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Thursday, March 10, 2022

Ancient Egyptian race controversy

From Wikipedia, the free encyclopedia
https://en.wikipedia.org/wiki/Ancient_Egyptian_race_controversy

The question of the race of ancient Egyptians was raised historically as a product of the early racial concepts of the 18th and 19th centuries, and was linked to models of racial hierarchy primarily based on craniometry and anthropometry. A variety of views circulated about the racial identity of the Egyptians and the source of their culture. Some scholars argued that ancient Egyptian culture was influenced by other Afroasiatic-speaking populations in North Africa or the Middle East, while others pointed to influences from various Nubian groups or populations in Europe. In more recent times some writers continued to challenge the mainstream view, some focusing on questioning the race of specific notable individuals such as the king represented in the Great Sphinx of Giza, native Egyptian pharaoh Tutankhamun, Egyptian Queen Tiye, and Greek Ptolemaic queen Cleopatra VII.

Mainstream scholars reject the notion that Egypt was a white or black civilization; they maintain that, despite the phenotypic diversity of Ancient and present-day Egyptians, applying modern notions of black or white races to ancient Egypt is anachronistic. In addition, scholars reject the notion, implicit in the notion of a black or white Egypt hypothesis, that Ancient Egypt was racially homogeneous; instead, skin color varied between the peoples of Lower Egypt, Upper Egypt, and Nubia, who in various eras rose to power in Ancient Egypt. Within Egyptian history, despite multiple foreign invasions, the demographics were not shifted substantially by large migrations.

Origins

In the 18th century, Constantin François de Chassebœuf, comte de Volney, wrote about his thoughts on the contentions regarding the race of the ancient Egyptians. In one translation, he noted that "the Copts are the proper representatives of the Ancient Egyptians" due to their "jaundiced and fumed skin, which is neither Greek, Negro nor Arab, their full faces, their puffy eyes, their crushed noses, and their thick lips...the ancient Egyptians were true negroes of the same type as all native born Africans". Volney also said that the Sphinx gave him the key to the riddle as to why all the Egyptians he saw across the country "have a bloated face, puffed-up eyes, flat nose, thick lips – in a word, the true face of the mulatto." He wrote he was tempted to attribute it to the climate, but upon visiting the Sphinx, its appearance gave him the answer; "seeing that head, typically negro in all its features", Volney saw it as the "true solution to the enigma (of how the modern Egyptians came to have their 'mulatto' appearance)". He goes on to postulate, "the Copts were "true negroes" of the same stock as all the autochthonous peoples of Africa" and they "after some centuries of mixing..., must have lost the full blackness of its original color."

Another early example of the controversy is an article published in The New-England Magazine of October 1833, where the authors dispute a claim that "Herodotus was given as authority for their being negroes." They point out with reference to tomb paintings: "It may be observed that the complexion of the men is invariably red, that of the women yellow; but neither of them can be said to have anything in their physiognomy at all resembling the Negro countenance."

A few years later, in 1839, Jean-François Champollion stated in his work Egypte Ancienne that the Egyptians and Nubians are represented in the same manner in tomb paintings and reliefs, further suggesting that: "In the Copts of Egypt, we do not find any of the characteristic features of the ancient Egyptian population. The Copts are the result of crossbreeding with all the nations that successfully dominated Egypt. It is wrong to seek in them the principal features of the old race." This memoir was made in the context of the first tribes that would have inhabited Egypt, his opinion was noted after his return from Nubia. In 1839, Champollion's and Volney's claims were disputed by Jacques Joseph Champollion-Figeac, who blamed a misunderstanding of the ancients for spreading a false impression of a "Negro" Egypt, stating "the two physical traits of black skin and wooly hair are not enough to stamp a race as negro" and "the opinion that the ancient population of Egypt belonged to the Negro African race, is an error long accepted as the truth. ... Volney's conclusion as to the Negro origin of the ancient Egyptian civilization is evidently forced and inadmissible."

Foster summarized the early 19th century "controversy over the ethnicity of the ancient Egyptians" as a debate of conflicting theories regarding the Hamites. "In ancient times, the Hamites, who developed the civilization of Egypt, were considered Black." Foster describes the 6th century CE curse of Ham theory, which began "in the Babylonian Talmud, a collection of oral traditions of the Jews, that the sons of Ham are cursed by being black." Foster said "throughout the Middle Ages and to the end of the eighteenth century, the Negro was seen by Europeans as a descendant of Ham." In the early 19th century, "after Napolean's expedition to Egypt, the Hamites began to be viewed as having been Caucasians." However, "Napolean's scientists concluded that the Egyptians were Negroid." Napoleon's colleagues referenced prior "well-known books" by Constantin François de Chassebœuf, comte de Volney and Vivant Denon that described Ancient Egyptians as "negroid". Finally, Foster concludes, "it was at this point that Egypt became the focus of much scientific and lay interest, the result of which was the appearance of many publications whose sole purpose was to prove that the Egyptians were not Black, and therefore capable of developing such a high civilization."

The debate over the race of the ancient Egyptians intensified during the 19th century movement to abolish slavery in the United States, as arguments relating to the justifications for slavery increasingly asserted the historical, mental and physical inferiority of black people. For example, in 1851, John Campbell directly challenged the claims by Champollion and others regarding the evidence for a black Egypt, asserting "There is one great difficulty, and to my mind an insurmountable one, which is that the advocates of the negro civilization of Egypt do not attempt to account for, how this civilization was lost.... Egypt progressed, and why, because it was Caucasian." The arguments regarding the race of the Egyptians became more explicitly tied to the debate over slavery in the United States, as tensions escalated towards the American Civil War. In 1854, Josiah C. Nott with George Glidden set out to prove: "that the Caucasian or white, and the Negro races were distinct at a very remote date, and that the Egyptians were Caucasians." Samuel George Morton, a physician and professor of anatomy, concluded that "Negroes were numerous in Egypt, but their social position in ancient times was the same that it now is [in the United States], that of servants and slaves." In the early 20th century, Flinders Petrie, a professor of Egyptology at the University of London, in turn spoke of "a black queen", Ahmose-Nefertari, who was the "divine ancestress of the XVIIIth dynasty". He described her physically as "the black queen Aohmes Nefertari had an aquiline nose, long and thin, and was of a type not in the least prognathous".

Position of modern scholarship

Modern scholars who have studied ancient Egyptian culture and population history have responded to the controversy over the race of the ancient Egyptians in various ways.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, the "Black Hypothesis" met with "profound" disagreement by scholars. Similarly, none of the participants voiced support for an earlier postulation that Egyptians were "white with a dark, even black, pigmentation." The arguments for all sides are recorded in the UNESCO publication General History of Africa, with the "Origin of the Egyptians" chapter being written by Cheikh Anta Diop, a proponent of the "Black Hypothesis". At the 1974 UNESCO conference, most participants concluded that the ancient Egyptian population was indigenous to the Nile Valley, and was made up of people from north and south of the Sahara who had a range of skin colors.

Since the late 20th century, as the science of human population genetics has advanced, most biological anthropologists have come to reject the notion of race as having any validity in the study of human biology.

Frank M. Snowden wrote: "Egyptians, Greeks and Romans attached no special stigma to the colour of the skin and developed no hierarchical notions of race whereby highest and lowest positions in the social pyramid were based on colour."

Frank J. Yurco wrote in 1990: "When you talk about Egypt, it's just not right to talk about black or white .... To take the terminology here in the United States and graft it onto Africa is anthropologically inaccurate". Yurco added that "We are applying a racial divisiveness to Egypt that they would never have accepted, They would have considered this argument absurd, and that is something we could really learn from." Yurco wrote in 1996 that "the peoples of Egypt, the Sudan, and much of North-East Africa are generally regarded as a Nilotic continuity, with widely ranging physical features (complexions light to dark, various hair and craniofacial types)". In a 1989 article, he elaborated: "In short, ancient Egypt, like modern Egypt, consisted of a very heterogeneous population".

Bernard R. Ortiz De Montellano wrote in 1993: "The claim that all Egyptians, or even all the pharaohs, were black, is not valid. Most scholars believe that Egyptians in antiquity looked pretty much as they look today, with a gradation of darker shades toward the Sudan".

Nancy Lovell wrote in 1999 that studies of skeletal remains indicate that the physical characteristics of ancient southern Egyptians and Nubians were "within the range of variation" for both ancient and modern indigenous peoples of the Sahara and tropical Africa, and that the distribution of population characteristics "seems to follow a clinal pattern from south to north", which may be explained by natural selection as well as gene flow between neighboring populations. She also wrote that the archaeological and inscriptional evidence for contact between Egypt and Syro-Palestine "suggests that gene flow between these areas was very likely," and that the early Nile Valley populations were "part of an African lineage, but exhibiting local variation".

Stuart Tyson Smith wrote in 2001: "Any characterization of race of the ancient Egyptians depends on modern cultural definitions, not on scientific study. Thus, by modern American standards it is reasonable to characterize the Egyptians as 'black', while acknowledging the scientific evidence for the physical diversity of Africans."

Donald B. Redford wrote in 2004 that: "The old notion of waves of "races" flowing up the Nile Valley, effecting cultural change and improvement, is now known to be as erroneous as it was simplistic. New ideas need not come by means of invasion: occasionally they are indigenous and may parallel similar discoveries elsewhere which are wholly unrelated." He also wrote: "It would be interesting to know how the Nubians reacted to the racial attitudes Egypt manifested towards them, and in particular whether they shared a similar xenophobic aversion to all Egyptians."

Barry J. Kemp wrote in 2007 that the black/white argument, though politically understandable, is an oversimplification that hinders an appropriate evaluation of the scientific data on the ancient Egyptians since it does not take into consideration the difficulty in ascertaining complexion from skeletal remains. It also ignores the fact that Africa is inhabited by many other populations besides Bantu-related ("Negroid") groups. He wrote that in reconstructions of life in ancient Egypt, modern Egyptians would therefore be the most logical and closest approximation to the ancient Egyptians.

S. O. Y. Keita wrote in 2008 that "There is no scientific reason to believe that the primary ancestors of the Egyptian population emerged and evolved outside of northeast Africa.... The basic overall genetic profile of the modern population is consistent with the diversity of ancient populations that would have been indigenous to northeastern Africa and subject to the range of evolutionary influences over time, although researchers vary in the details of their explanations of those influences."

Barbara Mertz wrote in 2011: "Egyptian civilization was not Mediterranean or African, Semitic or Hamitic, black or white, but all of them. It was, in short, Egyptian."

Kathryn Bard wrote in 2014: "Egyptians were the indigenous farmers of the lower Nile valley, neither black nor white as races are conceived of today".

Nicky Nielsen wrote in 2020: "Ancient Egypt was neither black nor white, and the repeated attempt by advocates of either ideology to seize the ownership of ancient Egypt simply perpetuates an old tradition: one of removing agency and control of their heritage from the modern population living along the banks of the Nile."

Marc Van De Mieroop wrote in 2021: "Some scholars have tried to determine what Egyptians could have looked like by comparing their skeletal remains with those of recent populations, but the samples are so limited and the interpretations so fraught with uncertainties that this is an unreliable approach". He concluded that ancient Egypt's "location at the edge of northeast Africa and its geography as a corridor between that continent and Asia opened it up to influences from all directions, in terms of both culture and of demography."

Some modern views on bias in Egyptology

Various scholars have highlighted the role of colonial racism in shaping the attitudes of early Egyptologists, and criticized the continued over-representation of North American and European perspectives in the field. Diop in his work, "The African Origin of Civilization" argued that the prevailing views in Egyptology were driven by biased scholarship and colonial attitudes. Similarily, Bruce Trigger wrote that early modern scholarship on the Nile Valley populations had been "marred by a confusion of race, language, and culture and by an accompanying racism".

Frank Martin, critical of bias in Western scholarship and the shifting consensus, stated in 1984 that "Egyptologists, archaeologists, anthropologists, classicists, and scholars from other disciplines who have touched on the question almost invariably have asserted that Pharaonic Egyptians were whites. At first, many even dogmatically asserted that Egyptians were of Asiatic origin. That untenable position seems to have been abandoned by most serious contemporary scholars, but these same scholars have been reluctant to acknowledge the fact of the African origin of Egyptians in the sense of their being racially as well as culturally and linguistically related to Africans to the south and west of Egypt, thus leaving us with a people who seem to stand by themselves and have no ethnic and linguistic ties with any other people”.

Toby Wilkinson in 1999 outlined that "The Theory of the 'dynastic race'- a 'master race" of invaders from the east, to be responsible for imposing civilisation on the 'primitive' and unsophisticated indigenous Egyptians - had been articlated by Petrie (1939)...was still being espoused enthusiastically by scholars such as Emery (1961) and Edwards (1971)" although he felt that there was a change in perception among scholars at the time.

Stephen Quirke wrote in 2011 that the UNESCO-sponsored conference on the General History of Africa in 1974 "did not change the Eurocentric climate of research" and of the need to incorporate both African-centred studies and White European, academic persepectives. He later outlines that "research conferences and publications on the history and language of Kemet [Egypt] remain dominated.....by those brought up and trained in European, not African societies and languages (which include Arabic)".

Sally-Ann Ashton wrote in 2011: "The fact that Ancient Egypt is forced to justify its African identity through its geographical location has not gone unnoticed....critics of the mainstream Eurocentric view of Ancient Egypt claim that not only is the connection between Egypt and Africa neglected, it is consciously denied". She later outlines "This is partly the legacy of the "rediscovery" of Egypt by Europe at the end of the 18th century. In addition to this historical context, Egyptology as a discipline is dominated by scholars who are White Europeans or North Americans".

Uros Matic wrote in 2018 that the residue of scientific racism and old approaches were still present in modern archaeology and historiography of ancient Egypt and Nubia. He outlines that "although later authors such as such as Trigger and Török distanced themselves from racial science and anthropology, they kept this approach to iconography. What we can notice here are the remnants of the old colonialist episteme which appear in the works of Trigger, Török, and Donald B. Redford, however now being part of a new episteme". Matic concludes that "Epistemological de-colonization of our discipline should not only be a postcolonial criticism of its past, but a fundamental rethinking of its colonial remnants in the present, and how these remnants still shape our interpretations of Egypt and Nubia".

Stuart Tyson Smith wrote in 2018 that a common practice among Egyptologists was to "divorce Egypt from its proper northeast African context, instead framing it as fundamentally part of a Near Eastern or “Mediterranean” economic, social and political sphere, hardly African at all or at best a crossroad between the Near East, the eastern Mediterranean and Africa, which carries with it the implication that it is ultimately not really part of Africa". He explicitly criticises Van De Mieroop's comments that ancient Egypt was clearly ‘in Africa’ it was not so clearly‘of Africa’ as reflecting "long-standing Egyptological biases". He concludes that the interrelated cultural features shared between northeast African dynamic and Pharonic Egypt are not “survivals” or coincidence, but shared traditions with common origins in the deep past".

Marc Van De Mieroop wrote in 2021: "It was only recently that traditional scholarship started to acknowledge the African background of Egyptian culture, partly in response to world history's aim to replace dominant western-centered narratives with others than focused more on the contributions of other regions, including Africa. At the same time, primarily African diaspora communities wanted the continent's ancient history to be approached outside a Eurocentric context, and insisted, for example, on the use of ancient Egyptian term kemet instead of the European one".

Keith Crawford in 2021 presented a critique of the "Black Pharaohs" narrative accepted among mainstream scholars in which the Twenty-fifth Dynasty rulers were the only dynasty of Black African origin and academic representation of Egyptian-Kushite interactions. He concludes "The separation of Egypt from Africa, beginning nearly two centuries ago, resulted from Egyptologists, historians, and anthropologists interpreting archaeological finds and physical remains through a prism blurred by the racism of the time. These views have persisted to this day, despite overwhelming evidence that refutes them".

Genetic studies have been criticised by several scholars for a range of methodological problems and providing misleading, interpretations on racial classifications. Specifically, Keita and Kittles argue that DNA studies applied to the Nile Valley region have downplayed or excluded data on comparable, African populations in order to maintain certain racial models along with pre-selected data categories. Boyce and Keita in a later study, argue that certain studies have adopted a selective approach in sampling, such as using samples drawn mostly from northern (Lower) Egypt, which has historically had the presence of more foreigners from the Mediterranean and the Near East, and using those samples as representing the rest of Egypt. Thus, excluding the 'darker' south or Upper Egypt which presents a false impression of Egyptian variability. The authors also note that chromosonial patterns have featured inconsistent labelling such as Haplotype V as seen the with use of misleading terms like "Arabic" to describe it, implying this haplotype is of 'Middle Eastern' origins. However, when the hapotype V variant is looked at in context, it does have a very high prevalence in African countries above the Sahara and in Ethiopia.

Present-day controversies

Today the issues regarding the race of the ancient Egyptians are "troubled waters which most people who write about ancient Egypt from within the mainstream of scholarship avoid." The debate, therefore, takes place mainly in the public sphere and tends to focus on a small number of specific issues.

Tutankhamun

Several scholars, including Diop, have claimed that Tutankhamun was black, and have protested that attempted reconstructions of Tutankhamun's facial features (as depicted on the cover of National Geographic magazine) have represented the king as "too white". Among these writers was Chancellor Williams, who argued that King Tutankhamun, his parents, and grandparents were black.

Forensic artists and physical anthropologists from Egypt, France, and the United States independently created busts of Tutankhamun, using a CT-scan of the skull. Biological anthropologist Susan Anton, the leader of the American team, said the race of the skull was "hard to call". She stated that the shape of the cranial cavity indicated an African, while the nose opening suggested narrow nostrils, which is usually considered to be a European characteristic. The skull was thus concluded to be that of a North African. Other experts have argued that neither skull shapes nor nasal openings are a reliable indication of race.

Although modern technology can reconstruct Tutankhamun's facial structure with a high degree of accuracy, based on CT data from his mummy, determining his skin tone and eye color is impossible. The clay model was therefore given a coloring, which, according to the artist, was based on an "average shade of modern Egyptians".

Queen Tiye, grandmother of Tutankhamun

Terry Garcia, National Geographic's executive vice president for mission programs, said, in response to some of those protesting against the Tutankhamun reconstruction:

The big variable is skin tone. North Africans, we know today, had a range of skin tones, from light to dark. In this case, we selected a medium skin tone, and we say, quite up front, 'This is midrange.' We will never know for sure what his exact skin tone was or the color of his eyes with 100% certainty.... Maybe in the future, people will come to a different conclusion.

When pressed on the issue by American activists in September 2007, the Secretary General of the Egyptian Supreme Council of Antiquities, Zahi Hawass stated "Tutankhamun was not black."

In a November 2007 publication of Ancient Egypt magazine, Hawass asserted that none of the facial reconstructions resemble Tut and that, in his opinion, the most accurate representation of the boy king is the mask from his tomb. The Discovery Channel commissioned a facial reconstruction of Tutankhamun, based on CT scans of a model of his skull, back in 2002.

In 2011, the genomics company iGENEA launched a Tutankhamun DNA project based on genetic markers that it indicated it had culled from a Discovery Channel special on the pharaoh. According to the firm, the microsatellite data suggested that Tutankhamun belonged to the haplogroup R1b1a2, the most common paternal clade among males in Western Europe. Carsten Pusch and Albert Zink, who led the unit that had extracted Tutankhamun's DNA, chided iGENEA for not liaising with them before establishing the project. After examining the footage, they also concluded that the methodology the company used was unscientific with Putsch calling them "simply impossible".

Cleopatra

The race and skin color of Cleopatra VII, the last active Hellenistic ruler of the Macedonian Greek Ptolemaic dynasty of Egypt, established in 323 BCE, has also caused some debate, although generally not in scholarly sources. For example, the article "Was Cleopatra Black?" was published in Ebony magazine in 2012, and an article about Afrocentrism from the St. Louis Post-Dispatch mentions the question, too. Mary Lefkowitz, Professor Emerita of Classical Studies at Wellesley College, traces the main origins of the black Cleopatra claim to the 1946 book by J.A. Rogers called "World's Great Men of Color.", although noting that the idea of Cleopatra as black goes back to at least the 19th century. Lefkowitz refutes Rogers' hypothesis, on various scholarly grounds. The black Cleopatra claim was further revived in an essay by afrocentrist John Henrik Clarke, chair of African history at Hunter College, entitled "African Warrior Queens." Lefkowitz notes the essay includes the claim that Cleopatra described herself as black in the New Testament's Book of Acts – when in fact Cleopatra had died more than sixty years before the death of Jesus Christ.

The Berlin Cleopatra, a Roman sculpture and marble portrait of the queen in the Altes Museum, 1st century BC 

Scholars identify Cleopatra as essentially of Greek ancestry with some Persian and Syrian ancestry, based on the fact that her Macedonian Greek family (the Ptolemaic dynasty) had intermingled with the Seleucid aristocracy of the time. Grant states that Cleopatra probably had not a drop of Egyptian blood and that she "would have described herself as Greek." Roller notes that "there is absolutely no evidence" that Cleopatra was racially black African as claimed by what he dismisses as generally not "credible scholarly sources." Cleopatra's official coinage (which she would have approved) and the three portrait busts of her which are considered authentic by scholars, all match each other, and they portray Cleopatra as a Greek woman. Polo writes that Cleopatra's coinage presents her image with certainty, and asserts that the sculpted portrait of the "Berlin Cleopatra" head is confirmed as having a similar profile.

In 2009, a BBC documentary speculated that Cleopatra might have been part North African. This was based largely on the claims of Hilke Thür of the Austrian Academy of Sciences, who in the 1990s had examined a headless skeleton of a female child in a 20 BCE tomb in Ephesus (modern Turkey), together with the old notes and photographs of the now-missing skull. Thür hypothesized the body as that of Arsinoe, half-sister to Cleopatra. Arsinoe and Cleopatra shared the same father (Ptolemy XII Auletes) but had different mothers, with Thür claiming the alleged African ancestry came from the skeleton's mother. To date it has never been definitively proved that the skeleton is that of Arsinoe IV. When a DNA test attempted to determine the identity of the child, it was impossible to get an accurate reading since the bones had been handled too many times, and the skull had been lost in Germany during World War II. Numerous studies have shown that cranial variation has a low correlation with race, and rather that cranial variation was strongly correlated with climate variables. Mary Beard states that the age of the skeleton is too young to be that of Arsinoe (the bones said to be that of a 15–18-year-old child, with Arsinoe being around her mid twenties at her death).

Great Sphinx of Giza

The Sphinx in profile in 2010

The identity of the model for the Great Sphinx of Giza is unknown. Most experts believe that the face of the Sphinx represents the likeness of the Pharaoh Khafra, although a few Egyptologists and interested amateurs have proposed different hypotheses.

An early description of the Sphinx, "typically negro in all its features", is recorded in the travel notes of a French scholar, Volney, who visited Egypt between 1783 and 1785 along with French novelist Gustave Flaubert. A similar description was given in the "well-known book" by Vivant Denon, where he described the sphinx as "the character is African; but the mouth, the lips of which are thick." Following Volney, Denon, and other early writers, numerous Afrocentric scholars, such as Du Bois, Diop and Asante have characterized the face of the Sphinx as Black, or "Negroid".

American geologist Robert M. Schoch has written that the "Sphinx has a distinctive African, Nubian, or Negroid aspect which is lacking in the face of Khafre", but he was described by others such as Ronald H. Fritze and Mark Lehner of being a "pseudoscientific writer". David S. Anderson writes in Lost City, Found Pyramid: Understanding Alternative Archaeologies and Pseudoscientific Practices that Van Sertima's claim that "the sphinx was a portrait statue of the black pharoah Khafre" is a form of "pseudoarchaeology" not supported by evidence. He compares it to the claim that Olmec colossal heads had "African origins", which is not taken seriously by Mesoamerican scholars such as Richard Diehl and Ann Cyphers.

Kemet

km in Egyptian hieroglyphs
km biliteral kmt (place) kmt (people)
km
km
t O49
km
t
A1B1Z3

Ancient Egyptians referred to their homeland as Kmt (conventionally pronounced as Kemet). According to Cheikh Anta Diop, the Egyptians referred to themselves as "Black" people or kmt, and km was the etymological root of other words, such as Kam or Ham, which refer to Black people in Hebrew tradition. A review of David Goldenberg's The Curse of Ham: Race and Slavery in Early Judaism, Christianity and Islam states that Goldenberg "argues persuasively that the biblical name Ham bears no relationship at all to the notion of blackness and as of now is of unknown etymology". Diop, William Leo Hansberry, and Aboubacry Moussa Lam have argued that kmt was derived from the skin color of the Nile valley people, which Diop claimed was black. The claim that the ancient Egyptians had black skin has become a cornerstone of Afrocentric historiography.

Mainstream scholars hold that kmt means "the black land" or "the black place", and that this is a reference to the fertile black soil that was washed down from Central Africa by the annual Nile inundation. By contrast the barren desert outside the narrow confines of the Nile watercourse was called dšrt (conventionally pronounced deshret) or "the red land". Raymond Faulkner's Concise Dictionary of Middle Egyptian translates kmt into "Egyptians", Gardiner translates it as "the Black Land, Egypt".

At the UNESCO Symposium in 1974, Sauneron, Obenga, and Diop concluded that KMT and KM meant black. However, Sauneron clarified that the adjective Kmtyw means "people of the black land" rather than "black people", and that the Egyptians never used the adjective Kmtyw to refer to the various black peoples they knew of, they only used it to refer to themselves.

Ancient Egyptian art

Ancient Egyptian tombs and temples contained thousands of paintings, sculptures, and written works, which reveal a great deal about the people of that time. However, their depictions of themselves in their surviving art and artifacts are rendered in sometimes symbolic, rather than realistic, pigments. As a result, ancient Egyptian artifacts provide sometimes conflicting and inconclusive evidence of the ethnicity of the people who lived in Egypt during dynastic times.

In their own art, "Egyptians are often represented in a color that is officially called dark red", according to Diop. Arguing against other theories, Diop quotes Champollion-Figeac, who states, "one distinguishes on Egyptian monuments several species of blacks, differing...with respect to complexion, which makes Negroes black or copper-colored." Regarding an expedition by King Sesostris, Cherubini states the following concerning captured southern Africans, "except for the panther skin about their loins, are distinguished by their color, some entirely black, others dark brown. University of Chicago scholars assert that Nubians are generally depicted with black paint, but the skin pigment used in Egyptian paintings to refer to Nubians can range "from dark red to brown to black". This can be observed in paintings from the tomb of the Egyptian Huy, as well as Ramses II's temple at Beit el-Wali. Also, Snowden indicates that Romans had accurate knowledge of "negroes of a red, copper-colored complexion ... among African tribes".

Conversely, in 2003 Najovits wrote that "Egyptian art depicted Egyptians on the one hand and Nubians and other blacks on the other hand with distinctly different ethnic characteristics and depicted this abundantly and often aggressively. The Egyptians accurately, arrogantly and aggressively made national and ethnic distinctions from a very early date in their art and literature." He continues, "There is an extraordinary abundance of Egyptian works of art which clearly depicted sharply contrasted reddish-brown Egyptians and black Nubians."

Barbara Mertz in 2011 wrote in Red Land, Black Land: Daily Life in Ancient Egypt: "The concept of race would have been totally alien to them [Ancient Egyptians] [..]The skin color that painters usually used for men is a reddish brown. Women were depicted as lighter in complexion, perhaps because they didn’t spend so much time out of doors. Some individuals are shown with black skins. I cannot recall a single example of the words “black,” “brown,” or “white” being used in an Egyptian text to describe a person." She gives the example of one of Thutmose III’s “sole companions”, who was Nubian or Kushite. In his funerary scroll, he is shown with dark brown skin instead of the conventional reddish brown used for Egyptians.

Table of Nations controversy

The "Table of Nations", from Lepsius: Top row, left to right: "Aamu" (Asiatics), "Nehesu" (Nubians), and "Themehu" (Libyans); middle row: a deity, "Reth" (Egyptians), "Aamu" (Asiatics).

However, Manu Ampim, a professor at Merritt College specializing in African and African American history and culture, claims in the book Modern Fraud: The Forged Ancient Egyptian Statues of Ra-Hotep and Nofret, that many ancient Egyptian statues and artworks are modern frauds that have been created specifically to hide the "fact" that the ancient Egyptians were black, while authentic artworks that demonstrate black characteristics are systematically defaced or even "modified". Ampim repeatedly makes the accusation that the Egyptian authorities are systematically destroying evidence that "proves" that the ancient Egyptians were black, under the guise of renovating and conserving the applicable temples and structures. He further accuses "European" scholars of wittingly participating in and abetting this process.

Ampim has a specific concern about the painting of the "Table of Nations" in the Tomb of Ramesses III (KV11). The "Table of Nations" is a standard painting that appears in a number of tombs, and they were usually provided for the guidance of the soul of the deceased. Among other things, it described the "four races of men" as follows: (translation by E.A. Wallis Budge) "The first are RETH, the second are AAMU, the third are NEHESU, and the fourth are THEMEHU. The RETH are Egyptians, the AAMU are dwellers in the deserts to the east and north-east of Egypt, the NEHESU are the black races, and the THEMEHU are the fair-skinned Libyans."

The archaeologist Karl Richard Lepsius documented many ancient Egyptian tomb paintings in his work Denkmäler aus Aegypten und Aethiopien. In 1913, after the death of Lepsius, an updated reprint of the work was produced, edited by Kurt Sethe. This printing included an additional section, called the "Ergänzungsband" in German, which incorporated many illustrations that did not appear in Lepsius' original work. One of them, plate 48, illustrated one example of each of the four "nations" as depicted in KV11, and shows the "Egyptian nation" and the "Nubian nation" as identical to each other in skin color and dress. Professor Ampim has declared that plate 48 is a true reflection of the original painting, and that it "proves" that the ancient Egyptians were identical in appearance to the Nubians, even though he admits no other examples of the "Table of Nations" show this similarity. He has further accused "Euro-American writers" of attempting to mislead the public on this issue.

The late Egyptologist Frank J. Yurco visited the tomb of Ramesses III (KV11), and in a 1996 article on the Ramesses III tomb reliefs he pointed out that the depiction of plate 48 in the Ergänzungsband section is not a correct depiction of what is actually painted on the walls of the tomb. Yurco notes, instead, that plate 48 is a "pastiche" of samples of what is on the tomb walls, arranged from Lepsius' notes after his death, and that a picture of a Nubian person has erroneously been labeled in the pastiche as an Egyptian person. Yurco points also to the much more recent photographs of Dr. Erik Hornung as a correct depiction of the actual paintings. (Erik Hornung, The Valley of the Kings: Horizon of Eternity, 1990). Ampim nonetheless continues to claim that plate 48 shows accurately the images that stand on the walls of KV11, and he categorically accuses both Yurco and Hornung of perpetrating a deliberate deception for the purposes of misleading the public about the true race of the ancient Egyptians.

Fayyum mummy portraits

The realistic Fayum mummy portraits show the diversity of Egyptians in the Roman period

The Roman era Fayum mummy portraits attached to coffins containing the latest dated mummies discovered in the Faiyum Oasis represent a population of both native Egyptians and those with mixed Greek heritage. The dental morphology of the mummies align more with the indigenous North African population than Greek or other later colonial European settlers.

Black queen controversy

The late British Africanist Basil Davidson stated "Whether the Ancient Egyptians were as black or as brown in skin color as other Africans may remain an issue of emotive dispute; probably, they were both. Their own artistic conventions painted them as pink, but pictures on their tombs show they often married queens shown as entirely black being from the south."

Queen Ahmose-Nefertari of the 18th Dynasty

Ahmose-Nefertari is an example. In most depictions of Ahmose-Nefertari, she is pictured with black skin, while in some instances her skin is blue or red. In 1939 Flinders Petrie said "an invasion from the south...established a black queen as the divine ancestress of the XVIIIth dynasty" He also said "a possibility of the black being symbolic has been suggested" and "Nefertari must have married a Libyan, as she was the mother of Amenhetep I, who was of fair Libyan style." In 1961 Alan Gardiner, in describing the walls of tombs in the Deir el-Medina area, noted in passing that Ahmose-Nefertari was "well represented" in these tomb illustrations, and that her countenance was sometimes black and sometimes blue. He did not offer any explanation for these colors, but noted that her probable ancestry ruled out that she might have had black blood. In 1974, Diop described Ahmose-Nefertari as "typically negroid." In the controversial book Black Athena, the hypotheses of which have been widely rejected by mainstream scholarship, Martin Bernal considered her skin color in paintings to be a clear sign of Nubian ancestry. In 1981 Michel Gitton noted that while in most artistic depictions of the queen she is pictured with black complexion, there are other cases in which she is shown with a pink, golden, blue, or dark red skin color.

Gitton called the issue of Ahmose-Nefertari's black color "a serious gap in the Egyptological research, which allows approximations or untruths". He pointed out that there is no known depiction of her painted during her lifetime (she is represented with the same light skin as other represented individuals in tomb TT15, before her deification); the earliest black skin depiction appears in tomb TT161, circa 150 years after her death. Barbara Lesko wrote in 1996 that Ahmose-Nefertari was "sometimes portrayed by later generations as having been black, although her coffin portrait gives her the typical light yellow skin of women."

In 2003, Betsy Bryan wrote in The Oxford History of Ancient Egypt that "the factors linking Amenhotep I and his mother with the necropolis region, with deified rulers, and with rejuvenation generally was visually transmitted by representations of the pair with black or blue skin – both colours of resurrection." In 2004 Aidan Dodson and Dyan Hilton recognized in a later depiction of the queen, "the black skin of a deity of resurrection" in connection to her role as a patron goddess of the Theban necropolis. Scholars such as Joyce Tyldesley, Sigrid Hodel-Hoenes, and Graciela Gestoso Singer, argued that the skin color of Ahmose-Nefertari is indicative of her role as a goddess of resurrection, since black is both the color of the fertile land of Egypt and that of Duat, the underworld. Singer recognizes that "Some scholars have suggested that this is a sign of Nubian ancestry." Singer also states a statuette of Ahmose-Nefertari at the Museo Egizio in Turin which shows her with a black face, though her arms and feet are not darkened, thus suggesting that the black coloring has an iconographic motive and does not reflect her actual appearance. In 2014, Margaret Bunson wrote that "the unusual depictions of Ahmose-Nefertari in blue-black tones of deification reflect her status and cult." In a wooden votive statue of Ahmose-Nefertari, currently in the Louvre museum, her skin was painted red, a color commonly seen symbolizing life or a higher being, or elevated status.

Historical hypotheses

Since the second half of the 20th century, typological and hierarchical models of race have increasingly been rejected by scientists, and most scholars have held that applying modern notions of race to ancient Egypt is anachronistic. The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt). At the UNESCO symposium in 1974, most participants concluded that the ancient Egyptian population was indigenous to the Nile Valley, and was made up of people from north and south of the Sahara who were differentiated by their color.

Black Egyptian hypothesis

The Black Egyptian hypothesis is the hypothesis that ancient Egypt was a "Black" civilization. Although there is consensus that Ancient Egypt was indigenous to Africa, the hypothesis that it was a "Black" civilization has met with "profound" disagreement, particularly because it rests upon a scientifically outdated conception of race.

The Black Egyptian hypothesis includes a particular focus on links to Sub Saharan cultures and the questioning of the race of specific notable individuals from Dynastic times, including Tutankhamun the person represented in the Great Sphinx of Giza, and the Greek Ptolemaic queen Cleopatra. Advocates of the Black African model rely heavily on writings from Classical Greek historians, including Strabo, Diodorus Siculus, and Herodotus. Advocates claim that these "classical" authors referred to Egyptians as "Black with woolly hair". The Greek word used was "melanchroes", and the English language translation of this Greek word is disputed, being translated by many as "dark skinned" and by many others as "black". Diop said "Herodotus applied melanchroes to both Ethiopians and Egyptians...and melanchroes is the strongest term in Greek to denote blackness." Snowden claims that Diop is distorting his classical sources and is quoting them selectively. There is dispute about the historical accuracy of the works of Herodotus – some scholars support the reliability of Herodotus while other scholars regard his works as being unreliable as historical sources, particularly those relating to Egypt.

Other claims used to support the Black Hypothesis included anthropological measurements of Egyptian mummies, testing melanin levels in a small sample of mummies, language affinities between ancient Egyptian language and sub-saharan languages, interpretations of the origin of the name Kmt, conventionally pronounced Kemet, used by the ancient Egyptians to describe themselves or their land (depending on points of view), biblical traditions, shared B blood group between Egyptians and West Africans, and interpretations of the depictions of the Egyptians in numerous paintings and statues. The hypothesis also claimed cultural affiliations, such as circumcision, matriarchy, totemism, hair braiding, head binding, and kingship cults. Artifacts found at Qustul (near Abu Simbel – Modern Sudan) in 1960–64 were seen as showing that ancient Egypt and the A-Group culture of Nubia shared the same culture and were part of the greater Nile Valley sub-stratum, but more recent finds in Egypt indicate that the Qustul rulers probably adopted/emulated the symbols of Egyptian pharaohs. Authors and critics state the hypothesis is primarily adopted by Afrocentrists.

The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt), and that modern racial categories like "Black" are social constructs inapplicable to Ancient Egypt.

Asiatic race theory

The Asiatic race theory, which has been rejected by mainstream scholarship, is the hypothesis that the ancient Egyptians were the lineal descendants of the biblical Ham, through his son Mizraim.

This theory was the most dominant view from the Early Middle Ages (c. 500 AD) all the way up to the early 19th century. The descendants of Ham were traditionally considered to be the darkest-skinned branch of humanity, either because of their geographic allotment to Africa or because of the Curse of Ham. Thus, Diop cites Gaston Maspero "Moreover, the Bible states that Mesraim, son of Ham, brother of Chus (Kush) ... and of Canaan, came from Mesopotamia to settle with his children on the banks of the Nile."

By the 20th century, the Asiatic race theory and its various offshoots were abandoned but were superseded by two related theories: the Eurocentric Hamitic hypothesis, asserting that a Caucasian racial group moved into North and East Africa from early prehistory subsequently bringing with them all advanced agriculture, technology and civilization, and the Dynastic race theory, proposing that Mesopotamian invaders were responsible for the dynastic civilization of Egypt (c. 3000 BC). In sharp contrast to the Asiatic race theory, neither of these theories proposes that Caucasians were the indigenous inhabitants of Egypt.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, none of the participants explicitly voiced support for any theory where Egyptians were Caucasian with a dark pigmentation.". The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Caucasian / Hamitic hypothesis

The Caucasian hypothesis, which has been rejected by mainstream scholarship, is the hypothesis that the Nile valley "was originally peopled by a branch of the Caucasian race". It was proposed in 1844 by Samuel George Morton, who acknowledged that Negroes were present in ancient Egypt but claimed they were either captives or servants. George Gliddon (1844) wrote: "Asiatic in their origin .... the Egyptians were white men, of no darker hue than a pure Arab, a Jew, or a Phoenician."

The similar Hamitic hypothesis, which has been rejected by mainstream scholarship, developed directly from the Asiatic Race Theory, and argued that the Ethiopid and Arabid populations of the Horn of Africa were the inventors of agriculture and had brought all civilization to Africa. It asserted that these people were Caucasians, not Negroid. It also rejected any Biblical basis despite using Hamitic as the theory's name. Charles Gabriel Seligman in his Some Aspects of the Hamitic Problem in the Anglo-Egyptian Sudan (1913) and later works argued that the ancient Egyptians were among this group of Caucasian Hamites, having arrived in the Nile Valley during early prehistory and introduced technology and agriculture to primitive natives they found there.

The Italian anthropologist Giuseppe Sergi (1901) believed that ancient Egyptians were the Eastern African (Hamitic) branch of the Mediterranean race, which he called "Eurafrican". According to Sergi, the Mediterranean race or "Eurafrican" contains three varieties or sub-races: the African (Hamitic) branch, the Mediterranean "proper" branch and the Nordic (depigmented) branch. Sergi maintained in summary that the Mediterranean race (excluding the depigmented Nordic or 'white') is: "a brown human variety, neither white nor Negroid, but pure in its elements, that is to say not a product of the mixture of Whites with Negroes or Negroid peoples". Grafton Elliot Smith modified the theory in 1911, stating that the ancient Egyptians were a dark haired "brown race", most closely "linked by the closest bonds of racial affinity to the Early Neolithic populations of the North African littoral and South Europe", and not Negroid. Smith's "brown race" is not synonymous or equivalent with Sergi's Mediterranean race. The Hamitic Hypothesis was still popular in the 1960s and late 1970s and was supported notably by Anthony John Arkell and George Peter Murdock.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, none of the participants explicitly voiced support for any theory where Egyptians were Caucasian with a dark pigmentation." The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Turanid race hypothesis

The Turanid race hypothesis, which has been rejected by mainstream scholarship, is the hypothesis that the ancient Egyptians belonged to the Turanid race, linking them to the Tatars.

It was proposed by Egyptologist Samuel Sharpe in 1846, who was "inspired" by some ancient Egyptian paintings, which depict Egyptians with sallow or yellowish skin. He said "From the colour given to the women in their paintings we learn that their skin was yellow, like that of the Mongul Tartars, who have given their name to the Mongolian variety of the human race.... The single lock of hair on the young nobles reminds us also of the Tartars."

The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Dynastic race theory

The Dynastic race theory, which has been rejected by mainstream scholarship, is the hypothesis that a Mesopotamian force had invaded Egypt in predynastic times, imposed itself on the indigenous Badarian people, and become their rulers. It further argued that the Mesopotamian-founded state or states then conquered both Upper and Lower Egypt and founded the First Dynasty of Egypt.

It was proposed in the early 20th century by Egyptologist Sir William Matthew Flinders Petrie, who deduced that skeletal remains found at pre-dynastic sites at Naqada (Upper Egypt) indicated the presence of two different races, with one race differentiated physically by a noticeably larger skeletal structure and cranial capacity. Petrie also noted new architectural styles—the distinctly Mesopotamian "niched-facade" architecture—pottery styles, cylinder seals and a few artworks, as well as numerous Predynastic rock and tomb paintings depicting Mesopotamian style boats, symbols, and figures. Based on plentiful cultural evidence, Petrie concluded that the invading ruling elite was responsible for the seemingly sudden rise of Egyptian civilization. In the 1950s, the Dynastic Race Theory was widely accepted by mainstream scholarship.

While there is clear evidence the Naqada II culture borrowed abundantly from Mesopotamia, the Naqada II period had a large degree of continuity with the Naqada I period, and the changes which did happen during the Naqada periods happened over significant amounts of time. The most commonly held view today is that the achievements of the First Dynasty were the result of a long period of cultural and political development, and the current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

The Senegalese Egyptologist Cheikh Anta Diop, fought against the Dynastic Race Theory with their own "Black Egyptian" theory and claimed, among other things, that Eurocentric scholars supported the Dynastic Race Theory "to avoid having to admit that Ancient Egyptians were black". Martin Bernal proposed that the Dynastic Race theory was conceived by European scholars to deny Egypt its African roots.

Genetic history of Africa

From Wikipedia, the free encyclopedia
 
Population structure of African populations in a broad context. ADMIXTURE analysis of 2,194 individuals from 81 populations for 16,420 SNPs reveals both well-established and novel ancestry components in African populations.

The genetic history of Africa is composed of the overall genetic history of African populations, including the regional genetic histories of North Africa, West Africa, East Africa, Central Africa, and Southern Africa, as well as the recent African origin of modern humans. The Sahara served as a trans-regional passageway and place of dwelling for people in Africa during various humid phases and periods throughout the history of Africa.

Overview

(A) the origin of the 46 African ethnic groups used in the analysis; ethnic groups from similar regions are given the same colour, but different shapes. (B) PCA shows that the first major axis of variation in Africa (PC1, y-axis) splits southern groups from the rest of Africa, each symbol represents an individual; PC2 (x-axis) reflects ethno-linguistic differences, with Niger-Congo and Nilo-Saharan speakers split from Afroasiatic speakers. (C) The third principle component (PC3, x-axis) represents geographical separation of Niger-Congo speakers, forming a cline from west to east Africans.

The people of Africa, the largest and culturally most diverse continent, are characterized by regional genetic substructure, and heterogeneity depending on the respective ethno-linguistic identity, which can be in part explained by the "multiregional evolution" of modern human lineages in various multiple regions of the African continent, as well as later admixture events, and back-migrations from Eurasia, including both highly differentiated West-Eurasian and East-Eurasian components.

Africans genetic ancestry is largely partitioned by geography and language family, with populations belonging to the same, or related, ethno-linguistic groupings showing high genetic homogeneity and coherence, however mixed ancestry in several individuals, consistent with both short- and long-range migration events followed by extensive admixture and bottleneck events, have influenced the genetic makeup and demographic structure of Africans. The historical Bantu expansion had lasting impacts on the modern demographic make up of Africa, resulting in a greater genetic and linguistic homogenization. Genetic, archeologic, and linguistic studies added extra insight into this movement: "Our results reveal a genetic continuum of Niger–Congo speaker populations across the continent and extend our current understanding of the routes, timing and extent of the Bantu migration."

Overall, different African populations display genetic diversity and substructure, but can be clustered in two distinct but partially overlapping groupings: The first and largest grouping consists of Niger-Congo-speakers and Nilo-Saharan-speakers from West Africa, Central Africa, and East Africa, which are occasionally designated as "Sub-Saharan Africans" in population genomics. The Khoisan, hunter-gatherers from Southern Africa, form an divergent branch, suggesting early isolation from other African groups. The second grouping consists of mostly, but not exclusively, Afroasiatic-speakers and Austronesian-speakers from Northern Africa and parts of the Horn of Africa, as well as from Madagascar respectively, which have received significant Eurasian admixture in varying degrees. The Niger-Congo-speakers, together with Nilo-Saharans, form a tight cluster, while the Khoisan, Afroasiatic and Austronesian groups are more diverged from other Africans, either through more diverged ancestry or from Eurasian admixture. Overall, African populations are closer to each other than to outside populations, except certain groups from coastal North Africa, which are relatively more close to Europeans and Middle Easterners.

Indigenous Africans

The indigenous population of Africa consists of Niger–Congo speakers, Nilo-Saharan speakers, the diverse Khoisan grouping, as well as several unclassified or isolated ethnolinguistic groupings (see unclassified languages of Africa). The origin of the Afroasiatic languages remains disputed, with some proposing an Middle Eastern origin, while others support an indigenous African origin.

PCA plot of genetic variation of worldwide populations. (A) Geographic coordinates of 53 populations. (B) Procrustes-transformed PCA plot of genetic variation.

The Niger–Congo languages probably originated in or near the area where these languages were spoken prior to Bantu expansion (i.e. West Africa or Central Africa). Its expansion may have been associated with the expansion of agriculture, during in the African Neolithic period, following the desiccation of the Sahara in c. 3500 BCE. Proto-Niger-Congo may have originated about 10,000 years before present in the "Green Sahara" of Africa (roughly the Sahel and southern Sahara), and that its dispersal can be correlated with the spread of the bow and arrow by migrating hunter-gatherers, which later developed agriculture.

Although the validity of the Nilo-Saharan family remains controversial, the region between Chad, Sudan, and the Central African Republic is seen as a likely candidate for its homeland prior to its dispersal around 10,000–8,000 BCE.

The Southern African hunter-gatherers (Khoisan) are suggested to represent the early diverged indigenous population of parts of Southern Africa, prior to the Bantu expansion, and were genetically isolated from surrounding populations until ~2,000 years ago, with many Khoisan samples showing high amounts of Bantu-related admixture, ranging from ~0% to up to ~87.1% of their ancestry, but also minor Eurasian-like admixture.

Out-of-Africa event

Human migration routes following Out-of-Africa.
 
Africans display a high level of genetic homogeneity with minor contributions from Eurasian populations, mostly concentrated in the Northeastern part of Africa.

The model proposes a "single origin" of Homo sapiens in Africa the taxonomic sense. Recent genetic and archeologic data suggests that Homo sapiens-subgroups originated in multiple regions of Africa, not confined to a single region of origin. The H. sapiens ancestral to proper Eurasians most likely developed in various regions of Northeastern Africa, between 300,000 and 200,000 years ago. The "recent African origin" model proposes that all modern non-African populations are substantially descended from one or several diverse waves of H. sapiens that left Africa, through the Arabian Peninsula, at about 110,000 years ago, with some proposing an earlier date.

According to a 2020 study by Durvasula et al., there are indications that 2% to 19% (≃6.6%) of the DNA of West African populations may have come from an unknown archaic hominin which split from the ancestor of humans and Neanderthals between 360 kya to 1.02 mya. However, the study also suggests that at least part of this archaic admixture is also present in Eurasians/non-Africans, and that the admixture event or events range from 0 to 124 ka B.P, which includes the period before the Out-of-Africa migration and prior to the African/Eurasian split (thus affecting in part the common ancestors of both Africans and Eurasians/non-Africans). Another academic paper from 2020 (Chen et al.) found that Africans have higher Neanderthal ancestry than previously thought. 2,504 African samples from all over Africa were analyzed and tested on Neanderthal ancestry. All African samples showed evidence for minor Neanderthal ancestry, but always at lower levels than observed in Eurasians.

Geneflow between Eurasia and African populations

Eurasian ancestry is found in many parts of Africa in varying degrees, but generally at low frequency. Eurasian admixture peaks amongst populations of Northern Africa, and among some ethnic groups of the Horn of Africa, as well as among the Malagasy people of Madagascar. A genome study (Busby et al. 2016) shows evidence for prehistoric back-migrations from Eurasian populations and following admixture with native groups.

Another study (Ramsay et al. 2018) also found evidence for Eurasian admixture in parts Africa, from both ancient and more recent migrations, being highest among populations from Northern Africa, parts of the Horn of Africa and parts of the Sahel zone:


This distinctive Eurasian admixture appears to have occurred over at least three time periods with ancient admixture in central west Africa (e.g. Yoruba from Nigeria) occurring between ~7.5 and 10.5 kya, older admixture in east Africa (e.g. Ethiopia) occurring between ~2.4 and 3.2 kya and more recent admixture between ~0.15 and 1.5 kya in some east African (e.g. Kenyan) populations.

Subsequent studies based on LD decay and haplotype sharing in an extensive set of African and Eurasian populations confirmed the presence of Eurasian signatures in west, east and southern Africans. In the west, in addition to Niger-Congo speakers from The Gambia and Mali, the Mossi from Burkina Faso showed the oldest Eurasian admixture event ~7 kya. In the east, these analyses inferred Eurasian admixture within the last 4000 years in Kenya.

A previous 2014 genome study by Hodgson et al. indicated the (West-) Eurasian admixture in the Northern Africa and the Horn of Africa was already from about 23,000 years ago. There is evidence that some Western Eurasian admixture arrived in Northeast Africa (particularly the Horn of Africa) in the Neolithic, with some moving south eventually reaching southern Africa.

Proposed migration routes of paternal lineage E, the most frequent haplogroup among Africans.
 
Proposed migration and expansion routes of the Afroasiatic languages according to the indigenous African origin model. West-Eurasian ancestry was introduced into the "North Afroasiatic heartland" and spreaded with later Afroasiatic populations, but remained largely absent from southern Afroasiatic populations.

Afroasiatic languages are suggested to have been spreaded by people with largely West Eurasian ancestry during the Neolithic Revolution, towards Northern Africa and the Horn of Africa, outgoing from the Middle East, specifically from the Levant. While there is no definitive agreement on when or where the original homeland of the Afroasiatic language family existed, many link the first speakers to the earliest farmers in the Levant, who would later spread to North and Northeastern Africa, spreading West-Eurasian ancestry and merging with local indigenous Africans. A 2015 study by Dobon et al. identified an ancestral autosomal component that is commonly found among many modern Afroasiatic-speaking populations in Northern Africa. This component formed from a majority of West-Eurasian ancestry and minor African ancestry. The authors proposed that Afroasiatic spread with this component and may be traced back to an Ancestral West-Eurasian source population. However some others argue that the first speakers of Afroasiatic (Proto-Afroasiatic) were of indigenous African origin, and based in North East Africa, probably near the Red sea. They argue that Proto-Afroasiatic migrated during the late Paleolithic into the Levant, merging with local West-Eurasians, resulting in a population which would later give rise to Natufian culture, which is frequently associated with early Afroasiatic-speaking communities, or specifically with early Semitic languages. This Natufian-related population would later spread Afroasiatic languages, resulting in their modern distribution and genetic makeup. A 2018 analysis of autosomal DNA using modern populations as a reference, found that the ancient Natufian samples were largely of local West-Eurasian origin, but harbored 6.8% Omotic-related ancestry (related to modern Sub-Saharan Africans). It is suggested that this Omotic/African component may be associated with the spread of Proto-Afroasiatic and the specific Y-haplogroup sublineage E-M215, also known as "E1b1b", to Western Eurasia. Further evidence for this are the results published in a 2017 paper about previous genetic data of Northern African populations, such as Berbers. Northern African populations have been described as a mosaic of Middle Eastern, European, and Sub-Saharan African-related ancestries.

Austronesian expansion, outgoing from Taiwan and the northern Philippines.
 
Population structure of Qataris in comparison to HGDP world populations. (A) Principal components 1 and 2 of all 55 HGDP populations along with Qataris colored by sampling origin.

Specific East Asian-related ancestry is found among the Malagasy speakers of Madagascar at low to medium frequency. The presence of this East Asian-related ancestry is mostly linked to the Austronesian peoples expansion from Southeast Asia. The peoples of Borneo were identified to resemble the East Asian voyagers, who arrived on Madagascar.

A 2020 study by Chen et al. analyzed 2,504 African samples from all over Africa, and found archaic Neanderthal ancestry, among all tested African samples at low frequency. They also identified an European-related (West-Eurasian) ancestry segment, which seems to largely correspond with the detected Neanderthal ancestry components. Eurasian admixture among Africans was estimated to be between ~0% to up to 30%.

According to the authors:

These data are consistent with the hypothesis that back-migration contributed to the signal of Neanderthal ancestry in Africans. Furthermore, the data indicates that this back-migration came after the split of Europeans and East Asians, from a population related to the European lineage."

A 2021 paper by Hollfelder et al., concluded that West African Yoruba people, which were previously used as "unadmixed reference population" for indigenous Africans, harbor minor levels of Neanderthal ancestry, which can be largely associated with back-migration of an "Ancestral European-like" source population.

Map of Sub-Saharan African and specific North African admixture in European populations

Multiple studies found also evidence for minor geneflow of indigenous African ancestry towards Eurasia, specifically Europe and the Middle East. The analysis of 40 different West-Eurasian populations found African admixture at a frequency of 1% to up to 15%. Modern Northern Africans (Egyptians) were found to carry significant indigenous African ancestry at ~25% on average, with a high variability depending on subgroups and region.

Regional genomic overview

North Africa

Archaic Human DNA

While Denisovan and Neanderthal ancestry in non-Africans outside of Africa are more certain, archaic human ancestry in Africans is less certain and is too early to be established with certainty.

Ancient DNA

Egypt

Amenhotep III, Akhenaten, and Tutankhamen carried the paternal haplogroup R1b. Thuya, Tiye, Tutankhamen's mother, and Tutankhamen carried the maternal haplogroup K.

Ramesses III and "Unknown Man E", possibly Pentawere, carried paternal haplogroup E1b1a.

Khnum-aa, Khnum-Nakht, and Nakht-Ankh carried maternal haplogroup M1a1.

Libya

At Takarkori rockshelter, in Libya, two naturally mummified women, dated to the Middle Pastoral Period (7000 BP), carried basal maternal haplogroup N.

Morocco

The Taforalts of Morocco, who have been radiocarbon dated between 15,100 cal BP and 13,900 cal BP, and were found to be 63.5% Natufian, were also found to be 36.5% Sub-Saharan African (e.g., Hadza), which is drawn out, most of all, by West Africans (e.g., Yoruba, Mende). In addition to having similarity with the remnant of a more basal Sub-Saharan African lineage (e.g., a basal West African lineage shared between Yoruba and Mende peoples), the Sub-Saharan African DNA in the Taforalt people of the Iberomaurusian culture may be best represented by modern West Africans (e.g., Yoruba).

Y-Chromosomal DNA

Mitochondrial DNA

Mitochondrial haplogroups L3, M, and N are found among Sudanese peoples (e.g., Beja, Nilotics, Nuba, Nubians), who have no known interaction (e.g., history of migration/admixture) with Europeans or Asians; rather than having developed in a post-Out-of-Africa migration context, mitochondrial macrohaplogroup L3/M/N and its subsequent development into distinct mitochondrial haplogroups (e.g., Haplogroup L3, Haplogroup M, Haplogroup N) may have occurred in East Africa at a time that considerably predates the Out-of-Africa migration event of 50,000 BP.

Autosomal DNA

Medical DNA

Lactase Persistence

Neolithic agriculturalists, who may have resided in Northeast Africa and the Near East, may have been the source population for lactase persistence variants, including –13910*T, and may have been subsequently supplanted by later migrations of peoples. The Sub-Saharan West African Fulani, the North African Tuareg, and European agriculturalists, who are descendants of these Neolithic agriculturalists, share the lactase persistence variant –13910*T. While shared by Fulani and Tuareg herders, compared to the Tuareg variant, the Fulani variant of –13910*T has undergone a longer period of haplotype differentiation. The Fulani lactase persistence variant –13910*T may have spread, along with cattle pastoralism, between 9686 BP and 7534 BP, possibly around 8500 BP; corroborating this timeframe for the Fulani, by at least 7500 BP, there is evidence of herders engaging in the act of milking in the Central Sahara.

West Africa

Archaic Human DNA

Archaic traits found in human fossils of West Africa (e.g., Iho Eleru fossils, which dates to 13,000 BP) and Central Africa (e.g., Ishango fossils, which dates between 25,000 BP and 20,000 BP) may have developed as a result of admixture between archaic humans and modern humans or may be evidence of late-persisting early modern humans. While Denisovan and Neanderthal ancestry in non-Africans outside of Africa are more certain, archaic human ancestry in Africans is less certain and is too early to be established with certainty.

Ancient DNA

As of 2017, human ancient DNA has not been found in the region of Western Africa. As of 2020, human ancient DNA has not been forthcoming in the region of Western Africa.

In 4000 BP, there may have been a population that traversed from Africa (e.g., West Africa or West-Central Africa), through the Strait of Gibraltar, into the Iberian peninsula, where admixing between Africans and Iberians (e.g., of northern Portugal, of southern Spain) occurred.

In Granada, a Muslim (Moor) of the Cordoba Caliphate, who was of haplogroups E1b1a1 and H1+16189, as well as estimated to date between 900 CE and 1000 CE, and a Morisco, who was of haplogroup L2e1, as well as estimated to date between 1500 CE and 1600 CE, were both found to be of Sub-Saharan West African (i.e., Gambian) and Iberian descent.

Y-Chromosomal DNA

As a result of haplogroup D0, a basal branch of haplogroup DE, being found in three Nigerian men, it may be the case that haplogroup DE, as well as its sublineages D and E, originated in Africa.

As of 19,000 years ago, Africans, bearing haplogroup E1b1a-V38, likely traversed across the Sahara, from east to west. E1b1a1-M2 likely originated in West Africa or Central Africa.

Mitochondrial DNA

Around 18,000 BP, Mende people, along with Gambian peoples, grew in population size.

In 15,000 BP, Niger-Congo speakers may have migrated from the Sahelian region of West Africa, along the Senegal River, and introduced L2a1 into North Africa, resulting in modern Mauritanian peoples and Berbers of Tunisia inheriting it.

Between 11,000 BP and 10,000 BP, Yoruba people and Esan people grew in population size.

Up to 11,000 years ago, Sub-Saharan West Africans, bearing macrohaplogroup L (e.g., L1b1a11, L1b1a6a, L1b1a8, L1b1a9a1, L2a1k, L3d1b1a), may have migrated through North Africa and into Europe, mostly into southern Europe (e.g., Iberia).

Autosomal DNA

Between 2000 BP and 1500 BP, Nilo-Saharan-speakers may have migrated across the Sahel, from East Africa into West Africa, and admixed with Niger-Congo-speaking Berom people.

Medical DNA

Sickle Cell

Amid the Green Sahara, the mutation for sickle cell originated in the Sahara or in the northwest forest region of western Central Africa (e.g., Cameroon) by at least 7,300 years ago, though possibly as early as 22,000 years ago. The ancestral sickle cell haplotype to modern haplotypes (e.g., Cameroon/Central African Republic and Benin/Senegal haplotypes) may have first arose in the ancestors of modern West Africans, bearing haplogroups E1b1a1-L485 and E1b1a1-U175 or their ancestral haplogroup E1b1a1-M4732. West Africans (e.g., Yoruba and Esan of Nigeria), bearing the Benin sickle cell haplotype, may have migrated through the northeastern region of Africa into the western region of Arabia. West Africans (e.g., Mende of Sierra Leone), bearing the Senegal sickle cell haplotype, may have migrated into Mauritania (77% modern rate of occurrence) and Senegal (100%); they may also have migrated across the Sahara, into North Africa, and from North Africa, into Southern Europe, Turkey, and a region near northern Iraq and southern Turkey. Some may have migrated into and introduced the Senegal and Benin sickle cell haplotypes into Basra, Iraq, where both occur equally. West Africans, bearing the Benin sickle cell haplotype, may have migrated into the northern region of Iraq (69.5%), Jordan (80%), Lebanon (73%), Oman (52.1%), and Egypt (80.8%).

Schistosomes

According to Steverding (2020), while not definite: Near the African Great Lakes, schistosomes (e.g., S. mansoni, S. haematobium) underwent evolution. Subsequently, there was an expansion alongside the Nile. From Egypt, the presence of schistosomes may have expanded, via migratory Yoruba people, into Western Africa. Thereafter, schistosomes may have expanded, via migratory Bantu peoples, into the rest of Sub-Saharan Africa (e.g., Southern Africa, Central Africa).

Thalassemia

Through pathways taken by caravans, or via travel amid the Almovarid period, a population (e.g., Sub-Saharan West Africans) may have introduced the –29 (A → G) β-thalassemia mutation (found in notable amounts among African-Americans) into the North African region of Morocco.

Central Africa

Archaic Human DNA

Archaic traits found in human fossils of West Africa (e.g., Iho Eleru fossils, which dates to 13,000 BP) and Central Africa (e.g., Ishango fossils, which dates between 25,000 BP and 20,000 BP) may have developed as a result of admixture between archaic humans and modern humans or may be evidence of late-persisting early modern humans. While Denisovan and Neanderthal ancestry in non-Africans outside of Africa are more certain, archaic human ancestry in Africans is less certain and is too early to be established with certainty.

Ancient DNA

In 4000 BP, there may have been a population that traversed from Africa (e.g., West Africa or West-Central Africa), through the Strait of Gibraltar, into the Iberian peninsula, where admixing between Africans and Iberians (e.g., of northern Portugal, of southern Spain) occurred.

Cameroon

West African hunter-gatherers, in the region of western Central Africa (e.g., Shum Laka, Cameroon), particularly between 8000 BP and 3000 BP, were found to be related to modern Central African hunter-gatherers (e.g., Baka, Bakola, Biaka, Bedzan).

Democratic Republic of Congo

At Kindoki, in the Democratic Republic of Congo, there were three individuals, dated to the protohistoric period (230 BP, 150 BP, 230 BP); one carried haplogroups E1b1a1a1d1a2 (E-CTS99, E-CTS99) and L1c3a1b, another carried haplogroup E (E-M96, E-PF1620), and the last carried haplogroups R1b1 (R-P25 1, R-M415) and L0a1b1a1.

At Ngongo Mbata, in the Democratic Republic of Congo, an individual, dated to the protohistoric period (220 BP), carried haplogroup L1c3a.

At Matangai Turu Northwest, in the Democratic Republic of Congo, an individual, dated to the Iron Age (750 BP), carried an undetermined haplogroup(s).

Y-Chromosomal DNA

Haplogroup R-V88 may have originated in western Central Africa (e.g., Equatorial Guinea), and, in the middle of the Holocene, arrived in North Africa through population migration.

Mitochondrial DNA

Mitochondrial haplogroup L1c is strongly associated with pygmies, especially with Bambenga groups. L1c prevalence was variously reported as: 100% in Ba-Kola, 97% in Aka (Ba-Benzélé), and 77% in Biaka, 100% of the Bedzan (Tikar), 97% and 100% in the Baka people of Gabon and Cameroon, respectively, 97% in Bakoya (97%), and 82% in Ba-Bongo. Mitochondrial haplogroups L2a and L0a are prevalent among the Bambuti.

Autosomal DNA

Genetically, African pygmies have some key difference between them and Bantu peoples.

Medical DNA

Evidence suggests that, when compared to other Sub-Saharan African populations, African pygmy populations display unusually low levels of expression of the genes encoding for human growth hormone and its receptor associated with low serum levels of insulin-like growth factor-1 and short stature.

Eastern Africa

Archaic Human DNA

While Denisovan and Neanderthal ancestry in non-Africans outside of Africa are more certain, archaic human ancestry in Africans is less certain and is too early to be established with certainty.

Ancient DNA

Ethiopia

At Mota, in Ethiopia, an individual, estimated to date to the 5th millennium BP, carried haplogroups E1b1 and L3x2a. The individual of Mota is genetically related to groups residing near the region of Mota, and in particular, are considerably genetically related to the Ari people.

Kenya

At Jawuoyo Rockshelter, in Kisumu County, Kenya, a forager of the Later Stone Age carried haplogroups E1b1b1a1b2/E-V22 and L4b2a2c.

At Ol Kalou, in Nyandarua County, Kenya, a pastoralist of the Pastoral Neolithic carried haplogroups E1b1b1b2b2a1/E-M293 and L3d1d.

At Kokurmatakore, in Marsabit County, Kenya, a pastoralist of the Pastoral Iron Age carried haplogroups E1b1b1/E-M35 and L3a2a.

At White Rock Point, in Homa Bay County, Kenya, there were two foragers of the Later Stone Age; one carried haplogroups BT (xCT), likely B, and L2a4, and another probably carried haplogroup L0a2.

At Nyarindi Rockshelter, in Kenya, there were two individuals, dated to the Later Stone Age (3500 BP); one carried haplogroup L4b2a and another carried haplogroup E (E-M96, E-P162).

At Lukenya Hill, in Kenya, there were two individuals, dated to the Pastoral Neolithic (3500 BP); one carried haplogroups E1b1b1b2b (E-M293, E-CTS10880) and L4b2a2b, and another carried haplogroup L0f1.

At Hyrax Hill, in Kenya, an individual, dated to the Pastoral Neolithic (2300 BP), carried haplogroups E1b1b1b2b (E-M293, E-M293) and L5a1b.

At Molo Cave, in Kenya, there were two individuals, dated to the Pastoral Neolithic (1500 BP); while one had haplogroups that went undetermined, another carried haplogroups E1b1b1b2b (E-M293, E-M293) and L3h1a2a1.

At Kakapel, in Kenya, there were three individuals, one dated to the Later Stone Age (3900 BP) and two dated to the Later Iron Age (300 BP, 900 BP); one carried haplogroups CT (CT-M168, CT-M5695) and L3i1, another carried haplogroup L2a1f, and the last carried haplogroup L2a5.

At Panga ya Saidi, in Kenya, an individual, estimated to date between 496 BP and 322 BP, carried haplogroups E1b1b1b2 and L4b2a2.

Laikipia County

At Kisima Farm/Porcupine Cave, in Laikipia County, Kenya, there were two pastoralists of the Pastoral Neolithic; one carried haplogroups E1b1b1b2b2a1/E-M293 and M1a1, and another carried haplogroup M1a1f.

At Kisima Farm/C4, in Laikipia County, Kenya, a pastoralist of the Pastoral Iron Age, carried haplogroups E2 (xE2b)/E-M75 and L3h1a1.

At Laikipia District Burial, in Laikipia County, Kenya, a pastoralist of the Pastoral Iron Age carried haplogroup L0a1c1.

Nakuru County

At Prettejohn's Gully, in Nakuru County, Kenya, there were two pastoralists of the early pastoral period; one carried haplogroups E2 (xE2b)/E-M75 and K1a, and another carried haplogroup L3f1b.

At Cole's Burial, in Nakuru County, Kenya, a pastoralist of the Pastoral Neolithic carried haplogroups E1b1b1a1a1b1/E-CTS3282 and L3i2.

At Rigo Cave, in Nakuru County, Kenya, there were three pastoralists of the Pastoral Neolithic/Elmenteitan, one carried haplogroups E1b1b1b2b2a1/E-M293 and L3f, another carried haplogroups E1b1b1b2b2/E-V1486, likely E-M293, and probably M1a1b, and the last carried haplogroups E1b1b1b2b2a1/E-M293 and L4b2a2c.

At Naishi Rockshelter, in Nakuru County, Kenya, there two pastoralists of the Pastoral Neolithic; one carried haplogroups E1b1b1b2b/E-V1515, likely E-M293, and L3x1a, and another carried haplogroups A1b (xA1b1b2a)/A-P108 and L0a2d.

At Keringet Cave, in Nakuru County, Kenya, a pastoralist of the Pastoral Neolithic carried haplogroups A1b1b2/A-L427 and L4b2a1, and another pastoralist of the Pastoral Neolithic/Elmenteitan carried haplogroup K1a.

At Naivasha Burial Site, in Nakuru County, Kenya, there were five pastoralists of the Pastoral Neolithic; one carried haplogroup L4b2a2b, another carried haplogroups xBT, likely A, and M1a1b, another carried haplogroups E1b1b1b2b2a1/E-M293 and L3h1a1, another carried haplogroups A1b1b2b/A-M13 and L4a1, and the last carried haplogroups E1b1b1b2b2a1/E-M293 and L3x1a.

At Njoro River Cave II, in Nakuru County, Kenya, a pastoralist of the Pastoral Neolithic carried haplogroup L3h1a2a1.

At Egerton Cave, in Nakuru County, Kenya, a pastoralist of the Pastoral Neolithic/Elmenteitan carried haplogroup L0a1d.

At Ilkek Mounds, in Nakuru County, Kenya, a pastoralist of the Pastoral Iron Age carried haplogroups E2 (xE2b)/E-M75 and L0f2a.

At Deloraine Farm, in Nakuru County, Kenya, an iron metallurgist of the Iron Age carried haplogroups E1b1a1a1a1a/E-M58 and L5b1.

Narok County

At Kasiole 2, in Narok County, Kenya, a pastoralist of the Pastoral Iron Age carried haplogroups E1b1b1b2b/E-V1515, likely E-M293, and L3h1a2a1.

At Emurua Ole Polos, in Narok County, Kenya, a pastoralist of the Pastoral Iron Age carried haplogroups E1b1b1b2b2a1/E-M293 and L3h1a2a1.

Tanzania

At Luxmanda, Tanzania, an individual, estimated to date between 3141 BP and 2890 BP, carried haplogroup L2a1.

At Kuumbi Cave, in Zanzibar, Tanzania, an individual, estimated to date between 1370 BP and 1303 BP, carried haplogroup L4b2a2c.

Karatu District

At Gishimangeda Cave, in Karatu District, Tanzania, there were eleven pastoralists of the Pastoral Neolithic; one carried haplogroups E1b1b1a1b2/E-V22 and HV1b1, another carried haplogroup L0a, another carried haplogroup L3x1, another carried haplogroup L4b2a2b, another carried haplogroups E1b1b1b2b2a1/E-M293 and L3i2, another carried haplogroup L3h1a2a1, another carried haplogroups E1b1b1b2b2/E-V1486, likely E-M293 and L0f2a1, and another carried haplogroups E1b1b1b2b2/E-V1486, likely E-M293, and T2+150; while most of the haplogroups among three pastoralists went undetermined, one was determined to carry haplogroup BT, likely B.

Pemba Island

At Makangale Cave, on Pemba Island, Tanzania, an individual, estimated to date between 1421 BP and 1307 BP, carried haplogroup L0a.

At Makangale Cave, on Pemba Island, Tanzania, an individual, estimated to date between 639 BP and 544 BP, carried haplogroup L2a1a2.

Uganda

At Munsa, in Uganda, an individual, dated to the Later Iron Age (500 BP), carried haplogroup L3b1a1.

Y-Chromosomal DNA

As of 19,000 years ago, Africans, bearing haplogroup E1b1a-V38, likely traversed across the Sahara, from east to west.

Before the slave trade period, East Africans, who carried haplogroup E1b1a-M2, expanded into Arabia, resulting in various rates of inheritance throughout Arabia (e.g., 2.8% Qatar, 3.2% Yemen, 5.5% United Arab Emirates, 7.4% Oman).

Mitochondrial DNA

Autosomal DNA

Across all areas of Madagascar, the average ancestry for the Malagasy people was found to be 4% West Eurasian, 37% Austronesian, and 59% Bantu.

Medical DNA

Southern Africa

Archaic Human DNA

While Denisovan and Neanderthal ancestry in non-Africans outside of Africa are more certain, archaic human ancestry in Africans is less certain and is too early to be established with certainty.

Ancient DNA

Three Later Stone Age hunter-gatherers carried ancient DNA similar to Khoisan-speaking hunter-gatherers. Prior to the Bantu migration into the region, as evidenced by ancient DNA from Botswana, East African herders migrated into Southern Africa. Out of four Iron Age Bantu agriculturalists of West African origin, two earlier agriculturalists carried ancient DNA similar to Tsonga and Venda peoples and the two later agriculturalists carried ancient DNA similar to Nguni people; this indicates that there were various movements of peoples in the overall Bantu migration, which resulted in increased interaction and admixing between Bantu-speaking peoples and Khoisan-speaking peoples.

Botswana

At Nqoma, in Botswana, an individual, dated to the Early Iron Age (900 BP), carried haplogroup L2a1f.

At Taukome, in Botswana, an individual, dated to the Early Iron Age (1100 BP), carried haplogroups E1b1a1 (E-M2, E-Z1123) and L0d3b1.

At Xaro, in Botswana, there were two individuals, dated to the Early Iron Age (1400 BP); one carried haplogroups E1b1a1a1c1a and L3e1a2, and another carried haplogroups E1b1b1b2b (E-M293, E-CTS10880) and L0k1a2.

Malawi
Fingira

At Fingira, in Malawi, an individual, estimated to date between 6175 BP and 5913 BP, carried haplogroups BT and L0d1b2b.

At Fingira, in Malawi, an individual, estimated to date between 6177 BP and 5923 BP, carried haplogroups BT and L0d1c.

At Fingira, in Malawi, an individual, estimated to date between 2676 BP and 2330 BP, carried haplogroup L0f.

Chencherere

At Chencherere, in Malawi, an individual, estimated to date between 5400 BP and 4800 BP, carried haplogroup L0k2.

At Chencherere, in Malawi, an individual, estimated to date between 5293 BP and 4979 BP, carried haplogroup L0k1.

Hora

At Hora, in Malawi, an individual, estimated to date between 10,000 BP and 5000 BP, carried haplogroups BT and L0k2.

At Hora, in Malawi, an individual, estimated to date between 8173 BP and 7957 BP, carried haplogroup L0a2.

South Africa

At Doonside, in South Africa, an individual, estimated to date between 2296 BP and 1910 BP, carried haplogroup L0d2.

At Champagne Castle, in South Africa, an individual, estimated to date between 448 BP and 282 BP, carried haplogroup L0d2a1a.

At Eland Cave, in South Africa, an individual, estimated to date between 533 BP and 453 BP, carried haplogroup L3e3b1.

At Mfongosi, in South Africa, an individual, estimated to date between 448 BP and 308 BP, carried haplogroup L3e1b2.

At Newcastle, in South Africa, an individual, estimated to date between 508 BP and 327 BP, carried haplogroup L3e2b1a2.

At St. Helena, in South Africa, an individual, estimated to date between 2241 BP and 1965 BP, carried haplogroups A1b1b2a and L0d2c1.

At Faraoskop Rock Shelter, in South Africa, an individual, estimated to date between 2017 BP and 1748 BP, carried haplogroups A1b1b2a and L0d1b2b1b.

At Kasteelberg, in South Africa, an individual, estimated to date between 1282 BP and 1069 BP, carried haplogroup L0d1a1a.

At Vaalkrans Shelter, in South Africa, an individual, estimated to date to 200 BP, is predominantly related to Khoisan speakers, partly related (15% - 32%) to East Africans, and carried haplogroups L0d3b1.

Ballito Bay

At Ballito Bay, South Africa, an individual, estimated to date between 1986 BP and 1831 BP, carried haplogroups A1b1b2 and L0d2c1.

At Ballito Bay, South Africa, an individual, estimated to date between 2149 BP and 1932 BP, carried haplogroups A1b1b2 and L0d2a1.

Y-Chromosomal DNA

Distribution of Y-Chromosome Haplogroup A in Africa.
 
Distribution of haplogroup B (M60) of the human Y chromosome in native populations.

Various Y chromosome studies show that the San carry some of the most divergent (oldest) human Y-chromosome haplogroups. These haplogroups are specific sub-groups of haplogroups A and B, the two earliest branches on the human Y-chromosome tree.

Mitochondrial DNA

Mitochondrial DNA studies also provide evidence that the San carry high frequencies of the earliest haplogroup branches in the human mitochondrial DNA tree. This DNA is inherited only from one's mother. The most divergent (oldest) mitochondrial haplogroup, L0d, has been identified at its highest frequencies in the southern African San groups.

Autosomal DNA

In a study published in March 2011, Brenna Henn and colleagues found that the ǂKhomani San, as well as the Sandawe and Hadza peoples of Tanzania, were the most genetically diverse of any living humans studied. This high degree of genetic diversity hints at the origin of anatomically modern humans.

Medical DNA

Among the ancient DNA from three hunter-gatherers sharing genetic similarity with San people and four Iron Age agriculturalists, their SNPs indicated that they bore variants for resistance against sleeping sickness and Plasmodium vivax. In particular, two out of the four Iron Age agriculturalists bore variants for resistance against sleeping sickness and three out of the four Iron Age agriculturalists bore Duffy negative variants for resistance against malaria. In contrast to the Iron Age agriculturalists, from among the San-related hunter-gatherers, a six-year-old boy may have died from schistosomiasis. In Botswana, a man, who dates to 1400 BP, may have also carried the Duffy negative variant for resistance against malaria.

Recent African origin of modern humans

Between 500,000 BP and 300,000 BP, anatomically modern humans may have emerged in Africa. As Africans (e.g., Y-Chromosomal Adam, Mitochondrial Eve) have migrated from their places of origin in Africa to other locations in Africa, and as the time of divergence for East African, Central African, and West African lineages are similar to the time of divergence for the Southern African lineage, there is insufficient evidence to identify a specific region for the origin of humans in Africa. In 100,000 BP, anatomically modern humans migrated from Africa into Eurasia. Subsequently, tens of thousands of years after, the ancestors of all present-day Eurasians migrated from Africa into Eurasia and eventually became admixed with Denisovans and Neanderthals.

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