystem and the efficiency of iron fertilization has slowed such experiments.[8]

Phytoplankton depend on other substances to survive as well. In particular, Vitamin B is crucial. Areas in the ocean have been identified as having a major lack of Vitamin B, and correspondingly, phytoplankton.[9]

While almost all phytoplankton species are obligate photoautotrophs, there are some that are mixotrophic and other, non-pigmented species that are actually heterotrophic (the latter are often viewed as zooplankton). Of these, the best known are dinoflagellate genera such as Noctiluca and Dinophysis, that obtain organic carbon by ingesting other organisms or detrital material.

The term phytoplankton encompasses all photoautotrophic microorganisms in aquatic food webs. Phytoplankton serve as the base of the aquatic food web, providing an essential ecological function for all aquatic life. However, unlike terrestrial communities, where most autotrophs are plants, phytoplankton are a diverse group, incorporating protistan eukaryotes and both eubacterial and archaebacterial prokaryotes. There are about 5,000 known species of marine phytoplankton.[10] There is uncertainty in how such diversity has evolved in an environment where competition for only a few resources would suggest limited potential for niche differentiation.[11]

In terms of numbers, the most important groups of phytoplankton include the diatoms, cyanobacteria and dinoflagellates, although many other groups of algae are represented. One group, the coccolithophorids, is responsible (in part) for the release of significant amounts of dimethyl sulfide (DMS) into the atmosphere. DMS is oxidized to form sulfate which, in areas where ambient aerosol particle concentrations are low, can contribute to the population of cloud condensation nuclei, mostly leading to increased cloud cover and cloud albedo according to the so-called CLAW Hypothesis.[12] [13] In oligotrophic oceanic regions such as the Sargasso Sea or the South Pacific Gyre, phytoplankton is dominated by the small sized cells, called picoplankton, mostly composed of cyanobacteria (Prochlorococcus, Synechococcus) and picoeucaryotes such as Micromonas.

Growth Strategy

In the early twentieth century, Alfred C. Redfield found the similarity of the phytoplankton’s elemental composition to the major dissolved nutrients in the deep ocean.[14] Redfield proposed that the ratio of nitrogen to phosphorus (16:1) in the ocean was controlled by the phytoplankton’s requirements which subsequently release nitrogen and phosphorus as they are remineralized. This so-called “Redfield ratio” in describing stoichiometry of phytoplankton and seawater has become a fundamental principle to understand the marine ecology, biogeochemistry and phytoplankton evolution.[15] However, Redfield ratio is not a universally value and it may diverge due to the changes in exogenous nutrient delivery [16] and microbial metabolisms in the ocean, such as nitrogen fixation, denitrification and anammox.

The dynamic stoichiometry shown in unicellular algae reflects their capability to stockpile nutrients in internal pool, shift between enzymes with various nutrient requirements and alter osmolyte composition.[17][18] Different cellular components have their own unique stoichiometry characteristics,[15] for instance, resource (light or nutrients) acquisition machinery such as proteins and chlorophyll contain high concentration of nitrogen but low in phosphorus. Meanwhile, growth machinery such as ribosomal RNA contains high nitrogen and phosphorus concentration.

Based on allocation of resources, phytoplankton is classified into three different growth strategies, namely survivalist, bloomer [19] and generalist. Survivalist phytoplankton has high ratio of N:P (>30) and contains numerous resource-acquisition machinery to sustain growth under scarce resources. Bloomer phytoplankton has low N:P ratio (<10), contains high proportion of growth machinery and adapted to exponential growth. Generalist phytoplankton has similar N:P to Redfield ratio and contain relatively equal resource-acquisition and growth machinery.

Environmental controversy

A 2010 study published in Nature reported that marine phytoplankton have declined substantially in the world's oceans over the past century. Phytoplankton concentrations in surface waters were estimated to have decreased by about 40% since 1950 alone, at a rate of around 1% per year, possibly in response to ocean warming.[20][21] The study generated debate among scientists and led to several communications and criticisms, also published in Nature.[22][23][24][25] The role of cadmium has been reviewed.[26]

Signs that such a large decline in phytoplankton has not occurred include not observing a comparable percentage decline in fish species which feed on phytoplankton.[27] Another global ocean primary productivity study found a net increase in phytoplankton, as judged from measured chlorophyll, when comparing observations in 1998–2002 to those conducted during a prior mission in 1979–1986.[28] The airborne fraction of CO2 from human emissions, the percentage neither sequestered by photosynthetic life on land and sea nor absorbed in the oceans abiotically, has been almost constant over the past century, and that suggests a moderate upper limit on how much a component of the carbon cycle as large as phytoplankton may have declined, if such declined in recent decades.[29] In the example of the northeast Atlantic, a case where chlorophyll measurements extend particularly far back, the location of the Continuous Plankton Recorder (CPR) survey, there was net increase over a 1948 to 2002 period examined.[30] During 1998–2005, global ocean net primary productivity rose during 1998 followed by primarily decline during the rest of that period, although still slightly higher at its end than at its start.[31] Using six different climate model simulations, a large multi-university study of ocean ecosystems predicts, by 2050 A.D., "a global increase in primary production of 0.7% at the low end to 8.1% at the high end," although with "very large regional differences" including "a contraction of the highly productive marginal sea ice biome by 42% in the Northern Hemisphere and 17% in the Southern Hemisphere."[32]

Aquaculture

Phytoplankton are a key food item in both aquaculture and mariculture. Both utilize phytoplankton as food for the animals being farmed. In mariculture, the phytoplankton is naturally occurring and is introduced into enclosures with the normal circulation of seawater. In aquaculture, phytoplankton must be obtained and introduced directly. The plankton can either be collected from a body of water or cultured, though the former method is seldom used. Phytoplankton is used as a foodstock for the production of rotifers,[33] which are in turn used to feed other organisms. Phytoplankton is also used to feed many varieties of aquacultured molluscs, including pearl oysters and giant clams.

The production of phytoplankton under artificial conditions is itself a form of aquaculture. Phytoplankton is cultured for a variety of purposes, including foodstock for other aquacultured organisms,[33] a nutritional supplement for captive invertebrates in aquaria. Culture sizes range from small-scale laboratory cultures of less than 1L to several tens of thousands of liters for commercial aquaculture.[33] Regardless of the size of the culture, certain conditions must be provided for efficient growth of plankton. The majority of cultured plankton is marine, and seawater of a specific gravity of 1.010 to 1.026 may be used as a culture medium. This water must be sterilized, usually by either high temperatures in an autoclave or by exposure to ultraviolet radiation, to prevent biological contamination of the culture. Various fertilizers are added to the culture medium to facilitate the growth of plankton. A culture must be aerated or agitated in some way to keep plankton suspended, as well as to provide dissolved carbon dioxide for photosynthesis. In addition to constant aeration, most cultures are manually mixed or stirred on a regular basis. Light must be provided for the growth of phytoplankton. The colour temperature of illumination should be approximately 6,500 K, but values from 4,000 K to upwards of 20,000 K have been used successfully. The duration of light exposure should be approximately 16 hours daily; this is the most efficient artificial day length.[33]

Lie point symmetry

From Wikipedia, the free encyclopedia https://en.wikipedia.org/wiki/Lie_point_symmetry     ...