Search This Blog

Sunday, March 1, 2015

Rings of Saturn


From Wikipedia, the free encyclopedia


The full set of rings, imaged as Saturn eclipsed the Sun from the vantage of the Cassini spacecraft, 1,200,000 km (746,000 mi) distant, on 19 July 2013 (brightness is exaggerated). Earth is visible as a "pale blue dot" at about the 4 o'clock position, between the G Ring and outer E Ring.

Simulated image using color to present radio occultation-derived information on particle sizes. Radio signals of 0.94, 3.6, and 13 cm wavelengths were sent from Cassini through the rings to Earth. The attenuation of each signal reflects the abundance of particles of sizes similar to the wavelength and larger. Purple (much of the B Ring and the inner A ring) indicates few particles smaller than 5 cm are present, i.e., similar attenuation of all signals. Green and blue (the C Ring and outer A Ring) indicate particles smaller than 5 cm and 1 cm, respectively, are common. White areas of the B Ring are densest and transmit too little signal for size estimation. Other evidence indicates all three rings have a broad range of particle sizes, up to meters across.

Voyager 2 view of Saturn casting a shadow across its rings. Four satellites and ring spokes are visible.

The rings of Saturn are the most extensive planetary ring system of any planet in the Solar System. They consist of countless small particles, ranging in size from micrometres to metres,[1] that orbit about Saturn. The ring particles are made almost entirely of water ice, with a trace component of rocky material. There is still no consensus as to their mechanism of formation; some features of the rings suggest a relatively recent origin, but theoretical models indicate they are likely to have formed early in the Solar System's history.[2]

Although reflection from the rings increases Saturn's brightness, they are not visible from Earth with unaided vision. In 1610, the year after Galileo Galilei first turned a telescope to the sky, he became the very first person to observe Saturn's rings, though he could not see them well enough to discern their true nature. In 1655, Christiaan Huygens was the first person to describe them as a disk surrounding Saturn.[3] Although many people think of Saturn's rings as being made up of a series of tiny ringlets (a concept that goes back to Laplace),[3] true gaps are few. It is more correct to think of the rings as an annular disk with concentric local maxima and minima in density and brightness.[2] On the scale of the clumps within the rings there is much empty space.

The rings have numerous gaps where particle density drops sharply: two opened by known moons embedded within them, and many others at locations of known destabilizing orbital resonances with Saturn's moons. Other gaps remain unexplained. Stabilizing resonances, on the other hand, are responsible for the longevity of several rings, such as the Titan Ringlet and the G Ring.

Well beyond the main rings is the Phoebe ring, which is tilted at an angle of 27 degrees to the other rings and, like Phoebe, orbits in retrograde fashion.

History

Galileo's work


Galileo first observed the rings in 1610.

Galileo Galilei was the first to observe the rings of Saturn in 1610 using his telescope, but was unable to identify them as such. He wrote to the Duke of Tuscany that "[t]he planet Saturn is not alone, but is composed of three, which almost touch one another and never move nor change with respect to one another. They are arranged in a line parallel to the zodiac, and the middle one (Saturn itself) is about three times the size of the lateral ones [the edges of the rings]." He also described Saturn as having "ears". In 1612, the plane of the rings was oriented directly at the Earth and the rings appeared to vanish. Mystified, Galileo wondered, "has Saturn swallowed his children?", referencing the myth of Saturn's consumption of his children to prevent them from overthrowing him.[4] They then reappeared in 1613, further confusing Galileo.[5]

Early astronomers used anagrams as a form of commitment scheme to lay claim to new discoveries before their results were ready for publication. Galileo used smaismrmilmepoetaleumibunenugttauiras for Altissimum planetam tergeminum observavi ("I have observed the most distant planet to have a triple form") for discovering the rings of Saturn.[6]

Ring theory and observations


Robert Hooke noted the shadows (a and b) cast by both the globe and the rings on each other in this 1666 drawing of Saturn.

In 1655, Christiaan Huygens became the first person to suggest that Saturn was surrounded by a ring. Using a 50 power refracting telescope that he designed himself, far superior to those available to Galileo, Huygens observed Saturn and wrote that "It [Saturn] is surrounded by a thin, flat, ring, nowhere touching, inclined to the ecliptic".[5] Robert Hooke was another early observer of the rings of Saturn, and noted the casting of shadows on the rings.[7]

In 1675, Giovanni Domenico Cassini determined that Saturn's ring was composed of multiple smaller rings with gaps between them; the largest of these gaps was later named the Cassini Division. This division is a 4,800 km-wide region between the A Ring and B Ring.[8]

In 1787, Pierre-Simon Laplace suggested that the rings were composed of a large number of solid ringlets.[3]

In 1859, James Clerk Maxwell demonstrated that the rings could not be solid or they would become unstable and break apart. He proposed that the rings must be composed of numerous small particles, all independently orbiting Saturn.[9] Later, Sofia Kovalevskaya found that Saturn's rings cannot be liquid ring-shaped bodies.[10] Maxwell's proposal was proven to be correct in 1895 through spectroscopic studies of the rings carried out by James Keeler of Allegheny Observatory and Aristarkh Belopolsky of Pulkovo Observatory.

The rings are named alphabetically in the order they were discovered.[11] The main rings are, working outward from the planet, C, B and A, with the Cassini Division, the largest gap, separating Rings B and A. Several fainter rings were discovered more recently. The D Ring is exceedingly faint and closest to the planet. The narrow F Ring is just outside the A Ring. Beyond that are two far fainter rings named G and E. The rings show a tremendous amount of structure on all scales, some related to perturbations by Saturn's moons, but much unexplained.[11]

Physical characteristics


The dark Cassini Division separates the wide inner B Ring and outer A Ring in this image from the HST's ACS (March 22, 2004). The less prominent C Ring is just inside the B Ring.

The dense main rings extend from 7,000 km to 80,000 km above Saturn's equator (see Major subdivisions of the rings; Saturn's equatorial radius is 60,300 km). With an estimated local thickness of as little as 10 metres[12] and as much as 1 kilometre,[13] they are composed of 99.9 percent pure water ice with a smattering of impurities that may include tholins or silicates.[14] The main rings are primarily composed of particles ranging in size from 1 centimetre to 10 meters.[15]

Based on Voyager observations, the total mass of the rings was estimated to be about 3 x 1019 kg. This is a small fraction of the total mass of Saturn (about 50 ppb) and is just a little less than the moon Mimas.[16] More-recent observations and computer modeling based on Cassini observations show that this may be an underestimate due to clumping in the rings and the mass may be three times this figure.[17] Although the largest gaps in the rings, such as the Cassini Division and Encke Gap, can be seen from Earth, both Voyager spacecraft discovered that the rings have an intricate structure of thousands of thin gaps and ringlets. This structure is thought to arise, in several different ways, from the gravitational pull of Saturn's many moons. Some gaps are cleared out by the passage of tiny moonlets such as Pan,[18] many more of which may yet be discovered, and some ringlets seem to be maintained by the gravitational effects of small shepherd satellites (similar to Prometheus and Pandora's maintenance of the F ring).[citation needed] Other gaps arise from resonances between the orbital period of particles in the gap and that of a more massive moon further out; Mimas maintains the Cassini Division in this manner.[19] Still more structure in the rings consists of spiral waves raised by the inner moons' periodic gravitational perturbations at less disruptive resonances.[citation needed]

Cassini mosaic of Saturn's rings on August 12, 2009, a day after equinox. With the rings pointed at the Sun, illumination is by light reflected off Saturn, except on thicker or out-of-plane sections, like the F Ring.

Data from the Cassini space probe indicate that the rings of Saturn possess their own atmosphere, independent of that of the planet itself. The atmosphere is composed of molecular oxygen gas (O2) produced when ultraviolet light from the Sun interacts with water ice in the rings. Chemical reactions between water molecule fragments and further ultraviolet stimulation create and eject, among other things, O2. According to models of this atmosphere, H2 is also present. The O2 and H2 atmospheres are so sparse that if the entire atmosphere were somehow condensed onto the rings, it would be about one atom thick.[20] The rings also have a similarly sparse OH (hydroxide) atmosphere. Like the O2, this atmosphere is produced by the disintegration of water molecules, though in this case the disintegration is done by energetic ions that bombard water molecules ejected by Saturn's moon Enceladus. This atmosphere, despite being extremely sparse, was detected from Earth by the Hubble Space Telescope.[21]

Cassini space probe view of the unilluminated side of Saturn's rings (May 9, 2007).

Saturn shows complex patterns in its brightness.[22] Most of the variability is due to the changing aspect of the rings,[23][24] and this goes through two cycles every orbit. However, superimposed on this is variability due to the eccentricity of the planet's orbit that causes the planet to display brighter oppositions in the northern hemisphere than it does in the southern.[25]

In 1980, Voyager 1 made a fly-by of Saturn that showed the F-ring to be composed of three narrow rings that appeared to be braided in a complex structure; it is now known that the outer two rings consist of knobs, kinks and lumps that give the illusion of braiding, with the less bright third ring lying inside them.[citation needed]

New images of the rings taken around the 11 August 2009 equinox of Saturn by NASA's Cassini spacecraft have shown that the rings extend significantly out of the nominal ring plane in a few places. This displacement reaches as much as 4 km (2.5 mi) at the border of the Keeler Gap, due to the out-of-plane orbit of Daphnis, the moon that creates the gap.[26]

Formation of main rings

Saturn's rings may be very old, dating to the formation of Saturn itself. There are two main theories regarding the origin of Saturn's inner rings. One theory, originally proposed by Édouard Roche in the 19th century, is that the rings were once a moon of Saturn (named Veritas, a Roman goddess who hid in a well) whose orbit decayed until it came close enough to be ripped apart by tidal forces (see Roche limit).[27] A variation of this theory is that the moon disintegrated after being struck by a large comet or asteroid.[28] The second theory is that the rings were never part of a moon, but are instead left over from the original nebular material from which Saturn formed.[citation needed]

A 2007 artist impression of the aggregates of icy particles that form the 'solid' portions of Saturn's rings. These elongated clumps are continually forming and dispersing. The largest particles are a few metres across.

A more traditional version of the disrupted moon theory is that the rings are composed of debris from a moon 400 to 600 km in diameter, slightly bigger than Mimas. The last time there were collisions large enough to be likely to disrupt a moon that large was during the Late Heavy Bombardment some four billion years ago.[29]

A more recent variant of this type of theory by R. M. Canup is that the rings could represent part of the remains of the icy mantle of a much larger, Titan-sized, differentiated moon that was stripped of its outer layer as it spiraled into the planet during the formative period when Saturn was still surrounded by a gaseous nebula.[30][31] This would explain the dearth of rocky material within the rings. The rings would initially have been much more massive (~1000 times) and broader than at present; material in the outer portions of the rings would have coalesced into the moons of Saturn out to Tethys, explaining the dearth of rocky material in the composition of most of these moons also.[31] Subsequent collisional or cryovolcanic evolution of Enceladus might then have caused selective loss of ice from this moon, raising its density to its current value of 1.61 g/cm3, compared to values of 1.15 for Mimas and 0.97 for Tethys.[31]

Saturn's rings and Tethys, Hyperion and Prometheus (14 July 2014).

The idea of massive early rings was subsequently extended to explain the formation of Saturn's moons out to Rhea.[32] If the initial massive rings contained chunks of rocky material (>100 km across) as well as ice, these silicate bodies would have accreted more ice and been expelled from the rings, due to gravitational interactions with the rings and tidal interaction with Saturn, into progressively wider orbits. Within the Roche limit, bodies of rocky material are dense enough to accrete additional material, while less dense bodies of ice are not. Once outside the rings, the newly formed moons could have continued to evolve through random mergers. This process may explain the variation in silicate content of Saturn' moons out to Rhea, as well as the trend towards less silicate content closer to Saturn. Rhea would then be the oldest of the moons formed from the primordial rings, with moons closer to Saturn being progressively younger.[32]

The brightness and purity of the water ice in Saturn's rings has been cited as evidence that the rings are much younger than Saturn, perhaps just 100 million years old, as the infall of meteoric dust would have led to darkening of the rings. However, new research indicates that the B Ring may be massive enough to have diluted infalling material and thus avoided substantial darkening over the age of the Solar System. Ring material may be recycled as clumps form within the rings and are then disrupted by impacts. This would explain the apparent youth of some of the material within the rings.[33]

The Cassini UVIS team, led by Larry Esposito, used stellar occultation to discover 13 objects, ranging from 27 metres to 10 km across, within the F ring. They are translucent, suggesting they are temporary aggregates of ice boulders a few metres across. Esposito believes this to be the basic structure of the Saturnian rings, particles clumping together, then being blasted apart.[34][35]

Subdivisions and structures within the rings

The densest parts of the Saturnian ring system are the A and B Rings, which are separated by the Cassini Division (discovered in 1675 by Giovanni Domenico Cassini). Along with the C Ring, which was discovered in 1850 and is similar in character to the Cassini Division, these regions comprise the main rings. The main rings are denser and contain larger particles than the tenuous dusty rings. The latter include the D Ring, extending inward to Saturn's cloud tops, the G and E Rings and others beyond the main ring system. These diffuse rings are characterised as "dusty" because of the small size of their particles (often about a micrometre); their chemical composition is, like the main rings, almost entirely of water ice. The narrow F Ring, just off the outer edge of the A Ring, is more difficult to categorize; parts of it are very dense, but it also contains a great deal of dust-size particles.
Natural-color mosaic of Cassini narrow-angle camera images of the unilluminated side of Saturn's D, C, B, A and F rings (left to right) taken on May 9, 2007.

The illuminated side of Saturn's rings with the major subdivisions labeled

Tabulated data

Notes:
(1) distance is to centre of gaps, rings and ringlets that are narrower than 1,000 km
(2) unofficial name
(3) Names as designated by the International Astronomical Union, unless otherwise noted. Broader separations between named rings are termed divisions, while narrower separations within named rings are called gaps.
(4) Data mostly from the Gazetteer of Planetary Nomenclature, a NASA factsheet and several papers.[36][37][38]

Major subdivisions of the rings

Name(3) Distance from Saturn's
center (km)(4)
Width (km)(4) Named after
D Ring 66,900   –  74,510 7,500
C Ring 74,658   –   92,000 17,500
B Ring 92,000   –  117,580 25,500
Cassini Division 117,580   –   122,170 4,700 Giovanni Cassini
A Ring 122,170   –   136,775 14,600
Roche Division 136,775   –   139,380 2,600 Édouard Roche
F Ring 140,180 (1) 30   –  500
Janus/Epimetheus Ring(2) 149,000   –  154,000 5,000 Janus and Epimetheus
G Ring 166,000   –  175,000 9,000
Methone Ring Arc(2) 194,230 ? Methone
Anthe Ring Arc(2) 197,665 ? Anthe
Pallene Ring(2) 211,000   –  213,500 2,500 Pallene
E Ring 180,000   –  480,000 300,000
Phoebe Ring ~4,000,000 – >13,000,000 Phoebe  

Structures within the C Ring

Name(3) Distance from Saturn's
center (km)(4)
Width (km)(4) Named after
Colombo Gap 77,870 (1) 150 Giuseppe "Bepi" Colombo
Titan Ringlet 77,870 (1) 25 Titan, moon of Saturn
Maxwell Gap 87,491 (1) 270 James Clerk Maxwell
Maxwell Ringlet 87,491 (1) 64 James Clerk Maxwell
Bond Gap 88,700 (1) 30 William Cranch Bond and George Phillips Bond
1.470RS Ringlet 88,716 (1) 16 its radius
1.495RS Ringlet 90,171 (1) 62 its radius
Dawes Gap 90,210 (1) 20 William Rutter Dawes

Structures within the Cassini Division

Name(3) Distance from Saturn's
center (km)(4)
Width (km)(4) Named after
Huygens Gap 117,680 (1) 285–400 Christiaan Huygens
Huygens Ringlet 117,848 (1) ~17 Christiaan Huygens
Herschel Gap 118,234 (1) 102 William Herschel
Russell Gap 118,614 (1) 33 Henry Norris Russell
Jeffreys Gap 118,950 (1) 38 Harold Jeffreys
Kuiper Gap 119,405 (1) 3 Gerard Kuiper
Laplace Gap 119,967 (1) 238 Pierre-Simon Laplace
Bessel Gap 120,241 (1) 10 Friedrich Bessel
Barnard Gap 120,312 (1) 13 Edward Emerson Barnard

Structures within the A Ring

Name(3) Distance from Saturn's
center (km)(4)
Width (km)(4) Named after
Encke Gap 133,589 (1) 325 Johann Encke
Keeler Gap 136,505 (1) 35 James Keeler

Oblique (4 degree angle) Cassini images of Saturn's C, B, and A rings (left to right; the F ring is faintly visible in the full size upper image if viewed at sufficient brightness). Upper image: natural color mosaic of Cassini narrow-angle camera photos of the illuminated side of the rings taken on December 12, 2004. Lower image: simulated view constructed from a radio occultation observation conducted on May 3, 2005. Color in the lower image is used to represent information about ring particle sizes (see the caption of the article's second image for an explanation).

D Ring


A Cassini image of Saturn's D ring processed to show faint ripples within it; the much brighter C ring appears in the upper left.

The D ring is the innermost ring, and is very faint. In 1980, Voyager 1 detected within this ring three ringlets designated D73, D72 and D68, with D68 being the discrete ringlet nearest to Saturn. Some 25 years later, Cassini images showed that D72 had become significantly broader and more diffuse, and had moved planetward by 200 kilometres.[40]

Present in the D ring is a finescale structure with waves 30 kilometres apart. First seen in the gap between the C ring and D73,[40] the structure was found during Saturn's 2009 equinox to extend a radial distance of 19000 km from the D ring to the inner edge of the B ring.[41][42] The waves are interpreted as a spiral pattern of vertical corrugations of 2 to 20 m amplitude;[43] the fact that the period of the waves is decreasing over time (from 60 km in 1995 to 30 km by 2006) allows a deduction that the pattern may have originated in late 1983 with the impact of a cloud of debris (with a mass of ~1012 kg) from a disrupted comet that tilted the rings out of the equatorial plane.[40][41][44] A similar spiral pattern in Jupiter's main ring has been attributed to a perturbation caused by impact of material from Comet Shoemaker-Levy 9 in 1994.[41][45][46]

C Ring


View of the outer C Ring; the Maxwell Gap with the Maxwell Ringlet on its right side are above and right of center. The Bond Gap is above a broad light band towards the upper right; the Dawes Gap is within a dark band just below the upper right corner.

The C Ring is a wide but faint ring located inward of the B Ring. It was discovered in 1850 by William and George Bond, though William R. Dawes and Johann Galle also saw it independently. William Lassell termed it the "Crepe Ring" because it seemed to be composed of darker material than the brighter A and B Rings.[47]

Its vertical thickness is estimated at 5 metres, its mass at around 1.1 × 1018 kilograms, and its optical depth varies from 0.05 to 0.12.[citation needed] That is, between 5 and 12 percent of light shining perpendicularly through the ring is blocked, so that when seen from above, the ring is close to transparent. The 30-kilometer wavelength spiral corrugations first seen in the D Ring were observed during Saturn's equinox of 2009 to extend throughout the C Ring (see above).

Colombo Gap and Titan Ringlets

The Colombo Gap lies in the inner C Ring. Within the gap lies the bright but narrow Colombo Ringlet, centered at 77,883 kilometres from Saturn's center, which is slightly elliptical rather than circular. This ringlet is also called the Titan Ringlet as it is governed by an orbital resonance with the moon Titan.[48] At this location within the rings, the length of a ring particle's apsidal precession is equal to the length of Titan's orbital motion, so that the outer end of this eccentric ringlet always points towards Titan.[48]

Maxwell Gap and Ringlet

The Maxwell Gap lies within the outer part of the C Ring. It also contains a dense non-circular ringlet, the Maxwell Ringlet. In many respects this ringlet is similar to the ε ring of Uranus. There are wave-like structures in the middle of both rings. While the wave in the ε ring is thought to be caused by Uranian moon Cordelia, no moon has been discovered in the Maxwell gap as of July 2008.[34]

B Ring


A location at the outer edge of the B Ring, viewed shortly before equinox, where shadows are cast by vertical structures up to 2.5 km high, probably created by unseen embedded moonlets. The Cassini Division is at top.

The B Ring is the largest, brightest, and most massive of the rings. Its thickness is estimated as 5 to 15 metres, its mass at 2.8 × 1019 kg, and its optical depth varies from 0.4 to 2.5, meaning that 91% of the light passing through some parts of the B Ring is blocked.[citation needed] The B Ring contains a great deal of variation in its density and brightness, nearly all of it unexplained. These are concentric, appearing as narrow ringlets, though the B Ring does not contain any gaps.[citation needed]. In places, the outer edge of the B Ring contains vertical structures deviating up to 2.5 kilometers from the main ring plane.

Spokes

Dark spokes mark the B ring's sunlit side in low phase angle Cassini images. This is a low bitrate video. Full size video with high bitrate of 471 kbit/s;
GIF version (400 × 400 pixels, file size: 2.21 MB)

Until 1980, the structure of the rings of Saturn was explained as being caused exclusively by the action of gravitational forces. Then images from the Voyager spacecraft showed radial features in the B ring, known as spokes,[49] which could not be explained in this manner, as their persistence and rotation around the rings was not consistent with gravitational orbital mechanics.[50] The spokes appear dark in backscattered light, and bright in forward-scattered light (see images in gallery); the transition occurs at a phase angle near 60°. The leading theory regarding the spokes' composition is that they consist of microscopic dust particles suspended away from the main ring by electrostatic repulsion, as they rotate almost synchronously with the magnetosphere of Saturn. The precise mechanism generating the spokes is still unknown, although it has been suggested that the electrical disturbances might be caused by either lightning bolts in Saturn's atmosphere or micrometeoroid impacts on the rings.[51]

The spokes were not observed again until some twenty-five years later, this time by the Cassini space probe. The spokes were not visible when Cassini arrived at Saturn in early 2004. Some scientists speculated that the spokes would not be visible again until 2007, based on models attempting to describe their formation. Nevertheless, the Cassini imaging team kept looking for spokes in images of the rings, and they were next seen in images taken on 5 September 2005.[52]

The spokes appear to be a seasonal phenomenon, disappearing in the Saturnian midwinter and midsummer and reappearing as Saturn comes closer to equinox. Suggestions that the spokes may be a seasonal effect, varying with Saturn's 29.7-year orbit, were supported by their gradual reappearance in the later years of the Cassini mission.[53]

Moonlet

In 2009, during equinox, a moonlet embedded in the B ring was discovered from the shadow it cast. It is estimated to be 400 meters (1,300 ft) in diameter.[54] The moonlet was given the provisional designation S/2009 S 1.

Cassini Division


The Cassini Division imaged from the Cassini spacecraft. The Huygens Gap lies at its right border; the Laplace Gap is towards the center. A number of other, narrower gaps are also present.

The Cassini Division is a 4,800 km (3,000 mi) wide region between the A Ring and B Ring. It was discovered in 1675 by Giovanni Cassini at the Paris Observatory using a refracting telescope that had a 2.5 inch objective lens with a 20 foot long focal length and a 90x magnification.[55][56] From Earth it appears as a thin black gap in the rings. However, Voyager discovered that the gap is itself populated by ring material bearing much similarity to the C Ring.[34] The division may appear bright in views of the unlit side of the rings, since the relatively low density of material allows more light to be transmitted through the thickness of the rings (see second image in gallery).[citation needed]

The inner edge of the Cassini Division is governed by a strong orbital resonance. Ring particles at this location orbit twice for every orbit of the moon Mimas.[57] The resonance causes Mimas' pulls on these ring particles to accumulate, destabilizing their orbits and leading to a sharp cutoff in ring density. Many of the other gaps between ringlets within the Cassini Division, however, are unexplained.[citation needed]

Huygens Gap

The Huygens Gap is located at the inner edge of the Cassini Division. It contains the dense, eccentric Huygens Ringlet in the middle. This ringlet exhibits irregular azimuthal variations of geometrical width and optical depth, which may be caused by the nearby 2:1 resonance with Mimas and the influence of the eccentric outer edge of the B-ring. There is an additional narrow ringlet just outside the Huygens Ringlet.[34]

A Ring

The central ringlet of the A ring's Encke Gap coincides with Pan's orbit, implying its particles oscillate in horseshoe orbits.

The A Ring is the outermost of the large, bright rings. Its inner boundary is the Cassini Division and its sharp outer boundary is close to the orbit of the small moon Atlas. The A Ring is interrupted at a location 22% of the ring width from its outer edge by the Encke Gap. A narrower gap 2% of the ring width from the outer edge is called the Keeler Gap.

The thickness of the A Ring is estimated as 10 to 30 metres, its mass as 6.2 × 1018 kg (about the mass of Hyperion), and its optical depth varies from 0.4 to 1.0.[citation needed]

Similarly to the B Ring, the A Ring's outer edge is maintained by an orbital resonance, in this case the 7:6 resonance with Janus and Epimetheus.[citation needed] Other orbital resonances also excite many spiral density waves in the A Ring (and, to a lesser extent, other rings as well), which account for most of its structure. These waves are described by the same physics that describes the spiral arms of galaxies. Spiral bending waves, also present in the A Ring and also described by the same theory, are vertical corrugations in the ring rather than compression waves.[citation needed]

In April 2014, NASA scientists reported observing the possible formative stage of a new moon near the outer edge of the A Ring.[58][59]

Encke Gap


Waves in the Keeler gap edges induced by the passage of Daphnis.

Near Saturn's equinox, Daphnis and its waves cast shadows on the A Ring.

The Encke Gap is a 325-kilometre-wide gap within the A Ring, centered at a distance of 133,590 kilometres from Saturn's center.[60] It is caused by the presence of the small moon Pan,[61] which orbits within it. Images from the Cassini probe have shown that there are at least three thin, knotted ringlets within the gap.[34] Spiral density waves visible on both sides of it are induced by resonances with nearby moons exterior to the rings, while Pan induces an additional set of spiraling wakes (see image in gallery).[34]

Johann Encke himself did not observe this gap; it was named in honour of his ring observations. The gap itself was discovered by James Edward Keeler in 1888.[47] The second major gap in the A Ring, discovered by Voyager, was named the Keeler Gap in his honor.[62]

The Encke Gap is a gap because it is entirely within the A Ring. There was some ambiguity between the terms gap and division until the IAU clarified the definitions in 2008; before that, the separation was sometimes called the "Encke Division".[63]

Keeler Gap

The Keeler Gap is a 42-kilometre-wide gap in the A Ring, approximately 250 kilometres from the ring's outer edge. The small moon Daphnis, discovered 1 May 2005, orbits within it, keeping it clear.[64] The moon induces waves in the edges of the gap.[34] Because the orbit of Daphnis is slightly inclined to the ring plane, the waves have a component that is perpendicular to the ring plane, reaching a distance of 1.5 km (0.93 mi) "above" the plane.[65][66]

The Keeler gap was discovered by Voyager, and named in honor of the astronomer James Edward Keeler. Keeler had in turn discovered and named the Encke Gap in honor of Johann Encke.[47]

Moonlets


Moonlet Bleriot

Location of the first four moonlets detected in the A ring.

In 2006, four tiny "moonlets" were found in Cassini images of the A Ring.[67] The moonlets themselves are only about a hundred metres in diameter, too small to be seen directly; what Cassini sees are the "propeller"-shaped disturbances the moonlets create, which are several kilometres across. It is estimated that the A Ring contains thousands of such objects. In 2007, the discovery of eight more moonlets revealed that they are largely confined to a 3000 km belt, about 130,000 km from Saturn's center,[68] and by 2008 over 150 propeller moonlets had been detected.[69] One that has been tracked for several years has been nicknamed Bleriot.[70]

Roche Division


The Roche Division (passing through image center) between the A Ring and the narrow F Ring. Atlas can be seen within it. The Encke and Keeler gaps are also visible.

The separation between the A Ring and the F Ring has been named the Roche Division in honor of the French physicist Édouard Roche.[71] The Roche Division should not be confused with the Roche limit, a physical concept that describes when a large object gets so close to a planet (such as Saturn) that the planet's tidal forces will pull it apart.[72] Lying at the outer edge of the main ring system, the Roche Division is in fact close to Saturn's Roche limit, which is why the rings have been unable to accrete into a moon.[73]

Like the Cassini Division, the Roche Division is not empty but contains a sheet of material.[citation needed] The character of this material is similar to the tenuous and dusty D, E, and G Rings.[citation needed] Two locations in the Roche Division have a higher concentration of dust than the rest of the region. These were discovered by the Cassini probe imaging team and were given temporary designations: R/2004 S 1, which lies along the orbit of the moon Atlas; and R/2004 S 2, centered at 138,900 km from Saturn's center, inward of the orbit of Prometheus.[74][75]

F Ring

The shepherd moons Pandora (left) and Prometheus (right) orbit on either side of the F ring; Prometheus is followed by dark channels that it has carved into the inner strands of the ring.

The F Ring is the outermost discrete ring of Saturn and perhaps the most active ring in the Solar System, with features changing on a timescale of hours.[76] It is located 3,000 km beyond the outer edge of the A Ring.[77] The ring was discovered in 1979 by the Pioneer 11 imaging team.[78] It is very thin, just a few hundred kilometres in radial extent, and is held together by two shepherd moons, Prometheus and Pandora, which orbit inside and outside it.[61]

Recent closeup images from the Cassini probe show that the F Ring consists of one core ring and a spiral strand around it.[79] They also show that when Prometheus encounters the ring at its apoapsis, its gravitational attraction creates kinks and knots in the F Ring as the moon 'steals' material from it, leaving a dark channel in the inner part of the ring (see video link and additional F Ring images in gallery). Since Prometheus orbits Saturn more rapidly than the material in the F ring, each new channel is carved about 3.2 degrees in front of the previous one.[76]

In 2008, further dynamism was detected, suggesting that small unseen moons orbiting within the F Ring are continually passing through its narrow core because of perturbations from Prometheus. One of the small moons was tentatively identified as S/2004 S 6.[76]
A mosaic of 107 images showing 255° (about 70%) of the F Ring as it would appear if straightened out. The radial width (top to bottom) is 1,500 km.

Outer rings


The outer rings seen back-illuminated by the Sun

The Anthe Ring Arc. The bright spot is Anthe.

The backlit E ring, with Enceladus silhouetted against it. The moon's south polar jets erupt brightly below it.

Janus/Epimetheus Ring

A faint dust ring is present around the region occupied by the orbits of Janus and Epimetheus, as revealed by images taken in forward-scattered light by the Cassini spacecraft in 2006. The ring has a radial extent of about 5,000 km.[80] Its source is particles blasted off the moons' surfaces by meteoroid impacts, which then form a diffuse ring around their orbital paths.[81]

G Ring

The G Ring (see last image in gallery) is a very thin, faint ring about halfway between the F Ring and the beginning of the E Ring, with its inner edge about 15,000 km inside the orbit of Mimas. It contains a single distinctly brighter arc near its inner edge (similar to the arcs in the rings of Neptune) that extends about one sixth of its circumference, centered on the half-kilometre diameter moonlet Aegaeon, which is held in place by a 7:6 orbital resonance with Mimas.[82][83] The arc is believed to be composed of icy particles up to a few metres in diameter, with the rest of the G Ring consisting of dust released from within the arc. The radial width of the arc is about 250 km, compared to a width of 9,000 km for the G Ring as a whole.[82] The arc is thought to contain matter equivalent to a small icy moonlet about a hundred metres in diameter.[82] Dust released from Aegaeon and other source bodies within the arc by micrometeoroid impacts drifts outward from the arc because of interaction with Saturn's magnetosphere (whose plasma corotates with Saturn's magnetic field, which rotates much more rapidly than the orbital motion of the G Ring). These tiny particles are steadily eroded away by further impacts and dispersed by plasma drag. Over the course of thousands of years the ring gradually loses mass,[84] which is replenished by further impacts on Aegaeon.

Methone Ring Arc

A faint ring arc, first detected in September 2006, covering a longitudinal extent of about 10 degrees is associated with the moon Methone. The material in the arc is believed to represent dust ejected from Methone by micrometeoroid impacts. The confinement of the dust within the arc is attributable to a 14:15 resonance with Mimas (similar to the mechanism of confinement of the arc within the G ring).[85][86] Under the influence of the same resonance, Methone librates back and forth in its orbit with an amplitude of 5° of longitude.

Anthe Ring Arc

A faint ring arc, first detected in June 2007, covering a longitudinal extent of about 20 degrees is associated with the moon Anthe. The material in the arc is believed to represent dust knocked off Anthe by micrometeoroid impacts. The confinement of the dust within the arc is attributable to a 10:11 resonance with Mimas. Under the influence of the same resonance, Anthe drifts back and forth in its orbit over 14° of longitude.[85][86]

Pallene Ring

A faint dust ring shares Pallene's orbit, as revealed by images taken in forward-scattered light by the Cassini spacecraft in 2006.[80] The ring has a radial extent of about 2,500 km. Its source is particles blasted off Pallene's surface by meteoroid impacts, which then form a diffuse ring around its orbital path.[81][86]

E Ring

The E Ring is the second outermost ring and is extremely wide; it consists of many tiny (micron and sub-micron) particles of water ice with silicates, carbon dioxide and ammonia.[87] The E ring is distributed between the orbits of Mimas and Titan.[88] Unlike the other rings, it is composed of microscopic particles rather than macroscopic ice chunks. In 2005, the source of the E Ring's material was determined to be cryovolcanic plumes[89][90] emanating from the "tiger stripes" of the south polar region of the moon Enceladus. Unlike the main rings, the E Ring is more than 2000 kilometers thick and increases with its distance from Enceladus.[88]

Particles of the E-ring tend to accumulate on moons that orbit within it. The equator of the leading hemisphere of Tethys is tinted slightly blue due to infalling material.[91] The trojan moons Telesto, Calypso, Helene and Polydeuces are particularly affected as their orbits move up and down the ring plane. This results in their surfaces being coated with bright material that smooths out features.[92]

Phoebe ring


The Phoebe ring's huge extent dwarfs the main rings. Inset: 24 micrometer Spitzer image of part of the ring

In October 2009, the discovery of a tenuous disk of material just interior to the orbit of Phoebe was reported. The disk was aligned edge-on to Earth at the time of discovery. This disk can be loosely described as another ring. Although very large (the apparent size of two full moons as seen from Earth), the ring is virtually invisible. It was discovered using NASA's infrared Spitzer Space Telescope,[93] and was seen over the entire range of the observations, which extended from 128 to 207 times the radius of Saturn,[94] with calculations indicating that it may extend outward up to 300 Saturn radii and inward to the orbit of Iapetus at 59 Saturn radii.[95] The ring was subsequently studied using the WISE, Herschel and Cassini spacecraft.[96]

Phoebe orbits the planet at an average distance of 215 radii. The ring is about 20 times as thick as the diameter of the planet.[97] Since the ring's particles are presumed to have originated from impacts (micrometeoroid and larger) on Phoebe, they should share its retrograde orbit,[95] which is opposite to the orbital motion of the next inner moon, Iapetus. This ring lies in the plane of Saturn's orbit, or roughly the ecliptic, and thus is tilted 27 degrees from Saturn's equatorial plane and the other rings. Phoebe is inclined by 5° with respect to Saturn's orbit plane (often written as 175°, due to Phoebe's retrograde orbital motion), and its resulting vertical excursions above and below the ring plane agree closely with the ring's observed thickness of 40 Saturn radii.

The existence of the ring was proposed in the 1970s by Steven Soter.[95] The discovery was made by Anne J. Verbiscer and Michael F. Skrutskie (of the University of Virginia) and Douglas P. Hamilton (of the University of Maryland, College Park).[94][98] The three had studied together at Cornell University as graduate students.[99]

Ring material migrates inward due to reemission of solar radiation,[94] and would thus strike the leading hemisphere of Iapetus. Infall of this material causes a slight darkening and reddening of the leading hemisphere of Iapetus (similar to what is seen on the Uranian moons Oberon and Titania) but does not directly create the dramatic two-tone coloration of that moon.[100] Rather, the infalling material initiates a positive feedback thermal self-segregation process of ice sublimation from warmer regions, followed by vapor condensation onto cooler regions. This leaves a dark residue of "lag" material covering most of the equatorial region of Iapetus's leading hemisphere, which contrasts with the bright ice deposits covering the polar regions and most of the trailing hemisphere.[101][102][103]

Possible ring system around Rhea

Saturn's second largest moon Rhea has been hypothesized to have a tenuous ring system of its own consisting of three narrow bands embedded in a disk of solid particles.[104][105] These putative rings have not been imaged, but their existence has been inferred from Cassini observations in November 2005 of a depletion of energetic electrons in Saturn's magnetosphere near Rhea. The Magnetospheric Imaging Instrument (MIMI) observed a gentle gradient punctuated by three sharp drops in plasma flow on each side of the moon in a nearly symmetric pattern. This could be explained if they were absorbed by solid material in the form of an equatorial disk containing denser rings or arcs, with particles perhaps several decimetres to approximately a metre in diameter. A more recent piece of evidence consistent with the presence of Rhean rings is a set of small ultraviolet-bright spots distributed in a line that extends three quarters of the way around the moon's circumference, within 2 degrees of the equator. The spots have been interpreted as the impact points of deorbiting ring material.[106] However, targeted observations by Cassini of the putative ring plane from several angles have turned up nothing, suggesting that another explanation for these enigmatic features is needed.[107]

Gallery


Cassini image mosaic of the unlit side of the inner B Ring (top) and outer C Ring (bottom) near Saturn's equinox, showing multiple views of the shadow of Mimas. The shadow is attenuated by the denser B ring. The Maxwell Gap is below center.

Dark B Ring spokes in a low-phase-angle Cassini image of the rings' unlit side. Left of center, two dark gaps (the larger being the Huygens Gap) and the bright (from this viewing geometry) ringlets to their left comprise the Cassini Division.

Cassini image of the sun-lit side of the rings taken in 2009 at a phase angle of 144°, with bright B Ring spokes.

Pan's motion through the A Ring's Encke Gap induces edge waves and (non-self-propagating) spiraling wakes ahead of and inward of it. The other more tightly wound bands are spiral density waves.

Prometheus near apoapsis carving a dark channel in the F Ring (with older channels to the right). A movie of the process may be viewed at the Cassini Imaging Team website[108] or YouTube[109]

Prometheus (at center) and Pandora are the inner and outer F Ring shepherds.

F ring dynamism, probably due to perturbing effects of small moonlets orbiting close to or through the ring's core.

Saturn's shadow truncates the backlit G Ring and its bright inner arc. A movie showing the arc's orbital motion may be viewed on YouTube[110] or the Cassini Imaging Team website[111]

Flower


From Wikipedia, the free encyclopedia


A poster with flowers or clusters of flowers produced by twelve species of flowering plants from different families

A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Magnoliophyta, also called angiosperms). The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs. Flowers may facilitate outcrossing (fusion of sperm and eggs from different individuals in a population) or allow selfing (fusion of sperm and egg from the same flower). Some flowers produce diaspores without fertilization (parthenocarpy). Flowers contain sporangia and are the site where gametophytes develop. Flowers give rise to fruit and seeds. Many flowers have evolved to be attractive to animals, so as to cause them to be vectors for the transfer of pollen.

In addition to facilitating the reproduction of flowering plants, flowers have long been admired and used by humans to beautify their environment, and also as objects of romance, ritual, religion, medicine and as a source of food.

Morphology


Main parts of a mature flower (Ranunculus glaberrimus)

Diagram of flower parts

Floral parts

The essential parts of a flower can be considered in two parts: the vegetative part, consisting of petals and associated structures in the perianth, and the reproductive or sexual parts. A stereotypical flower consists of four kinds of structures attached to the tip of a short stalk. Each of these kinds of parts is arranged in a whorl on the receptacle. The four main whorls (starting from the base of the flower or lowest node and working upwards) are as follows:

Vegetative (Perianth)

Collectively the calyx and corolla form the perianth (see diagram).
  • Calyx: the outermost whorl consisting of units called sepals; these are typically green and enclose the rest of the flower in the bud stage, however, they can be absent or prominent and petal-like in some species.
  • Corolla: the next whorl toward the apex, composed of units called petals, which are typically thin, soft and colored to attract animals that help the process of pollination.

Reproductive

Reproductive parts of Easter Lily (Lilium longiflorum). 1. Stigma, 2. Style, 3. Stamens, 4. Filament, 5. Petal
  • Androecium (from Greek andros oikia: man's house): the next whorl (sometimes multiplied into several whorls), consisting of units called stamens. Stamens consist of two parts: a stalk called a filament, topped by an anther where pollen is produced by meiosis and eventually dispersed.
  • Gynoecium (from Greek gynaikos oikia: woman's house): the innermost whorl of a flower, consisting of one or more units called carpels. The carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes. These give rise to egg cells. The gynoecium of a flower is also described using an alternative terminology wherein the structure one sees in the innermost whorl (consisting of an ovary, style and stigma) is called a pistil. A pistil may consist of a single carpel or a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen. The supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma. The relationship to the gynoecium on the receptacle is described as hypogynous (beneath a superior ovary), perigynous (surrounding a superior ovary), or epigynous (above inferior ovary).

Structure

Although the arrangement described above is considered "typical", plant species show a wide variation in floral structure.[1] These modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species.

The four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions and/or look like what is typically another part. In some families, like Ranunculaceae, the petals are greatly reduced and in many species the sepals are colorful and petal-like. Other flowers have modified stamens that are petal-like; the double flowers of Peonies and Roses are mostly petaloid stamens.[2] Flowers show great variation and plant scientists describe this variation in a systematic way to identify and distinguish species.

Specific terminology is used to describe flowers and their parts. Many flower parts are fused together; fused parts originating from the same whorl are connate, while fused parts originating from different whorls are adnate; parts that are not fused are free. When petals are fused into a tube or ring that falls away as a single unit, they are sympetalous (also called gamopetalous). Connate petals may have distinctive regions: the cylindrical base is the tube, the expanding region is the throat and the flaring outer region is the limb. A sympetalous flower, with bilateral symmetry with an upper and lower lip, is bilabiate. Flowers with connate petals or sepals may have various shaped corolla or calyx, including campanulate, funnelform, tubular, urceolate, salverform or rotate.

Referring to "fusion," as it is commonly done, appears questionable because at least some of the processes involved may be non-fusion processes. For example, the addition of intercalary growth at or below the base of the primordia of floral appendages such as sepals, petals, stamens and carpels may lead to a common base that is not the result of fusion.[3][4][5]

Left: A normal zygomorphic Streptocarpus flower. Right: An aberrant peloric Streptocarpus flower. Both of these flowers appeared on the Streptocarpus hybrid 'Anderson's Crows' Wings'.

Many flowers have a symmetry. When the perianth is bisected through the central axis from any point, symmetrical halves are produced, forming a radial symmetry. These flowers are also known to be actinomorphic or regular, e.g. rose or trillium. When flowers are bisected and produce only one line that produces symmetrical halves the flower is said to be irregular or zygomorphic, e.g. snapdragon or most orchids.

Flowers may be directly attached to the plant at their base (sessile—the supporting stalk or stem is highly reduced or absent). The stem or stalk subtending a flower is called a peduncle. If a peduncle supports more than one flower, the stems connecting each flower to the main axis are called pedicels.
The apex of a flowering stem forms a terminal swelling which is called the torus or receptacle.

Inflorescence


The familiar calla lily is not a single flower. It is actually an inflorescence of tiny flowers pressed together on a central stalk that is surrounded by a large petal-like bract.

In those species that have more than one flower on an axis, the collective cluster of flowers is termed an inflorescence. Some inflorescences are composed of many small flowers arranged in a formation that resembles a single flower. The common example of this is most members of the very large composite (Asteraceae) group. A single daisy or sunflower, for example, is not a flower but a flower head—an inflorescence composed of numerous flowers (or florets). An inflorescence may include specialized stems and modified leaves known as bracts.

Floral diagrams and floral formulae

A floral formula is a way to represent the structure of a flower using specific letters, numbers and symbols, presenting substantial information about the flower in a compact form. It can represent a taxon, usually giving ranges of the numbers of different organs, or particular species. Floral formulae have been developed in the early 19th century and their use has declined since. Prenner et al. (2010) devised an extension of the existing model to broaden the descriptive capability of the formula.[6] 
The format of floral formulae differs in different parts of the world, yet they convey the same information.[7][8][9][10]
The structure of a flower can also be expressed by the means of floral diagrams. The use of schematic diagrams can replace long descriptions or complicated drawings as a tool for understanding both floral structure and evolution. Such diagrams may show important features of flowers, including the relative positions of the various organs, including the presence of fusion and symmetry, as well as structural details.[7]

Development

A flower develops on a modified shoot or axis from a determinate apical meristem (determinate meaning the axis grows to a set size). It has compressed internodes, bearing structures that in classical plant morphology are interpreted as highly modified leaves.[11] Detailed developmental studies, however, have shown that stamens are often initiated more or less like modified stems (caulomes) that in some cases may even resemble branchlets.[5][12] Taking into account the whole diversity in the development of the androecium of flowering plants, we find a continuum between modified leaves (phyllomes), modified stems (caulomes), and modified branchlets (shoots).[13][14]

Flowering transition

The transition to flowering is one of the major phase changes that a plant makes during its life cycle. The transition must take place at a time that is favorable for fertilization and the formation of seeds, hence ensuring maximal reproductive success. To meet these needs a plant is able to interpret important endogenous and environmental cues such as changes in levels of plant hormones and seasonable temperature and photoperiod changes.[15] Many perennial and most biennial plants require vernalization to flower. The molecular interpretation of these signals is through the transmission of a complex signal known as florigen, which involves a variety of genes, including CONSTANS, FLOWERING LOCUS C and FLOWERING LOCUS T. Florigen is produced in the leaves in reproductively favorable conditions and acts in buds and growing tips to induce a number of different physiological and morphological changes.[16]

The first step of the transition is the transformation of the vegetative stem primordia into floral primordia. This occurs as biochemical changes take place to change cellular differentiation of leaf, bud and stem tissues into tissue that will grow into the reproductive organs. Growth of the central part of the stem tip stops or flattens out and the sides develop protuberances in a whorled or spiral fashion around the outside of the stem end. These protuberances develop into the sepals, petals, stamens, and carpels. Once this process begins, in most plants, it cannot be reversed and the stems develop flowers, even if the initial start of the flower formation event was dependent of some environmental cue.[17] Once the process begins, even if that cue is removed the stem will continue to develop a flower.

Organ development


The ABC model of flower development

The molecular control of floral organ identity determination appears to be fairly well understood in some species. In a simple model, three gene activities interact in a combinatorial manner to determine the developmental identities of the organ primordia within the floral meristem. These gene functions are called A, B and C-gene functions. In the first floral whorl only A-genes are expressed, leading to the formation of sepals. In the second whorl both A- and B-genes are expressed, leading to the formation of petals. In the third whorl, B and C genes interact to form stamens and in the center of the flower C-genes alone give rise to carpels. The model is based upon studies of homeotic mutants in Arabidopsis thaliana and snapdragon, Antirrhinum majus. For example, when there is a loss of B-gene function, mutant flowers are produced with sepals in the first whorl as usual, but also in the second whorl instead of the normal petal formation. In the third whorl the lack of B function but presence of C-function mimics the fourth whorl, leading to the formation of carpels also in the third whorl. See also The ABC Model of Flower Development.

Most genes central in this model belong to the MADS-box genes and are transcription factors that regulate the expression of the genes specific for each floral organ.

Floral function

An example of a "perfect flower", this Crateva religiosa flower has both stamens (outer ring) and a pistil (center).

The principal purpose of a flower is the reproduction of the individual and the species. All flowering plants are heterosporous, producing two types of spores. Microspores are produced by meiosis inside anthers while megaspores are produced inside ovules, inside an ovary. In fact, anthers typically consist of four microsporangia and an ovule is an integumented megasporangium. Both types of spores develop into gametophytes inside sporangia. As with all heterosporous plants, the gametophytes also develop inside the spores (are endosporic).

In the majority of species, individual flowers have both functional carpels and stamens. Botanists describe these flowers as being perfect or bisexual and the species as hermaphroditic. Some flowers lack one or the other reproductive organ and called imperfect or unisexual. If unisex flowers are found on the same individual plant but in different locations, the species is said to be monoecious. If each type of unisex flower is found only on separate individuals, the plant is dioecious.

Flower specialization and pollination

Flowering plants usually face selective pressure to optimize the transfer of their pollen, and this is typically reflected in the morphology of the flowers and the behaviour of the plants. Pollen may be transferred between plants via a number of 'vectors'. Some plants make use of abiotic vectors — namely wind (anemophily) or, much less commonly, water (hydrophily). Others use biotic vectors including insects (entomophily), birds (ornithophily), bats (chiropterophily) or other animals. Some plants make use of multiple vectors, but many are highly specialised.
Cleistogamous flowers are self-pollinated, after which they may or may not open. Many Viola and some Salvia species are known to have these types of flowers.

The flowers of plants that make use of biotic pollen vectors commonly have glands called nectaries that act as an incentive for animals to visit the flower. Some flowers have patterns, called nectar guides, that show pollinators where to look for nectar. Flowers also attract pollinators by scent and color. Still other flowers use mimicry to attract pollinators. Some species of orchids, for example, produce flowers resembling female bees in color, shape, and scent. Flowers are also specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to the bodies of the pollinator when it lands in search of its attractant (such as nectar, pollen, or a mate). In pursuing this attractant from many flowers of the same species, the pollinator transfers pollen to the stigmas—arranged with equally pointed precision—of all of the flowers it visits.

Anemophilous flowers use the wind to move pollen from one flower to the next. Examples include grasses, birch trees, ragweed and maples. They have no need to attract pollinators and therefore tend not to be "showy" flowers. Male and female reproductive organs are generally found in separate flowers, the male flowers having a number of long filaments terminating in exposed stamens, and the female flowers having long, feather-like stigmas. Whereas the pollen of animal-pollinated flowers tends to be large-grained, sticky, and rich in protein (another "reward" for pollinators), anemophilous flower pollen is usually small-grained, very light, and of little nutritional value to animals.

Pollination

Grains of pollen sticking to this bee will be transferred to the next flower it visits

The primary purpose of a flower is reproduction. Since the flowers are the reproductive organs of plant, they mediate the joining of the sperm, contained within pollen, to the ovules — contained in the ovary. Pollination is the movement of pollen from the anthers to the stigma. The joining of the sperm to the ovules is called fertilization. Normally pollen is moved from one plant to another, but many plants are able to self pollinate. The fertilized ovules produce seeds that are the next generation. Sexual reproduction produces genetically unique offspring, allowing for adaptation. Flowers have specific designs which encourages the transfer of pollen from one plant to another of the same species. Many plants are dependent upon external factors for pollination, including: wind and animals, and especially insects. Even large animals such as birds, bats, and pygmy possums can be employed. The period of time during which this process can take place (the flower is fully expanded and functional) is called anthesis. The study of pollination by insects is called anthecology.

Pollen

The types of pollen that most commonly cause allergic reactions are produced by the plain-looking plants (trees, grasses, and weeds) that do not have showy flowers. These plants make small, light, dry pollen grains that are custom-made for wind transport.

The type of allergens in the pollen is the main factor that determines whether the pollen is likely to cause hay fever. For example, pine tree pollen is produced in large amounts by a common tree, which would make it a good candidate for causing allergy. It is, however, a relatively rare cause of allergy because the types of allergens in pine pollen appear to make it less allergenic.[18]

Among North American plants, weeds are the most prolific producers of allergenic pollen.[19] Ragweed is the major culprit, but other important sources are sagebrush, redroot pigweed, lamb’s quarters, Russian thistle (tumbleweed), and English plantain.

There is much confusion about the role of flowers in allergies. For example the showy and entomophilous goldenrod (Solidago) is frequently blamed for respiratory allergies, of which it is innocent, since its pollen cannot be airborne. Instead the allergen is usually the pollen of the contemporary bloom of anemophilous ragweed (Ambrosia), which can drift for many kilometers.

Scientists have collected samples of ragweed pollen 400 miles out at sea and 2 miles high in the air.[18] A single ragweed plant can generate a million grains of pollen per day.[20]

It is common to hear people say they are allergic to colorful or scented flowers like roses. In fact, only florists, gardeners, and others who have prolonged, close contact with flowers are likely to be sensitive to pollen from these plants. Most people have little contact with the large, heavy, waxy pollen grains of such flowering plants because this type of pollen is not carried by wind but by insects such as butterflies and bees.

Attraction methods


A Bee orchid has evolved over many generations to better mimic a female bee to attract male bees as pollinators.

Plants cannot move from one location to another, thus many flowers have evolved to attract animals to transfer pollen between individuals in dispersed populations. Flowers that are insect-pollinated are called entomophilous; literally "insect-loving" in Greek. They can be highly modified along with the pollinating insects by co-evolution. Flowers commonly have glands called nectaries on various parts that attract animals looking for nutritious nectar. Birds and bees have color vision, enabling them to seek out "colorful" flowers.

Some flowers have patterns, called nectar guides, that show pollinators where to look for nectar; they may be visible only under ultraviolet light, which is visible to bees and some other insects. Flowers also attract pollinators by scent and some of those scents are pleasant to our sense of smell. Not all flower scents are appealing to humans; a number of flowers are pollinated by insects that are attracted to rotten flesh and have flowers that smell like dead animals, often called Carrion flowers, including Rafflesia, the titan arum, and the North American pawpaw (Asimina triloba). Flowers pollinated by night visitors, including bats and moths, are likely to concentrate on scent to attract pollinators and most such flowers are white.

Other flowers use mimicry to attract pollinators. Some species of orchids, for example, produce flowers resembling female bees in color, shape, and scent. Male bees move from one such flower to another in search of a mate.

Pollination mechanism

The pollination mechanism employed by a plant depends on what method of pollination is utilized.
Most flowers can be divided between two broad groups of pollination methods:

Entomophilous: flowers attract and use insects, bats, birds or other animals to transfer pollen from one flower to the next. Often they are specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to the bodies of the pollinator when it lands in search of its attractant (such as nectar, pollen, or a mate). In pursuing this attractant from many flowers of the same species, the pollinator transfers pollen to the stigmas—arranged with equally pointed precision—of all of the flowers it visits. Many flowers rely on simple proximity between flower parts to ensure pollination. Others, such as the Sarracenia or lady-slipper orchids, have elaborate designs to ensure pollination while preventing self-pollination.

Grass flower with vestigial perianth or lodicules

Anemophilous: flowers use the wind to move pollen from one flower to the next, examples include the grasses, Birch trees, Ragweed and Maples. They have no need to attract pollinators and therefore tend not to grow large blossoms. Whereas the pollen of entomophilous flowers tends to be large-grained, sticky, and rich in protein (another "reward" for pollinators), anemophilous flower pollen is usually small-grained, very light, and of little nutritional value to insects, though it may still be gathered in times of dearth. Honeybees and bumblebees actively gather anemophilous corn (maize) pollen, though it is of little value to them.

Some flowers are self-pollinated and use flowers that never open or are self-pollinated before the flowers open, these flowers are called cleistogamous. Many Viola species and some Salvia have these types of flowers.

Flower-pollinator relationships

Many flowers have close relationships with one or a few specific pollinating organisms. Many flowers, for example, attract only one specific species of insect, and therefore rely on that insect for successful reproduction. This close relationship is often given as an example of coevolution, as the flower and pollinator are thought to have developed together over a long period of time to match each other's needs.

This close relationship compounds the negative effects of extinction. The extinction of either member in such a relationship would mean almost certain extinction of the other member as well. Some endangered plant species are so because of shrinking pollinator populations.

Fertilization and dispersal

Some flowers with both stamens and a pistil are capable of self-fertilization, which does increase the chance of producing seeds but limits genetic variation. The extreme case of self-fertilization occurs in flowers that always self-fertilize, such as many dandelions. Conversely, many species of plants have ways of preventing self-fertilization. Unisexual male and female flowers on the same plant may not appear or mature at the same time, or pollen from the same plant may be incapable of fertilizing its ovules. The latter flower types, which have chemical barriers to their own pollen, are referred to as self-sterile or self-incompatible.

Evolution

Archaefructus liaoningensis, one of the earliest known flowering plants

Amborella trichopoda, the sister group to the rest of the flowering plants

While land plants have existed for about 425 million years, the first ones reproduced by a simple adaptation of their aquatic counterparts: spores. In the sea, plants—and some animals—can simply scatter out genetic clones of themselves to float away and grow elsewhere. This is how early plants reproduced. But plants soon evolved methods of protecting these copies to deal with drying out and other abuse which is even more likely on land than in the sea. The protection became the seed, though it had not yet evolved the flower. Early seed-bearing plants include the ginkgo and conifers. The earliest fossil of a flowering plant, Archaefructus liaoningensis, is dated about 125 million years old.[21]

Several groups of extinct gymnosperms, particularly seed ferns, have been proposed as the ancestors of flowering plants but there is no continuous fossil evidence showing exactly how flowers evolved. The apparently sudden appearance of relatively modern flowers in the fossil record posed such a problem for the theory of evolution that it was called an "abominable mystery" by Charles Darwin. Recently discovered angiosperm fossils such as Archaefructus, along with further discoveries of fossil gymnosperms, suggest how angiosperm characteristics may have been acquired in a series of steps.

Recent DNA analysis (molecular systematics)[22][23] shows that Amborella trichopoda, found on the Pacific island of New Caledonia, is the sister group to the rest of the flowering plants, and morphological studies[24] suggest that it has features which may have been characteristic of the earliest flowering plants.

The general assumption is that the function of flowers, from the start, was to involve animals in the reproduction process. Pollen can be scattered without bright colors and obvious shapes, which would therefore be a liability, using the plant's resources, unless they provide some other benefit. One proposed reason for the sudden, fully developed appearance of flowers is that they evolved in an isolated setting like an island, or chain of islands, where the plants bearing them were able to develop a highly specialized relationship with some specific animal (a wasp, for example), the way many island species develop today. This symbiotic relationship, with a hypothetical wasp bearing pollen from one plant to another much the way fig wasps do today, could have eventually resulted in both the plant(s) and their partners developing a high degree of specialization. Island genetics is believed to be a common source of speciation, especially when it comes to radical adaptations which seem to have required inferior transitional forms. Note that the wasp example is not incidental; bees, apparently evolved specifically for symbiotic plant relationships, are descended from wasps.

Likewise, most fruit used in plant reproduction comes from the enlargement of parts of the flower. This fruit is frequently a tool which depends upon animals wishing to eat it, and thus scattering the seeds it contains.

While many such symbiotic relationships remain too fragile to survive competition with mainland organisms, flowers proved to be an unusually effective means of production, spreading (whatever their actual origin) to become the dominant form of land plant life.

While there is only hard proof of such flowers existing about 130 million years ago, there is some circumstantial evidence that they did exist up to 250 million years ago. A chemical used by plants to defend their flowers, oleanane, has been detected in fossil plants that old, including gigantopterids,[25] which evolved at that time and bear many of the traits of modern, flowering plants, though they are not known to be flowering plants themselves, because only their stems and prickles have been found preserved in detail; one of the earliest examples of petrification.

The similarity in leaf and stem structure can be very important, because flowers are genetically just an adaptation of normal leaf and stem components on plants, a combination of genes normally responsible for forming new shoots.[26] The most primitive flowers are thought to have had a variable number of flower parts, often separate from (but in contact with) each other. The flowers would have tended to grow in a spiral pattern, to be bisexual (in plants, this means both male and female parts on the same flower), and to be dominated by the ovary (female part). As flowers grew more advanced, some variations developed parts fused together, with a much more specific number and design, and with either specific sexes per flower or plant, or at least "ovary inferior".

Flower evolution continues to the present day; modern flowers have been so profoundly influenced by humans that many of them cannot be pollinated in nature. Many modern, domesticated flowers used to be simple weeds, which only sprouted when the ground was disturbed. Some of them tended to grow with human crops, and the prettiest did not get plucked because of their beauty, developing a dependence upon and special adaptation to human affection.[27]

Symbolism


Red Rose

Lilies are often used to denote life or resurrection

Flowers are common subjects of still life paintings, such as this one by Ambrosius Bosschaert the Elder

Many flowers have important symbolic meanings in Western culture.[28] The practice of assigning meanings to flowers is known as floriography. Some of the more common examples include:
  • Red roses are given as a symbol of love, beauty, and passion.[29]
  • Poppies are a symbol of consolation in time of death. In the United Kingdom, New Zealand, Australia and Canada, red poppies are worn to commemorate soldiers who have died in times of war.
  • Irises/Lily are used in burials as a symbol referring to "resurrection/life". It is also associated with stars (sun) and its petals blooming/shining.
  • Daisies are a symbol of innocence.
Flowers within art are also representative of the female genitalia,[30] as seen in the works of artists such as Georgia O'Keeffe, Imogen Cunningham, Veronica Ruiz de Velasco, and Judy Chicago, and in fact in Asian and western classical art. Many cultures around the world have a marked tendency to associate flowers with femininity.

The great variety of delicate and beautiful flowers has inspired the works of numerous poets, especially from the 18th-19th century Romantic era. Famous examples include William Wordsworth's I Wandered Lonely as a Cloud and William Blake's Ah! Sun-Flower.

Because of their varied and colorful appearance, flowers have long been a favorite subject of visual artists as well. Some of the most celebrated paintings from well-known painters are of flowers, such as Van Gogh's sunflowers series or Monet's water lilies. Flowers are also dried, freeze dried and pressed in order to create permanent, three-dimensional pieces of flower art.

Their symbolism in dreams has also been discussed, with possible interpretations including "blossoming potential".[31]

The Roman goddess of flowers, gardens, and the season of Spring is Flora. The Greek goddess of spring, flowers and nature is Chloris.

In Hindu mythology, flowers have a significant status. Vishnu, one of the three major gods in the Hindu system, is often depicted standing straight on a lotus flower.[32] Apart from the association with Vishnu, the Hindu tradition also considers the lotus to have spiritual significance.[33] For example, it figures in the Hindu stories of creation.[34]

Usage


Flower market - Detroit's Eastern Market

A woman spreading flowers over a lingam in a temple in Varanasi

In modern times people have sought ways to cultivate, buy, wear, or otherwise be around flowers and blooming plants, partly because of their agreeable appearance and smell.[citation needed] Around the world, people use flowers for a wide range of events and functions that, cumulatively, encompass one's lifetime:
  • For new births or christenings
  • As a corsage or boutonniere worn at social functions or for holidays
  • As tokens of love or esteem
  • For wedding flowers for the bridal party, and for decorations for the hall
  • As brightening decorations within the home
  • As a gift of remembrance for bon voyage parties, welcome-home parties, and "thinking of you" gifts
  • For funeral flowers and expressions of sympathy for the grieving
  • For worshiping goddesses. In Hindu culture adherents commonly bring flowers as a gift to temples
People therefore grow flowers around their homes, dedicate entire parts of their living space to flower gardens, pick wildflowers, or buy flowers from florists who depend on an entire network of commercial growers and shippers to support their trade.

Flowers provide less food than other major plants parts (seeds, fruits, roots, stems and leaves) but they provide several important foods and spices. Flower vegetables include broccoli, cauliflower and artichoke. The most expensive spice, saffron, consists of dried stigmas of a crocus. Other flower spices are cloves and capers. Hops flowers are used to flavor beer. Marigold flowers are fed to chickens to give their egg yolks a golden yellow color, which consumers find more desirable; dried and ground marigold flowers are also used as a spice and colouring agent in Georgian cuisine. Flowers of the dandelion and elder are often made into wine. Bee pollen, pollen collected from bees, is considered a health food by some people. Honey consists of bee-processed flower nectar and is often named for the type of flower, e.g. orange blossom honey, clover honey and tupelo honey.

Hundreds of fresh flowers are edible but few are widely marketed as food. They are often used to add color and flavor to salads. Squash flowers are dipped in breadcrumbs and fried. Edible flowers include nasturtium, chrysanthemum, carnation, cattail, honeysuckle, chicory, cornflower, canna, and sunflower. Some edible flowers are sometimes candied such as daisy, rose, and violet (one may also come across a candied pansy).

Flowers can also be made into herbal teas. Dried flowers such as chrysanthemum, rose, jasmine, camomile are infused into tea both for their fragrance and medical properties. Sometimes, they are also mixed with tea leaves for the added fragrance.

Flowers have been used since as far back as 50,000 years in funeral rituals. Many cultures do draw a connection between flowers and life and death, and because of their seasonal return flowers also suggest rebirth, which may explain why many people place flowers upon graves. In ancient times the Greeks would place a crown of flowers on the head of the deceased as well as cover tombs with wreaths and flower petals. Rich and powerful women in ancient Egypt would wear floral headdresses and necklaces upon their death as representations of renewal and a joyful afterlife, and the Mexicans to this day use flowers prominently in their Day of the Dead celebrations in the same way that their Aztec ancestors did.
Eight Flowers, a painting by artist Qian Xuan, 13th century, Palace Museum, Beijing.

Evolutionary algorithm

From Wikipedia, the free encyclopedia https://en.wikipedia.org/wiki/Evolutionary_algorithm In computational int...