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Friday, April 25, 2025

Natural environment

From Wikipedia, the free encyclopedia
Land management has preserved the natural characteristics of Hopetoun Falls, Australia while allowing ample access for visitors.
An image of the Sahara Desert from satellite. It is the world's largest hot desert and third-largest desert after the polar deserts.

The natural environment or natural world encompasses all biotic and abiotic things occurring naturally, meaning in this case not artificial. The term is most often applied to Earth or some parts of Earth. This environment encompasses the interaction of all living species, climate, weather and natural resources that affect human survival and economic activity. The concept of the natural environment can be distinguished as components:

In contrast to the natural environment is the built environment. Built environments are where humans have fundamentally transformed landscapes such as urban settings and agricultural land conversion, the natural environment is greatly changed into a simplified human environment. Even acts which seem less extreme, such as building a mud hut or a photovoltaic system in the desert, the modified environment becomes an artificial one. Though many animals build things to provide a better environment for themselves, they are not human, hence beaver dams and the works of mound-building termites are thought of as natural.

People cannot find absolutely natural environments on Earth, naturalness usually varies in a continuum, from 100% natural in one extreme to 0% natural in the other. The massive environmental changes of humanity in the Anthropocene have fundamentally effected all natural environments including: climate change, biodiversity loss and pollution from plastic and other chemicals in the air and water. More precisely, we can consider the different aspects or components of an environment, and see that their degree of naturalness is not uniform. If, for instance, in an agricultural field, the mineralogic composition and the structure of its soil are similar to those of an undisturbed forest soil, but the structure is quite different.

Composition

Earth's layered structure: (1) inner core; (2) outer core; (3) lower mantle; (4) upper mantle; (5) lithosphere; (6) crust

Earth science generally recognizes four spheres, the lithosphere, the hydrosphere, the atmosphere and the biosphere as correspondent to rocks, water, air and life respectively. Some scientists include as part of the spheres of the Earth, the cryosphere (corresponding to ice) as a distinct portion of the hydrosphere, as well as the pedosphere (to soil) as an active and intermixed sphere. Earth science (also known as geoscience, the geographical sciences or the Earth Sciences), is an all-embracing term for the sciences related to the planet Earth. There are four major disciplines in earth sciences, namely geography, geology, geophysics and geodesy. These major disciplines use physics, chemistry, biology, chronology and mathematics to build a qualitative and quantitative understanding of the principal areas or spheres of Earth.

Geological activity

The Earth's crust or lithosphere, is the outermost solid surface of the planet and is chemically, physically and mechanically different from underlying mantle. It has been generated greatly by igneous processes in which magma cools and solidifies to form solid rock. Beneath the lithosphere lies the mantle which is heated by the decay of radioactive elements. The mantle though solid is in a state of rheic convection. This convection process causes the lithospheric plates to move, albeit slowly. The resulting process is known as plate tectonics. Volcanoes result primarily from the melting of subducted crust material or of rising mantle at mid-ocean ridges and mantle plumes.

Water on Earth

Coral reefs have significant marine biodiversity.

Most water is found in various kinds of natural body of water.

Oceans

An ocean is a major body of saline water and a component of the hydrosphere. Approximately 71% of the surface of the Earth (an area of some 362 million square kilometers) is covered by ocean, a continuous body of water that is customarily divided into several principal oceans and smaller seas. More than half of this area is over 3,000 meters (9,800 ft) deep. Average oceanic salinity is around 35 parts per thousand (ppt) (3.5%), and nearly all seawater has a salinity in the range of 30 to 38 ppt. Though generally recognized as several separate oceans, these waters comprise one global, interconnected body of salt water often referred to as the World Ocean or global ocean. The deep seabeds are more than half the Earth's surface, and are among the least-modified natural environments. The major oceanic divisions are defined in part by the continents, various archipelagos and other criteria, these divisions are : (in descending order of size) the Pacific Ocean, the Atlantic Ocean, the Indian Ocean, the Southern Ocean and the Arctic Ocean.

Rivers

A river is a natural watercourse, usually freshwater, flowing toward an ocean, a lake, a sea or another river. A few rivers simply flow into the ground and dry up completely without reaching another body of water.

Rocky stream in the U.S. state of Hawaii

The water in a river is usually in a channel, made up of a stream bed between banks. In larger rivers there is often also a wider floodplain shaped by waters over-topping the channel. Flood plains may be very wide in relation to the size of the river channel. Rivers are a part of the hydrological cycle. Water within a river is generally collected from precipitation through surface runoff, groundwater recharge, springs and the release of water stored in glaciers and snowpacks.

Small rivers may also be called by several other names, including stream, creek and brook. Their current is confined within a bed and stream banks. Streams play an important corridor role in connecting fragmented habitats and thus in conserving biodiversity. The study of streams and waterways in general is known as surface hydrology.

Lakes

Lácar Lake, of glacial origin, in the province of Neuquén, Argentina

A lake (from Latin lacus) is a terrain feature, a body of water that is localized to the bottom of basin. A body of water is considered a lake when it is inland, is not part of an ocean and is larger and deeper than a pond.

A swamp area in Everglades National Park, Florida, US

Natural lakes on Earth are generally found in mountainous areas, rift zones and areas with ongoing or recent glaciation. Other lakes are found in endorheic basins or along the courses of mature rivers. In some parts of the world, there are many lakes because of chaotic drainage patterns left over from the last ice age. All lakes are temporary over geologic time scales, as they will slowly fill in with sediments or spill out of the basin containing them.

Ponds

A pond is a body of standing water, either natural or human-made, that is usually smaller than a lake. A wide variety of human-made bodies of water are classified as ponds, including water gardens designed for aesthetic ornamentation, fish ponds designed for commercial fish breeding and solar ponds designed to store thermal energy. Ponds and lakes are distinguished from streams by their current speed. While currents in streams are easily observed, ponds and lakes possess thermally driven micro-currents and moderate wind-driven currents. These features distinguish a pond from many other aquatic terrain features, such as stream pools and tide pools.

Human impact on water

Humans impact the water in different ways such as modifying rivers (through dams and stream channelization), urbanization and deforestation. These impact lake levels, groundwater conditions, water pollution, thermal pollution, and marine pollution. Humans modify rivers by using direct channel manipulation. We build dams and reservoirs and manipulate the direction of the rivers and water path. Dams can usefully create reservoirs and hydroelectric power. However, reservoirs and dams may negatively impact the environment and wildlife. Dams stop fish migration and the movement of organisms downstream. Urbanization affects the environment because of deforestation and changing lake levels, groundwater conditions, etc. Deforestation and urbanization go hand in hand. Deforestation may cause flooding, declining stream flow and changes in riverside vegetation. The changing vegetation occurs because when trees cannot get adequate water they start to deteriorate, leading to a decreased food supply for the wildlife in an area.

Atmosphere, climate and weather

Atmospheric gases scatter blue light more than other wavelengths, creating a blue halo when seen from space.
A view of Earth's troposphere from an airplane
Lightning is an atmospheric discharge of electricity accompanied by thunder, which occurs during thunderstorms and certain other natural conditions.

The atmosphere of the Earth serves as a key factor in sustaining the planetary ecosystem. The thin layer of gases that envelops the Earth is held in place by the planet's gravity. Dry air consists of 78% nitrogen, 21% oxygen, 1% argon, inert gases and carbon dioxide. The remaining gases are often referred to as trace gases. The atmosphere includes greenhouse gases such as carbon dioxide, methane, nitrous oxide and ozone. Filtered air includes trace amounts of many other chemical compounds. Air also contains a variable amount of water vapor and suspensions of water droplets and ice crystals seen as clouds. Many natural substances may be present in tiny amounts in an unfiltered air sample, including dust, pollen and spores, sea spray, volcanic ash and meteoroids. Various industrial pollutants also may be present, such as chlorine (elementary or in compounds), fluorine compounds, elemental mercury, and sulphur compounds such as sulphur dioxide (SO2).

The ozone layer of the Earth's atmosphere plays an important role in reducing the amount of ultraviolet (UV) radiation that reaches the surface. As DNA is readily damaged by UV light, this serves to protect life at the surface. The atmosphere also retains heat during the night, thereby reducing the daily temperature extremes.

Layers of the atmosphere

Principal layers

Earth's atmosphere can be divided into five main layers. These layers are mainly determined by whether temperature increases or decreases with altitude. From highest to lowest, these layers are:

  • Exosphere: The outermost layer of Earth's atmosphere extends from the exobase upward, mainly composed of hydrogen and helium.
  • Thermosphere: The top of the thermosphere is the bottom of the exosphere, called the exobase. Its height varies with solar activity and ranges from about 350–800 km (220–500 mi; 1,150,000–2,620,000 ft). The International Space Station orbits in this layer, between 320 and 380 km (200 and 240 mi). In another way, the thermosphere is Earth's second highest atmospheric layer, extending from approximately 260,000 feet at the mesopause to the thermopause at altitudes ranging from 1,600,000 to 3,300,000 feet.
  • Mesosphere: The mesosphere extends from the stratopause to 80–85 km (50–53 mi; 262,000–279,000 ft). It is the layer where most meteors burn up upon entering the atmosphere.
  • Stratosphere: The stratosphere extends from the tropopause to about 51 km (32 mi; 167,000 ft). The stratopause, which is the boundary between the stratosphere and mesosphere, typically is at 50 to 55 km (31 to 34 mi; 164,000 to 180,000 ft).
  • Troposphere: The troposphere begins at the surface and extends to between 7 km (23,000 ft) at the poles and 17 km (56,000 ft) at the equator, with some variation due to weather. The troposphere is mostly heated by transfer of energy from the surface, so on average the lowest part of the troposphere is warmest and temperature decreases with altitude. The tropopause is the boundary between the troposphere and stratosphere.
Other layers

Within the five principal layers determined by temperature there are several layers determined by other properties.

  • The ozone layer is contained within the stratosphere. It is mainly located in the lower portion of the stratosphere from about 15–35 km (9.3–21.7 mi; 49,000–115,000 ft), though the thickness varies seasonally and geographically. About 90% of the ozone in our atmosphere is contained in the stratosphere.
  • The ionosphere: The part of the atmosphere that is ionized by solar radiation, stretches from 50 to 1,000 km (31 to 621 mi; 160,000 to 3,280,000 ft) and typically overlaps both the exosphere and the thermosphere. It forms the inner edge of the magnetosphere.
  • The homosphere and heterosphere: The homosphere includes the troposphere, stratosphere and mesosphere. The upper part of the heterosphere is composed almost completely of hydrogen, the lightest element.
  • The planetary boundary layer is the part of the troposphere that is nearest the Earth's surface and is directly affected by it, mainly through turbulent diffusion.

Effects of global warming

The retreat of glaciers since 1850 of Aletsch Glacier in the Swiss Alps (situation in 1979, 1991 and 2002), due to global warming

The dangers of global warming are being increasingly studied by a wide global consortium of scientists. These scientists are increasingly concerned about the potential long-term effects of global warming on our natural environment and on the planet. Of particular concern is how climate change and global warming caused by anthropogenic, or human-made releases of greenhouse gases, most notably carbon dioxide, can act interactively and have adverse effects upon the planet, its natural environment and humans' existence. It is clear the planet is warming, and warming rapidly. This is due to the greenhouse effect, which is caused by greenhouse gases, which trap heat inside the Earth's atmosphere because of their more complex molecular structure which allows them to vibrate and in turn trap heat and release it back towards the Earth. This warming is also responsible for the extinction of natural habitats, which in turn leads to a reduction in wildlife population. The most recent report from the Intergovernmental Panel on Climate Change (the group of the leading climate scientists in the world) concluded that the earth will warm anywhere from 2.7 to almost 11 degrees Fahrenheit (1.5 to 6 degrees Celsius) between 1990 and 2100. Efforts have been increasingly focused on the mitigation of greenhouse gases that are causing climatic changes, on developing adaptative strategies to global warming, to assist humans, other animal, and plant species, ecosystems, regions and nations in adjusting to the effects of global warming. Some examples of recent collaboration to address climate change and global warming include:

Another view of the Aletsch Glacier in the Swiss Alps, which because of global warming has been decreasing

A significantly profound challenge is to identify the natural environmental dynamics in contrast to environmental changes not within natural variances. A common solution is to adapt a static view neglecting natural variances to exist. Methodologically, this view could be defended when looking at processes which change slowly and short time series, while the problem arrives when fast processes turns essential in the object of the study.

Climate

Map of world dividing climate zones, largely influenced by latitude. The zones, going from the equator upward (and downward) are Tropical, Dry, Moderate, Continental and Polar. There are subzones within these zones.
Worldwide climate classifications map

Climate looks at the statistics of temperature, humidity, atmospheric pressure, wind, rainfall, atmospheric particle count and other meteorological elements in a given region over long periods of time. Weather, on the other hand, is the present condition of these same elements over periods up to two weeks.

Climates can be classified according to the average and typical ranges of different variables, most commonly temperature and precipitation. The most commonly used classification scheme is the one originally developed by Wladimir Köppen. The Thornthwaite system, in use since 1948, uses evapotranspiration as well as temperature and precipitation information to study animal species diversity and the potential impacts of climate changes.

Weather

A rainbow is an optical and meteorological phenomenon that causes a spectrum of light to appear in the sky when the Sun shines onto droplets of moisture in the Earth's atmosphere.

Weather is a set of all the phenomena occurring in a given atmospheric area at a given time. Most weather phenomena occur in the troposphere, just below the stratosphere. Weather refers, generally, to day-to-day temperature and precipitation activity, whereas climate is the term for the average atmospheric conditions over longer periods of time. When used without qualification, "weather" is understood to be the weather of Earth.

Weather occurs due to density (temperature and moisture) differences between one place and another. These differences can occur due to the sun angle at any particular spot, which varies by latitude from the tropics. The strong temperature contrast between polar and tropical air gives rise to the jet stream. Weather systems in the mid-latitudes, such as extratropical cyclones, are caused by instabilities of the jet stream flow. Because the Earth's axis is tilted relative to its orbital plane, sunlight is incident at different angles at different times of the year. On the Earth's surface, temperatures usually range ±40 °C (100 °F to −40 °F) annually. Over thousands of years, changes in the Earth's orbit have affected the amount and distribution of solar energy received by the Earth and influenced long-term climate.

Surface temperature differences in turn cause pressure differences. Higher altitudes are cooler than lower altitudes due to differences in compressional heating. Weather forecasting is the application of science and technology to predict the state of the atmosphere for a future time and a given location. The atmosphere is a chaotic system, and small changes to one part of the system can grow to have large effects on the system as a whole. Human attempts to control the weather have occurred throughout human history, and there is evidence that civilized human activity such as agriculture and industry has inadvertently modified weather patterns.

Life

There are many plant species on the planet.
An example of the many animal species on the Earth

Evidence suggests that life on Earth has existed for about 3.7 billion years. All known life forms share fundamental molecular mechanisms, and based on these observations, theories on the origin of life attempt to find a mechanism explaining the formation of a primordial single cell organism from which all life originates. There are many different hypotheses regarding the path that might have been taken from simple organic molecules via pre-cellular life to protocells and metabolism.

Although there is no universal agreement on the definition of life, scientists generally accept that the biological manifestation of life is characterized by organization, metabolism, growth, adaptation, response to stimuli and reproduction. Life may also be said to be simply the characteristic state of organisms. In biology, the science of living organisms, "life" is the condition which distinguishes active organisms from inorganic matter, including the capacity for growth, functional activity and the continual change preceding death.

A diverse variety of living organisms (life forms) can be found in the biosphere on Earth, and properties common to these organisms—plants, animals, fungi, protists, archaea, and bacteria—are a carbon- and water-based cellular form with complex organization and heritable genetic information. Living organisms undergo metabolism, maintain homeostasis, possess a capacity to grow, respond to stimuli, reproduce and, through natural selection, adapt to their environment in successive generations. More complex living organisms can communicate through various means.

Ecosystems

Rainforests often have a great deal of biodiversity with many plant and animal species. This is the Gambia River in Senegal's Niokolo-Koba National Park.

An ecosystem (also called an environment) is a natural unit consisting of all plants, animals, and micro-organisms (biotic factors) in an area functioning together with all of the non-living physical (abiotic) factors of the environment.

Central to the ecosystem concept is the idea that living organisms are continually engaged in a highly interrelated set of relationships with every other element constituting the environment in which they exist. Eugene Odum, one of the founders of the science of ecology, stated: "Any unit that includes all of the organisms (i.e.: the "community") in a given area interacting with the physical environment so that a flow of energy leads to clearly defined trophic structure, biotic diversity, and material cycles (i.e.: exchange of materials between living and nonliving parts) within the system is an ecosystem."

Old-growth forest and a creek on Larch Mountain, in the U.S. state of Oregon

The human ecosystem concept is then grounded in the deconstruction of the human/nature dichotomy, and the emergent premise that all species are ecologically integrated with each other, as well as with the abiotic constituents of their biotope.

A more significant number or variety of species or biological diversity of an ecosystem may contribute to greater resilience of an ecosystem because there are more species present at a location to respond to change and thus "absorb" or reduce its effects. This reduces the effect before the ecosystem's structure changes to a different state. This is not universally the case and there is no proven relationship between the species diversity of an ecosystem and its ability to provide goods and services on a sustainable level.

The term ecosystem can also pertain to human-made environments, such as human ecosystems and human-influenced ecosystems. It can describe any situation where there is relationship between living organisms and their environment. Fewer areas on the surface of the earth today exist free from human contact, although some genuine wilderness areas continue to exist without any forms of human intervention.

Biogeochemical cycles

Chloroplasts conduct photosynthesis and are found in plant cells and other eukaryotic organisms. These are chloroplasts visible in the cells of Plagiomnium affine — many-fruited thyme-moss.

Global biogeochemical cycles are critical to life, most notably those of water, oxygen, carbon, nitrogen and phosphorus.

  • The nitrogen cycle is the transformation of nitrogen and nitrogen-containing compounds in nature. It is a cycle which includes gaseous components.
  • The water cycle, is the continuous movement of water on, above, and below the surface of the Earth. Water can change states among liquid, vapour, and ice at various places in the water cycle. Although the balance of water on Earth remains fairly constant over time, individual water molecules can come and go.
  • The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of the Earth.
  • The oxygen cycle is the movement of oxygen within and between its three main reservoirs: the atmosphere, the biosphere, and the lithosphere. The main driving factor of the oxygen cycle is photosynthesis, which is responsible for the modern Earth's atmospheric composition and life.
  • The phosphorus cycle is the movement of phosphorus through the lithosphere, hydrosphere, and biosphere. The atmosphere does not play a significant role in the movements of phosphorus, because phosphorus and phosphorus compounds are usually solids at the typical ranges of temperature and pressure found on Earth.

Wilderness

A conifer forest in the Swiss Alps (National Park)
The Ahklun Mountains and the Togiak Wilderness within the Togiak National Wildlife Refuge in the U.S. state of Alaska

Wilderness is generally defined as a natural environment on Earth that has not been significantly modified by human activity. The WILD Foundation goes into more detail, defining wilderness as: "The most intact, undisturbed wild natural areas left on our planet – those last truly wild places that humans do not control and have not developed with roads, pipelines or other industrial infrastructure." Wilderness areas and protected parks are considered important for the survival of certain species, ecological studies, conservation, solitude, and recreation. Wilderness is deeply valued for cultural, spiritual, moral, and aesthetic reasons. Some nature writers believe wilderness areas are vital for the human spirit and creativity.

The word, "wilderness", derives from the notion of wildness; in other words that which is not controllable by humans. The word etymology is from the Old English wildeornes, which in turn derives from wildeor meaning wild beast (wild + deor = beast, deer). From this point of view, it is the wildness of a place that makes it a wilderness. The mere presence or activity of people does not disqualify an area from being "wilderness". Many ecosystems that are, or have been, inhabited or influenced by activities of people may still be considered "wild". This way of looking at wilderness includes areas within which natural processes operate without very noticeable human interference.

Wildlife includes all non-domesticated plants, animals and other organisms. Domesticating wild plant and animal species for human benefit has occurred many times all over the planet, and has a major impact on the environment, both positive and negative. Wildlife can be found in all ecosystems. Deserts, rain forests, plains, and other areas—including the most developed urban sites—all have distinct forms of wildlife. While the term in popular culture usually refers to animals that are untouched by civilized human factors, most scientists agree that wildlife around the world is (now) impacted by human activities.

A view of wilderness in Estonia

Challenges

Before flue-gas desulfurization was installed, the air-polluting emissions from this power plant in New Mexico contained excessive amounts of sulfur dioxide.
Amazon rainforest in Brazil. The tropical rainforests of South America contain the largest diversity of species on Earth, including some that have evolved within the past few hundred thousand years.

It is the common understanding of natural environment that underlies environmentalism — a broad political, social and philosophical movement that advocates various actions and policies in the interest of protecting what nature remains in the natural environment, or restoring or expanding the role of nature in this environment. While true wilderness is increasingly rare, wild nature (e.g., unmanaged forests, uncultivated grasslands, wildlife, wildflowers) can be found in many locations previously inhabited by humans.

Goals for the benefit of people and natural systems, commonly expressed by environmental scientists and environmentalists include:

Criticism

In some cultures the term environment is meaningless because there is no separation between people and what they view as the natural world, or their surroundings. Specifically in the United States and Arabian countries many native cultures do not recognize the "environment", or see themselves as environmentalists.

Androgenesis

From Wikipedia, the free encyclopedia
https://en.wikipedia.org/wiki/Androgenesis

Androgenesis is a system of asexual reproduction that requires the presence of eggs and occurs when a zygote is produced with only paternal nuclear genes. During standard sexual reproduction, one female parent and one male parent each produce haploid gametes (such as a sperm or egg cell, each containing only a single set of chromosomes), which recombine to create offspring with genetic material from both parents. However, in androgenesis, there is no recombination of maternal and paternal chromosomes, and only the paternal chromosomes are passed down to the offspring. (The inverse of this is gynogenesis, where only the maternal chromosomes are inherited, which is more common than androgenesis). The offspring produced in androgenesis will still have maternally inherited mitochondria, as is the case with most sexually reproducing species.

One of two things can occur to produce offspring with exclusively paternal genetic material: the maternal nuclear genome can be eliminated from the zygote, or the female can produce an egg with no nucleus, resulting in an embryo developing with only the genome of the male gamete. Although androgenesis requires both male and females gametes, it is not strictly considered a form of sexual reproduction because the offspring have uniparental nuclear DNA that has not undergone recombination, and the proliferation of androgenesis can lead to exclusively asexually reproducing species.

Androgenesis occurs in nature in many organisms such as plants (including trees, flowers, barley, algae or corn), invertebrates (including clams, stick insects, some ants, bees, flies and parasitic wasps) and vertebrates (mainly amphibians and fish). Androgenesis has also been observed in roosters and genetically modified laboratory mice.

Elimination of the maternal nuclear genome

When androgenesis occurs via elimination of the maternal nuclear genome, the elimination takes place after fertilization. The nuclei of the two gametes fuse as normal, but immediately afterwards the male nuclear genome then eliminates the female nuclear genome, leaving a fertilized ovum with only the nuclear genome of the male parent. If viable, the resulting offspring is a clone or sub-clone of the sperm- or pollen-producing parent.

Elimination of the maternal nuclear genome is evolutionarily advantageous for the male parent, because all offspring produced have entirely paternally-inherited alleles: in contrast, a male parent that reproduces sexually without androgenesis only passes down half of its genetic material to each of its offspring. A male allele promoting the elimination of the female gametic nucleus therefore has a high fitness advantage and can spread through a population and even reach fixation. However, this may be part of the reason androgenesis is very rarely observed in nature: despite being advantageous to the individual producing offspring, it is deleterious to the population as a whole: if an androgenesis-inducing allele reaches high frequencies, egg-producing individuals become rare. Because both egg- and sperm-producers are necessary for androgenesis, if the sex ratio becomes highly unbalanced and there are too few egg-producers, the population is driven to extinction. However, in hermaphrodites (species where a single individual produces both male and female gametes), this is less of a problem since there is no sex ratio.

Female production of a non-nuclear egg

Androgenesis can also occur through female production of an egg without a nucleus. Upon fertilization with pollen or sperm, there is no maternal nucleus to expel, and a zygote is produced that derives its nuclear genome entirely from its paternal parent. It is unclear why production of non-nucleate eggs would have evolved, because there is no fitness advantage to the egg parent: none of its nuclear genes are being passed onto its offspring. Therefore, any female allele causing non-nucleate egg production would be highly disadvantageous. This form of androgenesis could spread through genetic drift, or there may be some unknown benefit to the egg-producing parent. Species in which non-nucleate egg production occurs are less likely to go extinct than species where the maternal nuclear genome is eliminated. This is because females producing non-nucleate eggs are disfavored by natural selection, so their proportion in a population will remain low.

Male apomixis

Another type of androgenesis is male apomixis, which is a reproductive process in which a plant develops from a sperm cell (male gamete) without the participation of a female cell (ovum). In this process, the zygote is formed solely with genetic material from the father, resulting in offspring genetically identical to the male organism. This has been noted in many plants such as Nicotiana, Capsicum frutescens, Cicer arietinum, Poa arachnifera, Cupressus sempervirens, Solanum verrucosum, Phaeophyceae,Elodea canadensis, Barleys, Tripsacum dactyloides, and Zea mays, and occurs as the regular reproductive method in Cupressus dupreziana. This contrasts with the more common apomixis, where development occurs without fertilization, but with genetic material only from the mother.

There are also clonal species that reproduce through vegetative reproduction such as Lomatia tasmanica, Lagarostrobos franklinii, and Pando, where the genetic material is exclusively male.

Obligate androgenesis

Obligate androgenesis is the process by which males are only able to produce offspring exclusively through male genetic material, where mating with females of related species is not necessary to produce offspring. This leads to these species being able to survive in the absence of females. They are also capable of interbreeding with sexual and other androgenetic lineages in a phenomenon sometimes referred to as egg parasitism or androgenetic parasitism. This method of reproduction is relatively rare and has been found in several species of clams of the genus Corbicula, some plants like Elodea canadensis, Cupressus dupreziana, Lomatia tasmanica, Lagarostrobos franklinii, and Pando, algae of the genus Phaeophyceae, and recently in the all male fish species Squalius alburnoides. Although the most common term to refer to completely asexual reproduction in males is paternal apomixis, the term obligate androgenesis is more commonly used in animals and unlike paternal apomixis, obligate androgenesis implies that individuals are incapable of reproducing in a sexual manner, and therefore depend on androgenesis to reproduce.

Ploidy in androgenesis

Individuals produced through androgenesis can be either haploid or diploid (having one or two sets of chromosomes, respectively), depending on the species. Diploidy occurs through either the fusion of two haploid sperm cells or the duplication of chromosomes from one haploid sperm cell. In both cases, the offspring experience a loss of genetic variation: individuals with the genome of 2 fused sperm cells will suffer from inbreeding depression, and individuals with the genome of a duplicated sperm cell will be fully homozygous. In species with male heterogamety (males have XY or XO chromosomes and females have XX, like in most mammals), the doubling of male chromosomes will cause all offspring to be female: if the sperm carries an X chromosome, the embryo must be XX, and if it carries a Y or O, the embryo will be YY or OO, and nonviable. With sperm fusion, a quarter of fertilized eggs will be female (XX), half will be male (XO or XY), and a quarter will be nonviable (YY or OO).

Androgenesis is more common in haplodiploid species, (where sex is determined by ploidy such that males generally develop from an unfertilized egg and females from a fertilized egg), than in diploid species (where all sexes are diploid). This is because with haplodiploids, there is no requirement of the doubling of chromosomes from a haploid gamete, so no embryos are lost due to YY or OO chromosomes.

Androgenesis in non-gonochoristic species

Androgenesis is more likely to persist in hermaphrodites than in species with two distinct sexes (gonochorists) because all individuals have the ability to produce ova, so the spread of androgenesis-promoting alleles causing egg-producers to become scarce is not an issue. Androgenesis is also seen more frequently in species that already have uncommon modes of reproduction such as hybridogenesis and parthenogenesis, and is sometimes seen in interspecies hybridization.

Induced androgenesis

Humans sometimes induce androgenesis to create clonal lines in plants (specifically crops), fish, and silkworms. A common method of inducing androgenesis is through irradiation. Egg cells can have their nuclei inactivated by gamma ray, UV, or X-ray radiation before being fertilized with sperm or pollen. A 2015 study was successful in producing zebrafish androgenones by cold-shocking just-fertilized eggs, which prevents the first cleavage event that doubles the chromosome number after parthenogenesis, and then heat-shocking them to double their chromosome number.

Embryo

From Wikipedia, the free encyclopedia
 
Embryo
A human embryo seven weeks after conception (nine weeks gestational age)
 
An embryo is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres (4-cell stage) are arranged as a solid ball that when reaching a certain size, called a morula, (16-cell stage) takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals.

The mammalian blastocyst hatches before implantating into the endometrial lining of the womb. Once implanted the embryo will continue its development through the next stages of gastrulation, neurulation, and organogenesis. Gastrulation is the formation of the three germ layers that will form all of the different parts of the body. Neurulation forms the nervous system, and organogenesis is the development of all the various tissues and organs of the body.

A newly developing human is typically referred to as an embryo until the ninth week after conception, when it is then referred to as a fetus. In other multicellular organisms, the word "embryo" can be used more broadly to any early developmental or life cycle stage prior to birth or hatching.

Etymology

First attested in English in the mid-14c., the word embryon derives from Medieval Latin embryo, itself from Greek ἔμβρυον (embruon), lit. "young one", which is the neuter of ἔμβρυος (embruos), lit. "growing in", from ἐν (en), "in" and βρύω (bruō), "swell, be full"; the proper Latinized form of the Greek term would be embryum.

Development

Animal embryos

Embryos (and one tadpole) of the wrinkled frog (Rana rugosa)
Mouse and snake embryos

In animals, fertilization begins the process of embryonic development with the creation of a zygote, a single cell resulting from the fusion of gametes (e.g. egg and sperm). The development of a zygote into a multicellular embryo proceeds through a series of recognizable stages, often divided into cleavage, blastula, gastrulation, and organogenesis.

Cleavage is the period of rapid mitotic cell divisions that occur after fertilization. During cleavage, the overall size of the embryo does not change, but the size of individual cells decrease rapidly as they divide to increase the total number of cells. Cleavage results in a blastula.

Depending on the species, a blastula or blastocyst stage embryo can appear as a ball of cells on top of yolk, or as a hollow sphere of cells surrounding a middle cavity. The embryo's cells continue to divide and increase in number, while molecules within the cells such as RNAs and proteins actively promote key developmental processes such as gene expression, cell fate specification, and polarity. Before implanting into the uterine wall the embryo is sometimes known as the pre-implantation embryo or pre-implantation conceptus. Sometimes this is called the pre-embryo a term employed to differentiate from an embryo proper in relation to embryonic stem cell discourses.

Gastrulation is the next phase of embryonic development, and involves the development of two or more layers of cells (germinal layers). Animals that form two layers (such as Cnidaria) are called diploblastic, and those that form three (most other animals, from flatworms to humans) are called triploblastic. During gastrulation of triploblastic animals, the three germinal layers that form are called the ectoderm, mesoderm, and endoderm. All tissues and organs of a mature animal can trace their origin back to one of these layers. For example, the ectoderm will give rise to the skin epidermis and the nervous system, the mesoderm will give rise to the vascular system, muscles, bone, and connective tissues, and the endoderm will give rise to organs of the digestive system and epithelium of the digestive system and respiratory system. Many visible changes in embryonic structure happen throughout gastrulation as the cells that make up the different germ layers migrate and cause the previously round embryo to fold or invaginate into a cup-like appearance.

Past gastrulation, an embryo continues to develop into a mature multicellular organism by forming structures necessary for life outside of the womb or egg. As the name suggests, organogenesis is the stage of embryonic development when organs form. During organogenesis, molecular and cellular interactions prompt certain populations of cells from the different germ layers to differentiate into organ-specific cell types. For example, in neurogenesis, a subpopulation of cells from the ectoderm segregate from other cells and further specialize to become the brain, spinal cord, or peripheral nerves.

The embryonic period varies from species to species. In human development, the term fetus is used instead of embryo after the ninth week after conception, whereas in zebrafish, embryonic development is considered finished when a bone called the cleithrum becomes visible. In animals that hatch from an egg, such as birds, a young animal is typically no longer referred to as an embryo once it has hatched. In viviparous animals (animals whose offspring spend at least some time developing within a parent's body), the offspring is typically referred to as an embryo while inside of the parent, and is no longer considered an embryo after birth or exit from the parent. However, the extent of development and growth accomplished while inside of an egg or parent varies significantly from species to species, so much so that the processes that take place after hatching or birth in one species may take place well before those events in another. Therefore, according to one textbook, it is common for scientists to interpret the scope of embryology broadly as the study of the development of animals.

Plant embryos

The inside of a Ginkgo seed, showing the embryo

Flowering plants (angiosperms) create embryos after the fertilization of a haploid ovule by pollen. The DNA from the ovule and pollen combine to form a diploid, single-cell zygote that will develop into an embryo. The zygote, which will divide multiple times as it progresses throughout embryonic development, is one part of a seed. Other seed components include the endosperm, which is tissue rich in nutrients that will help support the growing plant embryo, and the seed coat, which is a protective outer covering. The first cell division of a zygote is asymmetric, resulting in an embryo with one small cell (the apical cell) and one large cell (the basal cell). The small, apical cell will eventually give rise to most of the structures of the mature plant, such as the stem, leaves, and roots. The larger basal cell will give rise to the suspensor, which connects the embryo to the endosperm so that nutrients can pass between them. The plant embryo cells continue to divide and progress through developmental stages named for their general appearance: globular, heart, and torpedo. In the globular stage, three basic tissue types (dermal, ground, and vascular) can be recognized. The dermal tissue will give rise to the epidermis or outer covering of a plant, ground tissue will give rise to inner plant material that functions in photosynthesis, resource storage, and physical support, and vascular tissue will give rise to connective tissue like the xylem and phloem that transport fluid, nutrients, and minerals throughout the plant. In heart stage, one or two cotyledons (embryonic leaves) will form. Meristems (centers of stem cell activity) develop during the torpedo stage, and will eventually produce many of the mature tissues of the adult plant throughout its life. At the end of embryonic growth, the seed will usually go dormant until germination. Once the embryo begins to germinate (grow out from the seed) and forms its first true leaf, it is called a seedling or plantlet.

Plants that produce spores instead of seeds, like bryophytes and ferns, also produce embryos. In these plants, the embryo begins its existence attached to the inside of the archegonium on a parental gametophyte from which the egg cell was generated. The inner wall of the archegonium lies in close contact with the "foot" of the developing embryo; this "foot" consists of a bulbous mass of cells at the base of the embryo which may receive nutrition from its parent gametophyte. The structure and development of the rest of the embryo varies by group of plants.

Since all land plants create embryos, they are collectively referred to as embryophytes (or by their scientific name, Embryophyta). This, along with other characteristics, distinguishes land plants from other types of plants, such as algae, which do not produce embryos.

Research and technology

Biological processes

Embryos from numerous plant and animal species are studied in biological research laboratories across the world to learn about topics such as stem cells, evolution and development, cell division, and gene expression. Examples of scientific discoveries made while studying embryos that were awarded the Nobel Prize in Physiology or Medicine include the Spemann-Mangold organizer, a group of cells originally discovered in amphibian embryos that give rise to neural tissues, and genes that give rise to body segments discovered in Drosophila fly embryos by Christiane Nüsslein-Volhard and Eric Wieschaus.

Assisted reproductive technology

Creating and/or manipulating embryos via assisted reproductive technology (ART) is used for addressing fertility concerns in humans and other animals, and for selective breeding in agricultural species. Between the years 1987 and 2015, ART techniques including in vitro fertilization (IVF) were responsible for an estimated one million human births in the United States alone. Other clinical technologies include preimplantation genetic diagnosis (PGD), which can identify certain serious genetic abnormalities, such as aneuploidy, prior to selecting embryos for use in IVF. Some have proposed (or even attempted - see He Jiankui affair) genetic editing of human embryos via CRISPR-Cas9 as a potential avenue for preventing disease; however, this has been met with widespread condemnation from the scientific community.

ART techniques are also used to improve the profitability of agricultural animal species such as cows and pigs by enabling selective breeding for desired traits and/or to increase numbers of offspring. For example, when allowed to breed naturally, cows typically produce one calf per year, whereas IVF increases offspring yield to 9–12 calves per year. IVF and other ART techniques, including cloning via interspecies somatic cell nuclear transfer (iSCNT), are also used in attempts to increase the numbers of endangered or vulnerable species, such as Northern white rhinos, cheetahs, and sturgeons.

Cryoconservation of plant and animal biodiversity

Cryoconservation of genetic resources involves collecting and storing the reproductive materials, such as embryos, seeds, or gametes, from animal or plant species at low temperatures in order to preserve them for future use. Some large-scale animal species cryoconservation efforts include "frozen zoos" in various places around the world, including in the UK's Frozen Ark, the Breeding Centre for Endangered Arabian Wildlife (BCEAW) in the United Arab Emirates, and the San Diego Zoo Institute for Conservation in the United States. As of 2018, there were approximately 1,700 seed banks used to store and protect plant biodiversity, particularly in the event of mass extinction or other global emergencies. The Svalbard Global Seed Vault in Norway maintains the largest collection of plant reproductive tissue, with more than a million samples stored at −18 °C (0 °F).

Fossilized embryos

Fossilized animal embryos are known from the Precambrian, and are found in great numbers during the Cambrian period. Even fossilized dinosaur embryos have been discovered.

Environmental remediation

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