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Thursday, March 10, 2022

Fish intelligence

From Wikipedia, the free encyclopedia
 
The elephantnose fish has the highest brain-to-body oxygen consumption ratio of all known vertebrates
 
The bony-eared assfish has the smallest brain-to-body weight ratio of all known vertebrates

Fish intelligence is "...the resultant of the process of acquiring, storing in memory, retrieving, combining, comparing, and using in new contexts information and conceptual skills" as it applies to fish.

According to Culum Brown from Macquarie University, "Fish are more intelligent than they appear. In many areas, such as memory, their cognitive powers match or exceed those of ‘higher’ vertebrates including non-human primates."

Fish hold records for the relative brain weights of vertebrates. Most vertebrate species have similar brain-to-body mass ratios. The deep sea bathypelagic bony-eared assfish, has the smallest ratio of all known vertebrates. At the other extreme, the electrogenic elephantnose fish, an African freshwater fish, has one of the largest brain-to-body weight ratios of all known vertebrates (slightly higher than humans) and the highest brain-to-body oxygen consumption ratio of all known vertebrates (three times that for humans).

Brain

Cross-section of the brain of a porbeagle shark, with the cerebellum highlighted
 
The brain of a cod

Fish typically have quite small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, most notably mormyrids and sharks, which have brains about as massive relative to body weight as birds and marsupials.

The cerebellum of cartilaginous and bony fishes is large and complex. In at least one important respect, it differs in internal structure from the mammalian cerebellum: The fish cerebellum does not contain discrete deep cerebellar nuclei. Instead, the primary targets of Purkinje cells are a distinct type of cell distributed across the cerebellar cortex, a type not seen in mammals. The circuits in the cerebellum are similar across all classes of vertebrates, including fish, reptiles, birds, and mammals. There is also an analogous brain structure in cephalopods with well-developed brains, such as octopuses. This has been taken as evidence that the cerebellum performs functions important to all animal species with a brain.

In mormyrid fish (a family of weakly electrosensitive freshwater fish), the cerebellum is considerably larger than the rest of the brain put together. The largest part of it is a special structure called the valvula, which has an unusually regular architecture and receives much of its input from the electrosensory system.

Memory

Individual carp captured by anglers have been shown to become less catchable thereafter. This suggests that fish use their memory of negative experiences to associate capture with stress and therefore become less easy to catch. This type of associative learning has also been shown in paradise fish (Macropodus opercularis) which avoid places where they have experienced a single attack by a predator and continue to avoid for many months.

Red Sea clownfish can recognize their mate after 30 days separation.

A number of studies have shown that fish can retain information for months or years. Anecdotally, channel catfish (Ictalurus punctatus) can remember the human voice call announcing food five years after last hearing that call. Goldfish remember the colour of a tube dispensing food one year after the last tube presentation. Sockeye salmon still react to a light signal that precedes food arrival up to eight months since the last reinforcement. Some common rudd and European chub could remember the person who trained them to feed from the hand, even after a 6-month break. Crimson-spotted rainbowfish can learn how to escape from a trawl by swimming through a small hole in the center and they remember this technique 11 months later. Rainbow trout can be trained to press a bar to get food, and they remember this three months after last seeing the bar. Red Sea clownfish can recognize their mate 30 days after it was experimentally removed from the home anemone.

Several fish species are capable of learning complex spatial relationships and forming cognitive maps. They can orient themselves using multiple landmarks or symbols and they are able to integrate experiences which enable them to generate appropriate avoidance responses.

Tool use

Video of an archerfish shooting at prey

Tool use is sometimes considered as an indication of intelligence in animals. There are few examples of tool use in fishes, perhaps because they have only their mouth in which to hold objects.

Several species of wrasse hold bivalves (scallops and clams) or sea urchins in their mouth and smash them against the surface of a rock (an "anvil") to break them up. This behaviour in an orange-dotted tuskfish (Choerodon anchorago) has been filmed; the fish fans sand to unearth the bivalve, takes it into its mouth, swims several metres to a rock which it uses as an anvil by smashing the mollusc apart with sideward thrashes of the head.

Archerfish (family Toxotidae) squirt jets of water at insects on plants above the surface to knock them into the water; they can adjust the size of the squirts to the size of the insect prey and learn to shoot at moving targets.

Whitetail damselfish clean the rock face where they intend to lay eggs by sucking up and blowing sand grains onto the surface. Triggerfish blow water at sea urchins to turn them over, thereby exposing their more vulnerable underside. River stingrays create water currents with their fins to suck food out of a PVC pipe. Banded acaras (Bujurquina vittata) lay their eggs on a loose leaf and carry the leaf away when a predator approaches.

In one laboratory study, Atlantic cod (Gadus morhua) given access to an operant feeding machine learned to pull a string to get food. The researchers had also tagged the fish by threading a bead in front of their dorsal fin. Some fish snagged the string with their bead, resulting in food delivery. These fish eventually learned to swim in a particular way to repeatedly make the bead snag the string and get food. Because the fish used an object external to their body in a goal-oriented way, this satisfies some definitions of tool use.

Construction

As for tool use, construction behaviour may be mostly innate. Yet it can be sophisticated, and the fact that fish can make judicious repairs to their creation suggests intelligence. Construction methods in fishes can be divided into three categories: excavations, pile-ups, and gluing.

Excavations may be simple depressions dug up in the substrate, such as the nests of bowfin, smallmouth bass, and Pacific salmon, but it can also consist of fairly large burrows used for shelter and for nesting. Burrowing species include the mudskippers, the red band-fish Cepola rubescens (burrows up to 1 m deep, often with a side branch), the yellowhead jawfish Opistognathus aurifrons (chambers up to 22 cm deep, lined with coral fragments to solidify it), the convict blenny Pholidichthys leucotaenia whose burrow is a maze of tunnels and chambers thought to be as much as 6 m long, and the Nicaragua cichlid, Hypsophrys nicaraguensis, who drills a tunnel by spinning inside of it. In the case of the mudskippers, the burrows are shaped like a J and can be as much as 2 m deep. Two species, the giant mudskipper Periophthalmodon schlosseri and the walking goby Scartelaos histophorus, build a special chamber at the bottom of their burrows into which they carry mouthfuls of air. Once released the air accumulates at the top of the chamber and forms a reserve from which the fish can breathe – like all amphibious fishes, mudskippers are good air breathers. If researchers experimentally extract air from the special chambers, the fish diligently replenish it. The significance of this behaviour stems from the facts that at high tide, when water covers the mudflats, the fish stay in their burrow to avoid predators, and water inside the confined burrow is often poorly oxygenated. At such times these air-breathing fishes can tap into the air reserve of their special chambers.

Mounds are easy to build, but can be quite extensive. In North American streams, the male cutlip minnow Exoglossum maxillingua, 90–115 mm (3.5–4.5 in) long, assembles mounds that are 75–150 mm (3.0–5.9 in) high, 30–45 cm (12–18 in) in diameter, made up of more than 300 pebbles 13–19 mm in diameter (a quarter to half an inch). The fish carry these pebbles one by one in their mouths, sometimes stealing some from the mounds of other males. The females deposit their eggs on the upstream slope of the mounds, and the males cover these eggs with more pebbles. Males of the hornyhead chub Nocomis biguttatus, 90 mm (3.5 in) long, and of the river chub Nocomis micropogon, 100 mm (3.9 in) long, also build mounds during the reproductive season. They start by clearing a slight depression in the substrate, which they overfill with up to 10,000 pebbles until the mounds are 60–90 cm (2.0–3.0 ft) long (in the direction of the water current), 30–90 cm (0.98–2.95 ft) wide, and 5–15 cm (2.0–5.9 in) high. Females lay their eggs among those pebbles. The stone accumulation is free of sand and it exposes the eggs to a good water current that supplies oxygen. Males of many mouthbrooding cichlid species in Lake Malawi and Lake Tanganyika build sand cones that are flattened or crater-shaped at the top. Some of these mounds can be 3 m in diameter and 40 cm high. The mounds serve to impress females or to allow species recognition during courtship.

Male pufferfish, Torquigener sp., also build sand mounds to attract females. The mounds, up to 2 m in diameter, are intricate with radiating ridges and valleys.

Several species build up mounds of coral pieces either to protect the entrance to their burrows, as in tilefishes and gobies of the genus Valenciennea, or to protect the patch of sand in which they will bury themselves for the night, as in the Jordan's tuskfish Choerodon jordani and the rockmover wrasse Novaculichthys taeniourus.

Male sticklebacks are well known for their habit of building an enclosed nest made of pieces of vegetation glued together with secretions from their kidneys. Some of them adorn the entrance of the nest with unusually colored algae or even shiny tinfoil experimentally introduced in the environment.

Foam nests, made up of air bubbles glued together with mucus from the mouth, are also well known in gouramis and armoured catfish.

Social intelligence

Fish can remember the attributes of other individuals, such as their competitive ability or past behavior, and modify their own behavior accordingly. For example, they can remember the identity of individuals to whom they have lost in a fight, and avoid these individuals in the future; or they can recognize territorial neighbors and show less aggression towards them as compared to strangers. They can recognize individuals in whose company they obtained less food in the past and preferentially associate with new partners in the future.

Fish can seem mindful of which individuals have watched them in the past. In an experiment with Siamese fighting fish, two males were made to fight each other while being watched by a female, whom the males could also see. The winner and the loser of the fight were then, separately, given a choice between spending time next to the watching female or to a new female. The winner courted both females equally, but the loser spent more time next to the new female, avoiding the watcher female. In this species, females prefer males they have seen win a fight over males they have seen losing, and it therefore makes sense for a male to prefer a female that has never seen him as opposed to a female that has seen him lose.

Social interactions also provides the context for a test of transitive inference, that is figuring out that if A > B and B > C, then A > C. In a study with the cichlid Astatotilapia burtoni, observer fish could watch aggressive interactions between pairs of other individuals. They witnessed individual A beat individual B, then individual B beat individual C, then C beat D, and D beat E. The observer fish were then given a choice of associating with either B or D (both of which they had seen win once and lose once). All eight observer fish tested spent more time next to D. Fish in this species prefer to associate with more subordinate individuals, so the preference for D showed that the observers had worked out that B was superior to C, and C to D, and therefore D was subordinate to B.

Deception

There are several examples of fish being deceptive, suggesting to some researchers that they may possess a theory of mind. However, most of the observations of deception can be understood as instinctive patterns of behavior that are triggered by specific environmental events, and they do not require a fish to understand of the point of view of other individuals.

Distraction display

Adult male bowfins distract potential predators away from their fry by thrashing as if injured.

In the threespine stickleback (Gasterosteus aculeatus), males sometimes see their nest full of eggs fall prey to groups of marauding females; some males, when they see a group of females approaching, swim away from their nest and start poking their snout into the substrate, as would a female raiding a nest. This distraction display commonly fools the females into behaving as if a nest has been discovered there and they rush to that site, leaving the male's real nest alone. Bowfin (Amia calva) males caring for their free-swimming fry exhibit a related distraction display when a potential fry-predator approaches; they move away and thrash about as if injured, drawing the predator's attention toward himself.

False courtship behaviour

In the Malili Lakes of Sulawesi, Indonesia, one species of sailfin silverside (Telmatherina sarasinorum) is an egg predator. They often follow courting pairs of the closely related species T. antoniae. When those pairs lay eggs, T. sarasinorum darts in and eats the eggs. On four different occasions in the field (out of 136 observation bouts in total), a male T. sarasinorum who was following a pair of courting T. antoniae eventually chased off the male T. antoniae and took his place, courting the heterospecific female. That female released eggs, at which point the male darted to the eggs and ate them.

Death feigning

Death feigning as a way to attract prey is another form of deception. In Lake Malawi, the predatory cichlid Nimbochromis livingstonii have been seen first remaining stationary with their abdomen on or near sand and that then dropping onto their sides. In a variant behaviour, some N. livingstonii fell through the water column and landed onto their side. The fish then remained immobile for several minutes. Their colour pattern was blotchy and suggested a rotting carcass. Small inquisitive cichlids of other species often came near and they were suddenly attacked by the predator. About a third of the death-feigning performances led to an attack, and about one-sixth of the attacks were successful. Another African cichlid, Lamprologus lemairii, from Lake Tanganyika, has been reported to do the same thing. A South American cichlid, the yellowjacket cichlid Parachromis friedrichsthalii, also uses death feigning. They turn over onto their sides at the bottom of the sinkholes they inhabit and remain immobile for as long as 15 minutes, during which they attack the small mollies that come too close to them. The comb grouper Mycteroperca acutirostris may also be an actor, though in this case the behaviour should be called dying or illness feigning, rather than death feigning, because while lying on its side the fish occasionally undulates its body. In 1999, off the coast of southeastern Brazil, one juvenile comb grouper was observed using this tactic to catch five small prey in 15 minutes.

Cooperation

Cooperative foraging reflects some mental flexibility and planning, and could therefore be interpreted as intelligence. There are a few examples in fishes.

Yellowtail amberjack can form packs of 7-15 individuals that maneuver in U-shaped formations to cut away the tail end of prey shoals (jack mackerels or Cortez grunts) and herd the downsized shoal next to seawalls where they proceed to capture the prey.

In the coral reefs of the Red Sea, roving coralgrouper that have spotted a small prey fish hiding in a crevice sometimes visit the sleeping hole of a giant moray and shake their head at the moray, and this seems to be an invitation to group hunting as the moray often swims away with the grouper, is led to the crevice where the prey hides, and proceeds to probe that crevice (which is too small to let the grouper in), either catching the prey by itself or flushing it into the open where the grouper grabs it. The closely related coral trout also enrolls the help of moray eels in this way, and they only do so when the prey they seek is hidden in crevices, where only the eel can flush them. They also quickly learn to invite preferentially those individual eels that collaborate most often.

Similarly, zebra lionfish that detect the presence of small prey fishes flare up their fins as an invitation to other zebra lionfish, or even to another species of lionfish (Pterois antennata), to join them in better cornering the prey and taking turns at striking the prey so that every individual hunter ends up with similar capture rates.

Numeracy

Mosquitofish (Gambusia holbrooki) can distinguish between doors marked with either two or three geometric symbols, only one of which allows the fish to rejoin its shoalmates. This can be achieved when the two symbols have the same total surface area, density and brightness as the three symbols. Further studies show this discrimination extends to 4 vs 8, 15 vs 30, 100 vs 200, 7 vs 14, and 8 vs 12 symbols, again controlling for non-numerical factors.

Many studies have shown that when given a choice, shoaling fish prefer to join the larger of two shoals. It has been argued that several aspects of such choice reflect an ability by fish to distinguish between numerical quantities.

Social learning

Fish can learn how to perform a behavior simply by watching other individuals in action. This is variously called observational learning, cultural transmission, or social learning. For example, fish can learn a particular route after following an experienced leader a few times. One study trained guppies to swim through a hole marked in red while ignoring another one marked in green in order to get food on the other side of a partition; when these experienced fish (“demonstrators”) were joined by a naive one (an “observer”), the observer followed the demonstrators through the red hole, and kept the habit once the demonstrators were removed, even when the green hole now allowed food access. In the wild, juvenile French grunt follow traditional migration routes, up to 1 km long, between their daytime resting sites and their nighttime foraging areas on coral reefs; if groups of 10-20 individuals are marked and then transplanted to new populations, they follow the residents along what is for them – the transplants – a new migration route, and if the residents are then removed two days later, the transplanted grunts continue to use the new route, as well as the resting and foraging sites at both ends.

Through cultural transmission, fishes could also learn where good food spots are. Ninespine stickleback, when given a choice between two food patches they have watched for a while, prefer the patch over which more fish have been seen foraging, or over which fish were seen feeding more intensively. Similarly, in a field experiment where Trinidadian guppies were given a choice between two distinctly marked feeders in their home rivers, the subjects chose the feeder where other guppies were already present, and in subsequent tests when both feeders were deserted, the subjects remembered the previously popular feeder and chose it.

Through social learning, fishes might learn not only where to get food, but also what to get and how to get it. Hatchery-raised salmon can be taught to quickly accept novel, live prey items similar to those they will encounter once they will be released in the wild, simply by watching an experienced salmon take such prey. The same is true of young perch. In the laboratory, juvenile European seabass can learn to push a lever in order to obtain food just by watching experienced individuals use the lever.

Fishes can also learn from others the identity of predatory species. Fathead minnows, for example, can learn the smell of a predatory pike just by being simultaneously exposed to that smell and the sight of experienced minnows reacting with fear, and brook stickleback can learn the visual identity of a predator by watching the fright reaction of experienced fathead minnows. Fish can also learn to recognize the odor of dangerous sites when they are simultaneously exposed to it and to other fish that suddenly show a fright reaction. Hatchery-raised salmon can learn the smell of a predator by being simultaneously exposed to it and to the alarm substance released by injured salmon.

Latent learning

Latent learning is a form of learning that is not immediately expressed in an overt response; it occurs without any obvious reinforcement of the behaviour or associations that are learned. One example in fish comes from research with male three spot gouramis (Trichopodus trichopterus). This species quickly form dominance hierarchies. To appease dominants, subordinates adopt a typical body posture angled at 15-60ยบ to the horizontal, all fins folded and pale body colors. Individuals trained to associate a light-stimulus with the imminent arrival of food exhibit this associative learning by approaching the surface where the food is normally dropped immediately the light-stimulus is presented. However, if a subordinate is placed in a tank with a dominant individual and the light-stimulus is presented, the subordinate immediately assumes the submissive posture rather than approaching the surface. The subordinate has predicted that going to the surface to get food would place it in competition with the dominant, and to avoid potential aggression, it immediately attempts to appease the dominant.

Cleaner fish

Bluestreak cleaner wrasse (bottom) with a client fish

The bluestreak cleaner wrasse (Labroides dimidiatus) performs a service for “client” fishes (belonging to other species) by removing and eating their ectoparasites. Clients can invite a cleaning session by adopting a typical posture or simply by remaining immobile near a wrasse's cleaning station. They can even form queues while doing so. But cleaning sessions do not always end up well, because wrasses (or wrasse-mimicking parasitic sabre-toothed blennies) may cheat and eat the nutritious body mucus of their clients, rather than just the ectoparasites, something that makes the client jolt and sometimes flee. This system has been the subject of extensive observations which have suggested cognitive abilities on the part of the cleaner wrasses and their clients. For example, clients refrain from soliciting a cleaning session if they have witnessed the cleaning session of the previous client ending badly. Cleaners give the impression of trying to maintain a good reputation, because they cheat less when they see a big audience (a long queue of clients) watching. Cleaners sometimes work as male-female teams, and when the smaller female cheats and bites the client, the larger male chases her off, as if to punish her for having tarnished their reputation.

Play

Play behaviour is often considered a correlate of intelligence. One possible example in fish is provided by the electrolocating Peters' elephantnose fish (mentioned above as having one of the largest brain-to-body weight ratios of all known vertebrates). One captive individual was observed carrying a small ball of aluminum foil (a good conductor of electricity) to the outflow tube of the aquarium filter, letting the current push the ball away before chasing after it and repeating the behaviour. Captive white-spotted cichlids have also been seen hitting a floating thermometer hundreds of times to make it wobble and bob.

Food stocking

Food stocking can be viewed potentially as an animal planning for the future. One example of short-term stocking involves climbing perch (Anabas testudineus). Individuals were kept singly in aquaria and fed with pellets dropped at the surface. When the pellets were dropped one after the other at 1-s intervals, the fish took them as they reached the surface and stocked them inside the mouth. On average, the fish placed 7 pellets in their mouth before moving away to consume them. When starved for 24-h before the feeding test, they doubled the number of pellets stocked (14 on average); the underside of their heads bulged under the load. The behaviour may be an indication that competition for food is normally severe in this species and that any adaptation to secure food would be beneficial.

Ancient Egyptian race controversy

From Wikipedia, the free encyclopedia
https://en.wikipedia.org/wiki/Ancient_Egyptian_race_controversy

The question of the race of ancient Egyptians was raised historically as a product of the early racial concepts of the 18th and 19th centuries, and was linked to models of racial hierarchy primarily based on craniometry and anthropometry. A variety of views circulated about the racial identity of the Egyptians and the source of their culture. Some scholars argued that ancient Egyptian culture was influenced by other Afroasiatic-speaking populations in North Africa or the Middle East, while others pointed to influences from various Nubian groups or populations in Europe. In more recent times some writers continued to challenge the mainstream view, some focusing on questioning the race of specific notable individuals such as the king represented in the Great Sphinx of Giza, native Egyptian pharaoh Tutankhamun, Egyptian Queen Tiye, and Greek Ptolemaic queen Cleopatra VII.

Mainstream scholars reject the notion that Egypt was a white or black civilization; they maintain that, despite the phenotypic diversity of Ancient and present-day Egyptians, applying modern notions of black or white races to ancient Egypt is anachronistic. In addition, scholars reject the notion, implicit in the notion of a black or white Egypt hypothesis, that Ancient Egypt was racially homogeneous; instead, skin color varied between the peoples of Lower Egypt, Upper Egypt, and Nubia, who in various eras rose to power in Ancient Egypt. Within Egyptian history, despite multiple foreign invasions, the demographics were not shifted substantially by large migrations.

Origins

In the 18th century, Constantin Franรงois de Chassebล“uf, comte de Volney, wrote about his thoughts on the contentions regarding the race of the ancient Egyptians. In one translation, he noted that "the Copts are the proper representatives of the Ancient Egyptians" due to their "jaundiced and fumed skin, which is neither Greek, Negro nor Arab, their full faces, their puffy eyes, their crushed noses, and their thick lips...the ancient Egyptians were true negroes of the same type as all native born Africans". Volney also said that the Sphinx gave him the key to the riddle as to why all the Egyptians he saw across the country "have a bloated face, puffed-up eyes, flat nose, thick lips – in a word, the true face of the mulatto." He wrote he was tempted to attribute it to the climate, but upon visiting the Sphinx, its appearance gave him the answer; "seeing that head, typically negro in all its features", Volney saw it as the "true solution to the enigma (of how the modern Egyptians came to have their 'mulatto' appearance)". He goes on to postulate, "the Copts were "true negroes" of the same stock as all the autochthonous peoples of Africa" and they "after some centuries of mixing..., must have lost the full blackness of its original color."

Another early example of the controversy is an article published in The New-England Magazine of October 1833, where the authors dispute a claim that "Herodotus was given as authority for their being negroes." They point out with reference to tomb paintings: "It may be observed that the complexion of the men is invariably red, that of the women yellow; but neither of them can be said to have anything in their physiognomy at all resembling the Negro countenance."

A few years later, in 1839, Jean-Franรงois Champollion stated in his work Egypte Ancienne that the Egyptians and Nubians are represented in the same manner in tomb paintings and reliefs, further suggesting that: "In the Copts of Egypt, we do not find any of the characteristic features of the ancient Egyptian population. The Copts are the result of crossbreeding with all the nations that successfully dominated Egypt. It is wrong to seek in them the principal features of the old race." This memoir was made in the context of the first tribes that would have inhabited Egypt, his opinion was noted after his return from Nubia. In 1839, Champollion's and Volney's claims were disputed by Jacques Joseph Champollion-Figeac, who blamed a misunderstanding of the ancients for spreading a false impression of a "Negro" Egypt, stating "the two physical traits of black skin and wooly hair are not enough to stamp a race as negro" and "the opinion that the ancient population of Egypt belonged to the Negro African race, is an error long accepted as the truth. ... Volney's conclusion as to the Negro origin of the ancient Egyptian civilization is evidently forced and inadmissible."

Foster summarized the early 19th century "controversy over the ethnicity of the ancient Egyptians" as a debate of conflicting theories regarding the Hamites. "In ancient times, the Hamites, who developed the civilization of Egypt, were considered Black." Foster describes the 6th century CE curse of Ham theory, which began "in the Babylonian Talmud, a collection of oral traditions of the Jews, that the sons of Ham are cursed by being black." Foster said "throughout the Middle Ages and to the end of the eighteenth century, the Negro was seen by Europeans as a descendant of Ham." In the early 19th century, "after Napolean's expedition to Egypt, the Hamites began to be viewed as having been Caucasians." However, "Napolean's scientists concluded that the Egyptians were Negroid." Napoleon's colleagues referenced prior "well-known books" by Constantin Franรงois de Chassebล“uf, comte de Volney and Vivant Denon that described Ancient Egyptians as "negroid". Finally, Foster concludes, "it was at this point that Egypt became the focus of much scientific and lay interest, the result of which was the appearance of many publications whose sole purpose was to prove that the Egyptians were not Black, and therefore capable of developing such a high civilization."

The debate over the race of the ancient Egyptians intensified during the 19th century movement to abolish slavery in the United States, as arguments relating to the justifications for slavery increasingly asserted the historical, mental and physical inferiority of black people. For example, in 1851, John Campbell directly challenged the claims by Champollion and others regarding the evidence for a black Egypt, asserting "There is one great difficulty, and to my mind an insurmountable one, which is that the advocates of the negro civilization of Egypt do not attempt to account for, how this civilization was lost.... Egypt progressed, and why, because it was Caucasian." The arguments regarding the race of the Egyptians became more explicitly tied to the debate over slavery in the United States, as tensions escalated towards the American Civil War. In 1854, Josiah C. Nott with George Glidden set out to prove: "that the Caucasian or white, and the Negro races were distinct at a very remote date, and that the Egyptians were Caucasians." Samuel George Morton, a physician and professor of anatomy, concluded that "Negroes were numerous in Egypt, but their social position in ancient times was the same that it now is [in the United States], that of servants and slaves." In the early 20th century, Flinders Petrie, a professor of Egyptology at the University of London, in turn spoke of "a black queen", Ahmose-Nefertari, who was the "divine ancestress of the XVIIIth dynasty". He described her physically as "the black queen Aohmes Nefertari had an aquiline nose, long and thin, and was of a type not in the least prognathous".

Position of modern scholarship

Modern scholars who have studied ancient Egyptian culture and population history have responded to the controversy over the race of the ancient Egyptians in various ways.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, the "Black Hypothesis" met with "profound" disagreement by scholars. Similarly, none of the participants voiced support for an earlier postulation that Egyptians were "white with a dark, even black, pigmentation." The arguments for all sides are recorded in the UNESCO publication General History of Africa, with the "Origin of the Egyptians" chapter being written by Cheikh Anta Diop, a proponent of the "Black Hypothesis". At the 1974 UNESCO conference, most participants concluded that the ancient Egyptian population was indigenous to the Nile Valley, and was made up of people from north and south of the Sahara who had a range of skin colors.

Since the late 20th century, as the science of human population genetics has advanced, most biological anthropologists have come to reject the notion of race as having any validity in the study of human biology.

Frank M. Snowden wrote: "Egyptians, Greeks and Romans attached no special stigma to the colour of the skin and developed no hierarchical notions of race whereby highest and lowest positions in the social pyramid were based on colour."

Frank J. Yurco wrote in 1990: "When you talk about Egypt, it's just not right to talk about black or white .... To take the terminology here in the United States and graft it onto Africa is anthropologically inaccurate". Yurco added that "We are applying a racial divisiveness to Egypt that they would never have accepted, They would have considered this argument absurd, and that is something we could really learn from." Yurco wrote in 1996 that "the peoples of Egypt, the Sudan, and much of North-East Africa are generally regarded as a Nilotic continuity, with widely ranging physical features (complexions light to dark, various hair and craniofacial types)". In a 1989 article, he elaborated: "In short, ancient Egypt, like modern Egypt, consisted of a very heterogeneous population".

Bernard R. Ortiz De Montellano wrote in 1993: "The claim that all Egyptians, or even all the pharaohs, were black, is not valid. Most scholars believe that Egyptians in antiquity looked pretty much as they look today, with a gradation of darker shades toward the Sudan".

Nancy Lovell wrote in 1999 that studies of skeletal remains indicate that the physical characteristics of ancient southern Egyptians and Nubians were "within the range of variation" for both ancient and modern indigenous peoples of the Sahara and tropical Africa, and that the distribution of population characteristics "seems to follow a clinal pattern from south to north", which may be explained by natural selection as well as gene flow between neighboring populations. She also wrote that the archaeological and inscriptional evidence for contact between Egypt and Syro-Palestine "suggests that gene flow between these areas was very likely," and that the early Nile Valley populations were "part of an African lineage, but exhibiting local variation".

Stuart Tyson Smith wrote in 2001: "Any characterization of race of the ancient Egyptians depends on modern cultural definitions, not on scientific study. Thus, by modern American standards it is reasonable to characterize the Egyptians as 'black', while acknowledging the scientific evidence for the physical diversity of Africans."

Donald B. Redford wrote in 2004 that: "The old notion of waves of "races" flowing up the Nile Valley, effecting cultural change and improvement, is now known to be as erroneous as it was simplistic. New ideas need not come by means of invasion: occasionally they are indigenous and may parallel similar discoveries elsewhere which are wholly unrelated." He also wrote: "It would be interesting to know how the Nubians reacted to the racial attitudes Egypt manifested towards them, and in particular whether they shared a similar xenophobic aversion to all Egyptians."

Barry J. Kemp wrote in 2007 that the black/white argument, though politically understandable, is an oversimplification that hinders an appropriate evaluation of the scientific data on the ancient Egyptians since it does not take into consideration the difficulty in ascertaining complexion from skeletal remains. It also ignores the fact that Africa is inhabited by many other populations besides Bantu-related ("Negroid") groups. He wrote that in reconstructions of life in ancient Egypt, modern Egyptians would therefore be the most logical and closest approximation to the ancient Egyptians.

S. O. Y. Keita wrote in 2008 that "There is no scientific reason to believe that the primary ancestors of the Egyptian population emerged and evolved outside of northeast Africa.... The basic overall genetic profile of the modern population is consistent with the diversity of ancient populations that would have been indigenous to northeastern Africa and subject to the range of evolutionary influences over time, although researchers vary in the details of their explanations of those influences."

Barbara Mertz wrote in 2011: "Egyptian civilization was not Mediterranean or African, Semitic or Hamitic, black or white, but all of them. It was, in short, Egyptian."

Kathryn Bard wrote in 2014: "Egyptians were the indigenous farmers of the lower Nile valley, neither black nor white as races are conceived of today".

Nicky Nielsen wrote in 2020: "Ancient Egypt was neither black nor white, and the repeated attempt by advocates of either ideology to seize the ownership of ancient Egypt simply perpetuates an old tradition: one of removing agency and control of their heritage from the modern population living along the banks of the Nile."

Marc Van De Mieroop wrote in 2021: "Some scholars have tried to determine what Egyptians could have looked like by comparing their skeletal remains with those of recent populations, but the samples are so limited and the interpretations so fraught with uncertainties that this is an unreliable approach". He concluded that ancient Egypt's "location at the edge of northeast Africa and its geography as a corridor between that continent and Asia opened it up to influences from all directions, in terms of both culture and of demography."

Some modern views on bias in Egyptology

Various scholars have highlighted the role of colonial racism in shaping the attitudes of early Egyptologists, and criticized the continued over-representation of North American and European perspectives in the field. Diop in his work, "The African Origin of Civilization" argued that the prevailing views in Egyptology were driven by biased scholarship and colonial attitudes. Similarily, Bruce Trigger wrote that early modern scholarship on the Nile Valley populations had been "marred by a confusion of race, language, and culture and by an accompanying racism".

Frank Martin, critical of bias in Western scholarship and the shifting consensus, stated in 1984 that "Egyptologists, archaeologists, anthropologists, classicists, and scholars from other disciplines who have touched on the question almost invariably have asserted that Pharaonic Egyptians were whites. At first, many even dogmatically asserted that Egyptians were of Asiatic origin. That untenable position seems to have been abandoned by most serious contemporary scholars, but these same scholars have been reluctant to acknowledge the fact of the African origin of Egyptians in the sense of their being racially as well as culturally and linguistically related to Africans to the south and west of Egypt, thus leaving us with a people who seem to stand by themselves and have no ethnic and linguistic ties with any other people”.

Toby Wilkinson in 1999 outlined that "The Theory of the 'dynastic race'- a 'master race" of invaders from the east, to be responsible for imposing civilisation on the 'primitive' and unsophisticated indigenous Egyptians - had been articlated by Petrie (1939)...was still being espoused enthusiastically by scholars such as Emery (1961) and Edwards (1971)" although he felt that there was a change in perception among scholars at the time.

Stephen Quirke wrote in 2011 that the UNESCO-sponsored conference on the General History of Africa in 1974 "did not change the Eurocentric climate of research" and of the need to incorporate both African-centred studies and White European, academic persepectives. He later outlines that "research conferences and publications on the history and language of Kemet [Egypt] remain dominated.....by those brought up and trained in European, not African societies and languages (which include Arabic)".

Sally-Ann Ashton wrote in 2011: "The fact that Ancient Egypt is forced to justify its African identity through its geographical location has not gone unnoticed....critics of the mainstream Eurocentric view of Ancient Egypt claim that not only is the connection between Egypt and Africa neglected, it is consciously denied". She later outlines "This is partly the legacy of the "rediscovery" of Egypt by Europe at the end of the 18th century. In addition to this historical context, Egyptology as a discipline is dominated by scholars who are White Europeans or North Americans".

Uros Matic wrote in 2018 that the residue of scientific racism and old approaches were still present in modern archaeology and historiography of ancient Egypt and Nubia. He outlines that "although later authors such as such as Trigger and Tรถrรถk distanced themselves from racial science and anthropology, they kept this approach to iconography. What we can notice here are the remnants of the old colonialist episteme which appear in the works of Trigger, Tรถrรถk, and Donald B. Redford, however now being part of a new episteme". Matic concludes that "Epistemological de-colonization of our discipline should not only be a postcolonial criticism of its past, but a fundamental rethinking of its colonial remnants in the present, and how these remnants still shape our interpretations of Egypt and Nubia".

Stuart Tyson Smith wrote in 2018 that a common practice among Egyptologists was to "divorce Egypt from its proper northeast African context, instead framing it as fundamentally part of a Near Eastern or “Mediterranean” economic, social and political sphere, hardly African at all or at best a crossroad between the Near East, the eastern Mediterranean and Africa, which carries with it the implication that it is ultimately not really part of Africa". He explicitly criticises Van De Mieroop's comments that ancient Egypt was clearly ‘in Africa’ it was not so clearly‘of Africa’ as reflecting "long-standing Egyptological biases". He concludes that the interrelated cultural features shared between northeast African dynamic and Pharonic Egypt are not “survivals” or coincidence, but shared traditions with common origins in the deep past".

Marc Van De Mieroop wrote in 2021: "It was only recently that traditional scholarship started to acknowledge the African background of Egyptian culture, partly in response to world history's aim to replace dominant western-centered narratives with others than focused more on the contributions of other regions, including Africa. At the same time, primarily African diaspora communities wanted the continent's ancient history to be approached outside a Eurocentric context, and insisted, for example, on the use of ancient Egyptian term kemet instead of the European one".

Keith Crawford in 2021 presented a critique of the "Black Pharaohs" narrative accepted among mainstream scholars in which the Twenty-fifth Dynasty rulers were the only dynasty of Black African origin and academic representation of Egyptian-Kushite interactions. He concludes "The separation of Egypt from Africa, beginning nearly two centuries ago, resulted from Egyptologists, historians, and anthropologists interpreting archaeological finds and physical remains through a prism blurred by the racism of the time. These views have persisted to this day, despite overwhelming evidence that refutes them".

Genetic studies have been criticised by several scholars for a range of methodological problems and providing misleading, interpretations on racial classifications. Specifically, Keita and Kittles argue that DNA studies applied to the Nile Valley region have downplayed or excluded data on comparable, African populations in order to maintain certain racial models along with pre-selected data categories. Boyce and Keita in a later study, argue that certain studies have adopted a selective approach in sampling, such as using samples drawn mostly from northern (Lower) Egypt, which has historically had the presence of more foreigners from the Mediterranean and the Near East, and using those samples as representing the rest of Egypt. Thus, excluding the 'darker' south or Upper Egypt which presents a false impression of Egyptian variability. The authors also note that chromosonial patterns have featured inconsistent labelling such as Haplotype V as seen the with use of misleading terms like "Arabic" to describe it, implying this haplotype is of 'Middle Eastern' origins. However, when the hapotype V variant is looked at in context, it does have a very high prevalence in African countries above the Sahara and in Ethiopia.

Present-day controversies

Today the issues regarding the race of the ancient Egyptians are "troubled waters which most people who write about ancient Egypt from within the mainstream of scholarship avoid." The debate, therefore, takes place mainly in the public sphere and tends to focus on a small number of specific issues.

Tutankhamun

Several scholars, including Diop, have claimed that Tutankhamun was black, and have protested that attempted reconstructions of Tutankhamun's facial features (as depicted on the cover of National Geographic magazine) have represented the king as "too white". Among these writers was Chancellor Williams, who argued that King Tutankhamun, his parents, and grandparents were black.

Forensic artists and physical anthropologists from Egypt, France, and the United States independently created busts of Tutankhamun, using a CT-scan of the skull. Biological anthropologist Susan Anton, the leader of the American team, said the race of the skull was "hard to call". She stated that the shape of the cranial cavity indicated an African, while the nose opening suggested narrow nostrils, which is usually considered to be a European characteristic. The skull was thus concluded to be that of a North African. Other experts have argued that neither skull shapes nor nasal openings are a reliable indication of race.

Although modern technology can reconstruct Tutankhamun's facial structure with a high degree of accuracy, based on CT data from his mummy, determining his skin tone and eye color is impossible. The clay model was therefore given a coloring, which, according to the artist, was based on an "average shade of modern Egyptians".

Queen Tiye, grandmother of Tutankhamun

Terry Garcia, National Geographic's executive vice president for mission programs, said, in response to some of those protesting against the Tutankhamun reconstruction:

The big variable is skin tone. North Africans, we know today, had a range of skin tones, from light to dark. In this case, we selected a medium skin tone, and we say, quite up front, 'This is midrange.' We will never know for sure what his exact skin tone was or the color of his eyes with 100% certainty.... Maybe in the future, people will come to a different conclusion.

When pressed on the issue by American activists in September 2007, the Secretary General of the Egyptian Supreme Council of Antiquities, Zahi Hawass stated "Tutankhamun was not black."

In a November 2007 publication of Ancient Egypt magazine, Hawass asserted that none of the facial reconstructions resemble Tut and that, in his opinion, the most accurate representation of the boy king is the mask from his tomb. The Discovery Channel commissioned a facial reconstruction of Tutankhamun, based on CT scans of a model of his skull, back in 2002.

In 2011, the genomics company iGENEA launched a Tutankhamun DNA project based on genetic markers that it indicated it had culled from a Discovery Channel special on the pharaoh. According to the firm, the microsatellite data suggested that Tutankhamun belonged to the haplogroup R1b1a2, the most common paternal clade among males in Western Europe. Carsten Pusch and Albert Zink, who led the unit that had extracted Tutankhamun's DNA, chided iGENEA for not liaising with them before establishing the project. After examining the footage, they also concluded that the methodology the company used was unscientific with Putsch calling them "simply impossible".

Cleopatra

The race and skin color of Cleopatra VII, the last active Hellenistic ruler of the Macedonian Greek Ptolemaic dynasty of Egypt, established in 323 BCE, has also caused some debate, although generally not in scholarly sources. For example, the article "Was Cleopatra Black?" was published in Ebony magazine in 2012, and an article about Afrocentrism from the St. Louis Post-Dispatch mentions the question, too. Mary Lefkowitz, Professor Emerita of Classical Studies at Wellesley College, traces the main origins of the black Cleopatra claim to the 1946 book by J.A. Rogers called "World's Great Men of Color.", although noting that the idea of Cleopatra as black goes back to at least the 19th century. Lefkowitz refutes Rogers' hypothesis, on various scholarly grounds. The black Cleopatra claim was further revived in an essay by afrocentrist John Henrik Clarke, chair of African history at Hunter College, entitled "African Warrior Queens." Lefkowitz notes the essay includes the claim that Cleopatra described herself as black in the New Testament's Book of Acts – when in fact Cleopatra had died more than sixty years before the death of Jesus Christ.

The Berlin Cleopatra, a Roman sculpture and marble portrait of the queen in the Altes Museum, 1st century BC 

Scholars identify Cleopatra as essentially of Greek ancestry with some Persian and Syrian ancestry, based on the fact that her Macedonian Greek family (the Ptolemaic dynasty) had intermingled with the Seleucid aristocracy of the time. Grant states that Cleopatra probably had not a drop of Egyptian blood and that she "would have described herself as Greek." Roller notes that "there is absolutely no evidence" that Cleopatra was racially black African as claimed by what he dismisses as generally not "credible scholarly sources." Cleopatra's official coinage (which she would have approved) and the three portrait busts of her which are considered authentic by scholars, all match each other, and they portray Cleopatra as a Greek woman. Polo writes that Cleopatra's coinage presents her image with certainty, and asserts that the sculpted portrait of the "Berlin Cleopatra" head is confirmed as having a similar profile.

In 2009, a BBC documentary speculated that Cleopatra might have been part North African. This was based largely on the claims of Hilke Thรผr of the Austrian Academy of Sciences, who in the 1990s had examined a headless skeleton of a female child in a 20 BCE tomb in Ephesus (modern Turkey), together with the old notes and photographs of the now-missing skull. Thรผr hypothesized the body as that of Arsinoe, half-sister to Cleopatra. Arsinoe and Cleopatra shared the same father (Ptolemy XII Auletes) but had different mothers, with Thรผr claiming the alleged African ancestry came from the skeleton's mother. To date it has never been definitively proved that the skeleton is that of Arsinoe IV. When a DNA test attempted to determine the identity of the child, it was impossible to get an accurate reading since the bones had been handled too many times, and the skull had been lost in Germany during World War II. Numerous studies have shown that cranial variation has a low correlation with race, and rather that cranial variation was strongly correlated with climate variables. Mary Beard states that the age of the skeleton is too young to be that of Arsinoe (the bones said to be that of a 15–18-year-old child, with Arsinoe being around her mid twenties at her death).

Great Sphinx of Giza

The Sphinx in profile in 2010

The identity of the model for the Great Sphinx of Giza is unknown. Most experts believe that the face of the Sphinx represents the likeness of the Pharaoh Khafra, although a few Egyptologists and interested amateurs have proposed different hypotheses.

An early description of the Sphinx, "typically negro in all its features", is recorded in the travel notes of a French scholar, Volney, who visited Egypt between 1783 and 1785 along with French novelist Gustave Flaubert. A similar description was given in the "well-known book" by Vivant Denon, where he described the sphinx as "the character is African; but the mouth, the lips of which are thick." Following Volney, Denon, and other early writers, numerous Afrocentric scholars, such as Du Bois, Diop and Asante have characterized the face of the Sphinx as Black, or "Negroid".

American geologist Robert M. Schoch has written that the "Sphinx has a distinctive African, Nubian, or Negroid aspect which is lacking in the face of Khafre", but he was described by others such as Ronald H. Fritze and Mark Lehner of being a "pseudoscientific writer". David S. Anderson writes in Lost City, Found Pyramid: Understanding Alternative Archaeologies and Pseudoscientific Practices that Van Sertima's claim that "the sphinx was a portrait statue of the black pharoah Khafre" is a form of "pseudoarchaeology" not supported by evidence. He compares it to the claim that Olmec colossal heads had "African origins", which is not taken seriously by Mesoamerican scholars such as Richard Diehl and Ann Cyphers.

Kemet

km in Egyptian hieroglyphs
km biliteral kmt (place) kmt (people)
km
km
t O49
km
t
A1B1Z3

Ancient Egyptians referred to their homeland as Kmt (conventionally pronounced as Kemet). According to Cheikh Anta Diop, the Egyptians referred to themselves as "Black" people or kmt, and km was the etymological root of other words, such as Kam or Ham, which refer to Black people in Hebrew tradition. A review of David Goldenberg's The Curse of Ham: Race and Slavery in Early Judaism, Christianity and Islam states that Goldenberg "argues persuasively that the biblical name Ham bears no relationship at all to the notion of blackness and as of now is of unknown etymology". Diop, William Leo Hansberry, and Aboubacry Moussa Lam have argued that kmt was derived from the skin color of the Nile valley people, which Diop claimed was black. The claim that the ancient Egyptians had black skin has become a cornerstone of Afrocentric historiography.

Mainstream scholars hold that kmt means "the black land" or "the black place", and that this is a reference to the fertile black soil that was washed down from Central Africa by the annual Nile inundation. By contrast the barren desert outside the narrow confines of the Nile watercourse was called dลกrt (conventionally pronounced deshret) or "the red land". Raymond Faulkner's Concise Dictionary of Middle Egyptian translates kmt into "Egyptians", Gardiner translates it as "the Black Land, Egypt".

At the UNESCO Symposium in 1974, Sauneron, Obenga, and Diop concluded that KMT and KM meant black. However, Sauneron clarified that the adjective Kmtyw means "people of the black land" rather than "black people", and that the Egyptians never used the adjective Kmtyw to refer to the various black peoples they knew of, they only used it to refer to themselves.

Ancient Egyptian art

Ancient Egyptian tombs and temples contained thousands of paintings, sculptures, and written works, which reveal a great deal about the people of that time. However, their depictions of themselves in their surviving art and artifacts are rendered in sometimes symbolic, rather than realistic, pigments. As a result, ancient Egyptian artifacts provide sometimes conflicting and inconclusive evidence of the ethnicity of the people who lived in Egypt during dynastic times.

In their own art, "Egyptians are often represented in a color that is officially called dark red", according to Diop. Arguing against other theories, Diop quotes Champollion-Figeac, who states, "one distinguishes on Egyptian monuments several species of blacks, differing...with respect to complexion, which makes Negroes black or copper-colored." Regarding an expedition by King Sesostris, Cherubini states the following concerning captured southern Africans, "except for the panther skin about their loins, are distinguished by their color, some entirely black, others dark brown. University of Chicago scholars assert that Nubians are generally depicted with black paint, but the skin pigment used in Egyptian paintings to refer to Nubians can range "from dark red to brown to black". This can be observed in paintings from the tomb of the Egyptian Huy, as well as Ramses II's temple at Beit el-Wali. Also, Snowden indicates that Romans had accurate knowledge of "negroes of a red, copper-colored complexion ... among African tribes".

Conversely, in 2003 Najovits wrote that "Egyptian art depicted Egyptians on the one hand and Nubians and other blacks on the other hand with distinctly different ethnic characteristics and depicted this abundantly and often aggressively. The Egyptians accurately, arrogantly and aggressively made national and ethnic distinctions from a very early date in their art and literature." He continues, "There is an extraordinary abundance of Egyptian works of art which clearly depicted sharply contrasted reddish-brown Egyptians and black Nubians."

Barbara Mertz in 2011 wrote in Red Land, Black Land: Daily Life in Ancient Egypt: "The concept of race would have been totally alien to them [Ancient Egyptians] [..]The skin color that painters usually used for men is a reddish brown. Women were depicted as lighter in complexion, perhaps because they didn’t spend so much time out of doors. Some individuals are shown with black skins. I cannot recall a single example of the words “black,” “brown,” or “white” being used in an Egyptian text to describe a person." She gives the example of one of Thutmose III’s “sole companions”, who was Nubian or Kushite. In his funerary scroll, he is shown with dark brown skin instead of the conventional reddish brown used for Egyptians.

Table of Nations controversy

The "Table of Nations", from Lepsius: Top row, left to right: "Aamu" (Asiatics), "Nehesu" (Nubians), and "Themehu" (Libyans); middle row: a deity, "Reth" (Egyptians), "Aamu" (Asiatics).

However, Manu Ampim, a professor at Merritt College specializing in African and African American history and culture, claims in the book Modern Fraud: The Forged Ancient Egyptian Statues of Ra-Hotep and Nofret, that many ancient Egyptian statues and artworks are modern frauds that have been created specifically to hide the "fact" that the ancient Egyptians were black, while authentic artworks that demonstrate black characteristics are systematically defaced or even "modified". Ampim repeatedly makes the accusation that the Egyptian authorities are systematically destroying evidence that "proves" that the ancient Egyptians were black, under the guise of renovating and conserving the applicable temples and structures. He further accuses "European" scholars of wittingly participating in and abetting this process.

Ampim has a specific concern about the painting of the "Table of Nations" in the Tomb of Ramesses III (KV11). The "Table of Nations" is a standard painting that appears in a number of tombs, and they were usually provided for the guidance of the soul of the deceased. Among other things, it described the "four races of men" as follows: (translation by E.A. Wallis Budge) "The first are RETH, the second are AAMU, the third are NEHESU, and the fourth are THEMEHU. The RETH are Egyptians, the AAMU are dwellers in the deserts to the east and north-east of Egypt, the NEHESU are the black races, and the THEMEHU are the fair-skinned Libyans."

The archaeologist Karl Richard Lepsius documented many ancient Egyptian tomb paintings in his work Denkmรคler aus Aegypten und Aethiopien. In 1913, after the death of Lepsius, an updated reprint of the work was produced, edited by Kurt Sethe. This printing included an additional section, called the "Ergรคnzungsband" in German, which incorporated many illustrations that did not appear in Lepsius' original work. One of them, plate 48, illustrated one example of each of the four "nations" as depicted in KV11, and shows the "Egyptian nation" and the "Nubian nation" as identical to each other in skin color and dress. Professor Ampim has declared that plate 48 is a true reflection of the original painting, and that it "proves" that the ancient Egyptians were identical in appearance to the Nubians, even though he admits no other examples of the "Table of Nations" show this similarity. He has further accused "Euro-American writers" of attempting to mislead the public on this issue.

The late Egyptologist Frank J. Yurco visited the tomb of Ramesses III (KV11), and in a 1996 article on the Ramesses III tomb reliefs he pointed out that the depiction of plate 48 in the Ergรคnzungsband section is not a correct depiction of what is actually painted on the walls of the tomb. Yurco notes, instead, that plate 48 is a "pastiche" of samples of what is on the tomb walls, arranged from Lepsius' notes after his death, and that a picture of a Nubian person has erroneously been labeled in the pastiche as an Egyptian person. Yurco points also to the much more recent photographs of Dr. Erik Hornung as a correct depiction of the actual paintings. (Erik Hornung, The Valley of the Kings: Horizon of Eternity, 1990). Ampim nonetheless continues to claim that plate 48 shows accurately the images that stand on the walls of KV11, and he categorically accuses both Yurco and Hornung of perpetrating a deliberate deception for the purposes of misleading the public about the true race of the ancient Egyptians.

Fayyum mummy portraits

The realistic Fayum mummy portraits show the diversity of Egyptians in the Roman period

The Roman era Fayum mummy portraits attached to coffins containing the latest dated mummies discovered in the Faiyum Oasis represent a population of both native Egyptians and those with mixed Greek heritage. The dental morphology of the mummies align more with the indigenous North African population than Greek or other later colonial European settlers.

Black queen controversy

The late British Africanist Basil Davidson stated "Whether the Ancient Egyptians were as black or as brown in skin color as other Africans may remain an issue of emotive dispute; probably, they were both. Their own artistic conventions painted them as pink, but pictures on their tombs show they often married queens shown as entirely black being from the south."

Queen Ahmose-Nefertari of the 18th Dynasty

Ahmose-Nefertari is an example. In most depictions of Ahmose-Nefertari, she is pictured with black skin, while in some instances her skin is blue or red. In 1939 Flinders Petrie said "an invasion from the south...established a black queen as the divine ancestress of the XVIIIth dynasty" He also said "a possibility of the black being symbolic has been suggested" and "Nefertari must have married a Libyan, as she was the mother of Amenhetep I, who was of fair Libyan style." In 1961 Alan Gardiner, in describing the walls of tombs in the Deir el-Medina area, noted in passing that Ahmose-Nefertari was "well represented" in these tomb illustrations, and that her countenance was sometimes black and sometimes blue. He did not offer any explanation for these colors, but noted that her probable ancestry ruled out that she might have had black blood. In 1974, Diop described Ahmose-Nefertari as "typically negroid." In the controversial book Black Athena, the hypotheses of which have been widely rejected by mainstream scholarship, Martin Bernal considered her skin color in paintings to be a clear sign of Nubian ancestry. In 1981 Michel Gitton noted that while in most artistic depictions of the queen she is pictured with black complexion, there are other cases in which she is shown with a pink, golden, blue, or dark red skin color.

Gitton called the issue of Ahmose-Nefertari's black color "a serious gap in the Egyptological research, which allows approximations or untruths". He pointed out that there is no known depiction of her painted during her lifetime (she is represented with the same light skin as other represented individuals in tomb TT15, before her deification); the earliest black skin depiction appears in tomb TT161, circa 150 years after her death. Barbara Lesko wrote in 1996 that Ahmose-Nefertari was "sometimes portrayed by later generations as having been black, although her coffin portrait gives her the typical light yellow skin of women."

In 2003, Betsy Bryan wrote in The Oxford History of Ancient Egypt that "the factors linking Amenhotep I and his mother with the necropolis region, with deified rulers, and with rejuvenation generally was visually transmitted by representations of the pair with black or blue skin – both colours of resurrection." In 2004 Aidan Dodson and Dyan Hilton recognized in a later depiction of the queen, "the black skin of a deity of resurrection" in connection to her role as a patron goddess of the Theban necropolis. Scholars such as Joyce Tyldesley, Sigrid Hodel-Hoenes, and Graciela Gestoso Singer, argued that the skin color of Ahmose-Nefertari is indicative of her role as a goddess of resurrection, since black is both the color of the fertile land of Egypt and that of Duat, the underworld. Singer recognizes that "Some scholars have suggested that this is a sign of Nubian ancestry." Singer also states a statuette of Ahmose-Nefertari at the Museo Egizio in Turin which shows her with a black face, though her arms and feet are not darkened, thus suggesting that the black coloring has an iconographic motive and does not reflect her actual appearance. In 2014, Margaret Bunson wrote that "the unusual depictions of Ahmose-Nefertari in blue-black tones of deification reflect her status and cult." In a wooden votive statue of Ahmose-Nefertari, currently in the Louvre museum, her skin was painted red, a color commonly seen symbolizing life or a higher being, or elevated status.

Historical hypotheses

Since the second half of the 20th century, typological and hierarchical models of race have increasingly been rejected by scientists, and most scholars have held that applying modern notions of race to ancient Egypt is anachronistic. The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt). At the UNESCO symposium in 1974, most participants concluded that the ancient Egyptian population was indigenous to the Nile Valley, and was made up of people from north and south of the Sahara who were differentiated by their color.

Black Egyptian hypothesis

The Black Egyptian hypothesis is the hypothesis that ancient Egypt was a "Black" civilization. Although there is consensus that Ancient Egypt was indigenous to Africa, the hypothesis that it was a "Black" civilization has met with "profound" disagreement, particularly because it rests upon a scientifically outdated conception of race.

The Black Egyptian hypothesis includes a particular focus on links to Sub Saharan cultures and the questioning of the race of specific notable individuals from Dynastic times, including Tutankhamun the person represented in the Great Sphinx of Giza, and the Greek Ptolemaic queen Cleopatra. Advocates of the Black African model rely heavily on writings from Classical Greek historians, including Strabo, Diodorus Siculus, and Herodotus. Advocates claim that these "classical" authors referred to Egyptians as "Black with woolly hair". The Greek word used was "melanchroes", and the English language translation of this Greek word is disputed, being translated by many as "dark skinned" and by many others as "black". Diop said "Herodotus applied melanchroes to both Ethiopians and Egyptians...and melanchroes is the strongest term in Greek to denote blackness." Snowden claims that Diop is distorting his classical sources and is quoting them selectively. There is dispute about the historical accuracy of the works of Herodotus – some scholars support the reliability of Herodotus while other scholars regard his works as being unreliable as historical sources, particularly those relating to Egypt.

Other claims used to support the Black Hypothesis included anthropological measurements of Egyptian mummies, testing melanin levels in a small sample of mummies, language affinities between ancient Egyptian language and sub-saharan languages, interpretations of the origin of the name Kmt, conventionally pronounced Kemet, used by the ancient Egyptians to describe themselves or their land (depending on points of view), biblical traditions, shared B blood group between Egyptians and West Africans, and interpretations of the depictions of the Egyptians in numerous paintings and statues. The hypothesis also claimed cultural affiliations, such as circumcision, matriarchy, totemism, hair braiding, head binding, and kingship cults. Artifacts found at Qustul (near Abu Simbel – Modern Sudan) in 1960–64 were seen as showing that ancient Egypt and the A-Group culture of Nubia shared the same culture and were part of the greater Nile Valley sub-stratum, but more recent finds in Egypt indicate that the Qustul rulers probably adopted/emulated the symbols of Egyptian pharaohs. Authors and critics state the hypothesis is primarily adopted by Afrocentrists.

The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt), and that modern racial categories like "Black" are social constructs inapplicable to Ancient Egypt.

Asiatic race theory

The Asiatic race theory, which has been rejected by mainstream scholarship, is the hypothesis that the ancient Egyptians were the lineal descendants of the biblical Ham, through his son Mizraim.

This theory was the most dominant view from the Early Middle Ages (c. 500 AD) all the way up to the early 19th century. The descendants of Ham were traditionally considered to be the darkest-skinned branch of humanity, either because of their geographic allotment to Africa or because of the Curse of Ham. Thus, Diop cites Gaston Maspero "Moreover, the Bible states that Mesraim, son of Ham, brother of Chus (Kush) ... and of Canaan, came from Mesopotamia to settle with his children on the banks of the Nile."

By the 20th century, the Asiatic race theory and its various offshoots were abandoned but were superseded by two related theories: the Eurocentric Hamitic hypothesis, asserting that a Caucasian racial group moved into North and East Africa from early prehistory subsequently bringing with them all advanced agriculture, technology and civilization, and the Dynastic race theory, proposing that Mesopotamian invaders were responsible for the dynastic civilization of Egypt (c. 3000 BC). In sharp contrast to the Asiatic race theory, neither of these theories proposes that Caucasians were the indigenous inhabitants of Egypt.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, none of the participants explicitly voiced support for any theory where Egyptians were Caucasian with a dark pigmentation.". The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Caucasian / Hamitic hypothesis

The Caucasian hypothesis, which has been rejected by mainstream scholarship, is the hypothesis that the Nile valley "was originally peopled by a branch of the Caucasian race". It was proposed in 1844 by Samuel George Morton, who acknowledged that Negroes were present in ancient Egypt but claimed they were either captives or servants. George Gliddon (1844) wrote: "Asiatic in their origin .... the Egyptians were white men, of no darker hue than a pure Arab, a Jew, or a Phoenician."

The similar Hamitic hypothesis, which has been rejected by mainstream scholarship, developed directly from the Asiatic Race Theory, and argued that the Ethiopid and Arabid populations of the Horn of Africa were the inventors of agriculture and had brought all civilization to Africa. It asserted that these people were Caucasians, not Negroid. It also rejected any Biblical basis despite using Hamitic as the theory's name. Charles Gabriel Seligman in his Some Aspects of the Hamitic Problem in the Anglo-Egyptian Sudan (1913) and later works argued that the ancient Egyptians were among this group of Caucasian Hamites, having arrived in the Nile Valley during early prehistory and introduced technology and agriculture to primitive natives they found there.

The Italian anthropologist Giuseppe Sergi (1901) believed that ancient Egyptians were the Eastern African (Hamitic) branch of the Mediterranean race, which he called "Eurafrican". According to Sergi, the Mediterranean race or "Eurafrican" contains three varieties or sub-races: the African (Hamitic) branch, the Mediterranean "proper" branch and the Nordic (depigmented) branch. Sergi maintained in summary that the Mediterranean race (excluding the depigmented Nordic or 'white') is: "a brown human variety, neither white nor Negroid, but pure in its elements, that is to say not a product of the mixture of Whites with Negroes or Negroid peoples". Grafton Elliot Smith modified the theory in 1911, stating that the ancient Egyptians were a dark haired "brown race", most closely "linked by the closest bonds of racial affinity to the Early Neolithic populations of the North African littoral and South Europe", and not Negroid. Smith's "brown race" is not synonymous or equivalent with Sergi's Mediterranean race. The Hamitic Hypothesis was still popular in the 1960s and late 1970s and was supported notably by Anthony John Arkell and George Peter Murdock.

At the UNESCO "Symposium on the Peopling of Ancient Egypt and the Deciphering of the Meroitic Script" in Cairo in 1974, none of the participants explicitly voiced support for any theory where Egyptians were Caucasian with a dark pigmentation." The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Turanid race hypothesis

The Turanid race hypothesis, which has been rejected by mainstream scholarship, is the hypothesis that the ancient Egyptians belonged to the Turanid race, linking them to the Tatars.

It was proposed by Egyptologist Samuel Sharpe in 1846, who was "inspired" by some ancient Egyptian paintings, which depict Egyptians with sallow or yellowish skin. He said "From the colour given to the women in their paintings we learn that their skin was yellow, like that of the Mongul Tartars, who have given their name to the Mongolian variety of the human race.... The single lock of hair on the young nobles reminds us also of the Tartars."

The current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

Dynastic race theory

The Dynastic race theory, which has been rejected by mainstream scholarship, is the hypothesis that a Mesopotamian force had invaded Egypt in predynastic times, imposed itself on the indigenous Badarian people, and become their rulers. It further argued that the Mesopotamian-founded state or states then conquered both Upper and Lower Egypt and founded the First Dynasty of Egypt.

It was proposed in the early 20th century by Egyptologist Sir William Matthew Flinders Petrie, who deduced that skeletal remains found at pre-dynastic sites at Naqada (Upper Egypt) indicated the presence of two different races, with one race differentiated physically by a noticeably larger skeletal structure and cranial capacity. Petrie also noted new architectural styles—the distinctly Mesopotamian "niched-facade" architecture—pottery styles, cylinder seals and a few artworks, as well as numerous Predynastic rock and tomb paintings depicting Mesopotamian style boats, symbols, and figures. Based on plentiful cultural evidence, Petrie concluded that the invading ruling elite was responsible for the seemingly sudden rise of Egyptian civilization. In the 1950s, the Dynastic Race Theory was widely accepted by mainstream scholarship.

While there is clear evidence the Naqada II culture borrowed abundantly from Mesopotamia, the Naqada II period had a large degree of continuity with the Naqada I period, and the changes which did happen during the Naqada periods happened over significant amounts of time. The most commonly held view today is that the achievements of the First Dynasty were the result of a long period of cultural and political development, and the current position of modern scholarship is that the Egyptian civilization was an indigenous Nile Valley development (see population history of Egypt).

The Senegalese Egyptologist Cheikh Anta Diop, fought against the Dynastic Race Theory with their own "Black Egyptian" theory and claimed, among other things, that Eurocentric scholars supported the Dynastic Race Theory "to avoid having to admit that Ancient Egyptians were black". Martin Bernal proposed that the Dynastic Race theory was conceived by European scholars to deny Egypt its African roots.

Fearmongering

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