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Saturday, March 21, 2026

American nationalism

From Wikipedia, the free encyclopedia
Schoolchildren in the United States reciting the Pledge of Allegiance in 1899

American nationalism is a form of nationalism in the United States, which can have civic, ethnic, cultural or economic influences. Essentially, it indicates the aspects that characterize and distinguish the United States as an autonomous political community. The term often explains efforts to reinforce its national identity and self-determination within its national and international affairs.

American nationalism has found expression throughout American history. The first Naturalization Act of 1790 passed by Congress and George Washington defined American identity and citizenship on racial lines, declaring that only "free white men of good character" could become citizens, and denying citizenship to enslaved black people and anyone of non-European stock; thus it was a form of ethnic nationalism. Some American scholars have argued that the United States government institutionalized a civic nationalism founded upon legal and rational concepts of citizenship, being based on common language and cultural traditions, and that the Founding Fathers of the United States established the country upon liberal and individualist principles.

History

Colonial

The United States traces its origins to the Thirteen Colonies founded by Britain in the 17th and early 18th century. Residents identified with Britain until the mid-18th century when the first sense of being "American" emerged. The Albany Plan proposed a union between the colonies in 1754. Although unsuccessful, it served as a reference for future discussions of independence.

John Trumbull's Declaration of Independence is often identified as a depiction of the signing of the Declaration, but it actually shows the five person drafting committee presenting its work to the Second Continental Congress.

American Revolution

Soon afterward, the colonies faced several common grievances over acts passed by the British Parliament, including taxation without representation. Americans were in general agreement that only their own colonial legislatures—and not Parliament in London—could pass internal taxes. Parliament vigorously insisted otherwise and no compromise was found. The London government punished Boston for the Boston Tea Party, and the Thirteen Colonies united and formed the Continental Congress, which lasted from 1774 to 1789. Fighting broke out in 1775 and the sentiment swung to independence in early 1776, influenced significantly by the appeal to American nationalism by Thomas Paine. His pamphlet Common Sense was a runaway best seller in 1776, read aloud in taverns and coffee houses. Congress unanimously issued a Declaration of Independence announcing a new nation of independent states had formed, the United States of America. American Patriots won the American Revolutionary War and received generous peace terms from Britain in 1783. The minority of Loyalists (loyal to King George III) could remain or leave, but about 80% remained and became full American citizens. Frequent parades along with new rituals and ceremonies—and a new flag—provided popular occasions for expressing a spirit of American nationalism.

The new nation operated under the very weak national government set up by the Articles of Confederation, and most Americans prioritized their state over the nation. Nationalists led by George Washington, Alexander Hamilton, and James Madison had Congress call a constitutional convention in 1787. It produced the Constitution for a strong national government which was debated in every state and unanimously adopted. It went into effect in 1789 as the first modern constitutional liberal democracy based on the consent of the governed, with Washington as the first President.

Westward Expansion

In an 1858 speech, future President Abraham Lincoln alluded to a form of American civic nationalism originating from the tenets of the Declaration of Independence as a force for national unity in the United States, stating that it was a method for uniting diverse peoples of different ethnic ancestries into a common nationality:

If they look back through this history to trace their connection with those days by blood, they find they have none, they cannot carry themselves back into that glorious epoch and make themselves feel that they are part of us, but when they look through that old Declaration of Independence they find that those old men say that "We hold these truths to be self-evident, that all men are created equal", and then they feel that moral sentiment taught in that day evidences their relation to those men, that it is the father of all moral principle in them, and that they have a right to claim it as though they were blood of the blood, and flesh of the flesh of the men who wrote the Declaration, and so they are. That is the electric cord in that Declaration that links the hearts of patriotic and liberty-loving men together, that will link those patriotic hearts as long as the love of freedom exists in the minds of men throughout the world.

— Abraham Lincoln, address to Chicagoan voters, July 10, 1858

American Civil War

White Southerners increasingly felt alienated—they saw themselves as becoming second-class citizens as aggressive anti-slavery Northerners tried to end their ability to enslave people to the fast-growing western territories. They questioned whether their loyalty to the nation trumped their commitment to their state and their way of life since it was so intimately bound up with slavery and whether they could enslave people. A sense of Southern nationalism was starting to emerge; however, it was rudimentary as late as 1860 when the election of Lincoln was a signal for most of the slave states in the South to secede and form a new nation. The Confederate government insisted the nationalism was real and imposed increasing burdens on the population in the name of independence and nationalism. The fierce combat record of the Confederates demonstrates their commitment to the death for independence. The government and army refused to compromise and were militarily overwhelmed in 1865. By the 1890s, the white South felt vindicated through its belief in the newly constructed memory of the Lost Cause of the Confederacy. The North came to accept or at least tolerate racial segregation and disfranchisement of black voters in the South. The spirit of American nationalism had returned to Dixie.

Reconstruction, Gilded Age and Progressive Era

An 1869 Thomas Nast cartoon espousing American exceptionalism shows Americans of different ancestries and ethnic backgrounds sit together at a dinner table with Columbia to enjoy a Thanksgiving meal as equal members of the American citizenry while Uncle Sam prepares and sets the table, thus espousing an inclusive form of American nationalism that is civic, where membership in the nation is not dependent upon ethnicity.

The North's triumph in the American Civil War marked a significant transition in American national identity. The ratification of the Fourteenth Amendment settled the fundamental question of national identity, such as the criteria for becoming a citizen of the United States. Everyone born in the territorial boundaries of the United States or those areas and subject to its jurisdiction was an American citizen, regardless of ethnicity or social status (indigenous people on reservations became citizens in 1924, while indigenous people off reservations had always been citizens).

In the early 20th century, one of the highest-profile advocates for American nationalism was Theodore Roosevelt. Roosevelt's policies both at home and abroad, which came to be known as the New Nationalism, included an element of a strong national identity. He insisted that one had to be 100% American, not a hyphenated American who juggled multiple loyalties.

With a fast-growing industrial economy, immigrants were welcome from Europe, Canada, Mexico, and Cuba, and millions came. Becoming a full citizen was easy, requiring the completion of paperwork over five years.  However, new Asian arrivals were not welcome. The U.S. imposed restrictions on most Chinese immigrants in the 1880s and informal restrictions on most Japanese in 1907. By 1924, it was difficult for any Asian to enter the United States, but children born in the United States to Asian parents were full citizens. The restrictions were ended on the Chinese in the 1940s and on other Asians in 1965.

Newspaper reporting the annexation of the Republic of Hawaii in 1898

World Wars and interwar period

After the United States entered World War I, nationalism surged. Americans enlisted in the military en masse, motivated by propaganda and war films. There was very little resistance to conscription.

By the First World War, many native-born Protestants were skeptical of recent immigrants to the United States, who were often Catholic or Jewish and spoke languages other than English in their daily lives. There was a strong belief among many in favor of "one hundred percent Americanism", in contrast to "hyphenated Americanism". This was exemplified by the film The Birth of a Nation in 1915 and the rise of the Ku Klux Klan in the 1910s and 20s.  In the early- to-mid 20th century, public school education became compulsory in many jurisdictions, with parochial schools being restricted or outlawed. The school day typically began with the Pledge of Allegiance. It was in this milieu that the Immigration Act of 1924 was passed to regulate immigration from Southern and Eastern Europe. However, during this period, citizenship was also extended to Native Americans, both on- and off-reservation, for the first time.

World War II led to unprecedented nationalism in the United States. After the 1941 Attack on Pearl Harbor, many Americans enlisted in the military. During the war, much of American life centered on contributing to the war effort, mainly through volunteer efforts, entry into the labor force, rationing, price controls, and income saving. Citizens willingly accepted these sacrifices out of a sense of nationalism, feeling they were for the greater good. Even members of anti-war groups like the pacifist churches, anti-war movement, and conscientious objectors abandoned their pacifism for the sake of the war, feeling that World War II was a just war.

Cold War

Following World War II and beginning with the Cold War, the United States emerged as a world superpower and abandoned its traditional policy of isolationism in favor of interventionism. With this, nationalism took on a new form in the U.S., as Americans began to view their country as a world police with the ultimate goal of eradicating communism from the world. This nationalist fervor was fueled by US involvement in the Korean War, Vietnam War, Bay of Pigs Invasion, and many other conflicts.[30]

Modern era

The September 11 attacks of 2001 led to a wave of nationalist expression in the United States. The start of the war on terror was accompanied by a rise in military enlistment that included not only lower-income Americans but also middle-class and upper-class citizens. This nationalism continued long into the War in Afghanistan and Iraq War.

Contemporary United States

Nationalism and Americanism remain topics in the modern United States. Political scientist Paul McCartney, for instance, argues that as a nation defined by a creed and sense of mission, Americans tend to equate their interests with those of humanity, which informs their global posture. In some instances, it may be considered a form of ethnocentrism and American exceptionalism.

Due to the distinctive circumstances involved throughout history in American politics, its nationalism has developed concerning loyalty to a set of liberal, universal political ideals and perceived accountability to propagate those principles globally. Acknowledging the conception of the United States as accountable for spreading liberal change and promoting democracy throughout the world's politics and governance has defined practically all of American foreign policy. Therefore, democracy promotion is not just another measure of foreign policy, but it is instead the fundamental characteristic of their national identity and political determination.

Varieties of American nationalism

In a 2016 paper in the American Sociological Review, "Varieties of American Popular Nationalism", sociologists Bart Bonikowski and Paul DiMaggio report on research findings supporting the existence of at least four kinds of American nationalists, including, groups which range from the smallest to the largest: (1) the disengaged, (2) creedal or civic nationalists, (3) ardent nationalists, and (4) restrictive nationalists.

Bonikowski and Dimaggio's analysis of these four groups found that ardent nationalists made up about 24% of their study, and they comprised the largest of the two groups Bonikowski and Dimaggio consider "extreme". Members of this group closely identified with the United States, were very proud of their country and strongly associated themselves with factors of national hubris. They felt that a "true American" must speak English and live in the U.S. for most of their life. Fewer, but 75%, believe that a "true American" must be a Christian, and 86% believe a "true American" must be born in the country. Further, ardent nationalists thought that Jews, Muslims, agnostics and naturalized citizens were something less than genuinely American. The second class, Bonikowski and DiMaggio considered "extreme", was the smallest of the four classes because its members comprised 17% of their respondents. The disengaged showed low pride in the government institutions, and they did not fully identify with the United States. Their lack of pride extended to American democracy, American history, political equality in the U.S., and the country's political influence. This group was the least nationalistic of all of the four groups which they identified.

Studies supported the idea that the American flag increased nationalism.

The two remaining classes were less homogeneous in their responses than the ardent nationalists and disengaged were. Restrictive nationalists had low levels of pride in America and its institutions, but they defined a "true American" in ways that were markedly "exclusionary". This group was the largest of the four because its members comprised 38% of the study's respondents. While their levels of national identification and pride were moderate, they espoused beliefs that caused them to hold restrictive definitions of "true Americans"; for instance, their definitions excluded non-Christians."

The final group to be identified was creedal nationalists (also known as civic nationalists), whose members made up 22% of the study's respondents who were studied. This group believed in liberal values, was proud of the United States, and its members held the fewest restrictions on who could be considered a true American. They closely identified with their country, which they felt "very close" to, and were proud of its achievements. Bonikowski and Dimaggio dubbed the group "creedal" because their beliefs most closely approximated the precepts of what is widely considered the American creed.

As part of their findings, the authors report that the connection between big money, religious belief, and national identity is significant. The belief that being a Christian is an integral part of what it means to be a "true American" is the most significant factor which separates the creedal nationalists and the disengaged from the restrictive and ardent nationalists. They also determined that their groupings cut across partisan boundaries, and they also help to explain what they perceive is the recent success of populist, nativist, and racist rhetoric in American politics.

According to a 2021 American Journal of Sociology study by Bart Bonikowski, Yuval Feinstein, and Sean Bock, competing understandings of American nationhood had emerged in the United States in the prior two decades. They find, "nationalism has become sorted by party, as Republican identifiers have come to define America in more exclusionary and critical terms and Democrats have increasingly endorsed inclusive and positive conceptions of nationhood."

Cultural nationalism

Cultural nationalism has historically been an integral element of American nationalism. Such cultural nationalists form group allegiances based on a common cultural heritage rather than race or political party. This heritage may include culture (Culture of the United States), language (English language), religion (Christianity), history (History of the United States), ideology (Democracy), and symbols (National symbols of the United States). Cultural nationalism is distinct from ethnic nationalism, in which race and ethnicity are emphasized over culture and language.

Nationalism gained a cultural character beginning in the late 18th century. Multiple historical ideas have shaped modern cultural nationalism in the U.S., including the concept of the nation state, the fusion of nationalism and religion into religious nationalism, and identity politics.

Civic nationalism

American nationalism sometimes takes the form of Civic nationalism, a liberal form of nationalism based on values such as freedom, equality, and individual rights. Civic nationalists view nationhood as a political identity. They argue that liberal democratic principles and loyalty define a civic nation. Membership is open to every citizen, regardless of culture, ethnicity, or language, as long as they believe in these values.

Trumpism

President Donald Trump was described as a nationalist, and he embraced the term himself. Several officials within his administration were described as representing a "nationalist wing" within the federal government, including former White House Chief Strategist Steve BannonSenior Advisor to the President Stephen MillerDirector of the National Trade Council Peter Navarro, former Deputy Assistant to the President Sebastian GorkaSpecial Assistant to the President Julia Hahn, former Deputy Assistant to the President for Strategic Communications Michael AntonSecretary of State Mike PompeoSecretary of Commerce Wilbur RossTrade Representative Robert Lighthizer, former acting Director of National Intelligence Richard Grenell, former National Security Advisor John R. Bolton and former National Security Advisor Michael Flynn.

In a February 2017 article in The Atlantic, journalist Uri Friedman described "populist economic nationalist" as a new nationalist movement "modeled on the 'populism' of the 19th-century U.S. President Andrew Jackson" which was introduced in Trump's remarks to the Republican National Convention in a speech written by Stephen Miller and Steve Bannon. Miller had adopted Senator Jeff Sessions' form of "nation-state populism" while working as his aide. By September 2017, The Washington Post journalist Greg Sargent observed that "Trump's nationalism" as "defined" by Bannon, Breitbart, Miller and "the rest of the 'populist economic nationalist' contingent around Trump" was beginning to have wavering support among Trump voters. Some Republican members of Congress were also described as nationalists during the Trump era, such as Representative Steve King, Representative Matt Gaetz, Senator Tom Cotton and Senator Josh Hawley.

During the Trump era, commonly identified American nationalist political commentators included Rush Limbaugh, Ann CoulterMichelle MalkinLou DobbsAlex JonesCharlie KirkLaura IngrahamCandace OwensMichael SavageTucker CarlsonMike Cernovich, and Nick Fuentes.

Culture jamming

From Wikipedia, the free encyclopedia

Culture jamming (sometimes also guerrilla communication) is a form of protest used by many anti-consumerist social movements to disrupt or subvert media culture and its mainstream cultural institutions, including corporate advertising. It attempts to "expose the methods of domination" of mass society.

Culture jamming employs techniques originally associated with Letterist International, and later Situationist International known as détournement. It uses the language and rhetoric of mainstream culture to subversively critique the social institutions that produce that culture. Tactics include editing company logos to critique the respective companies, products, or concepts they represent, or wearing fashion statements that criticize the current fashion trends by deliberately clashing with them. Culture jamming often entails using mass media to produce ironic or satirical commentary about itself, commonly using the original medium's communication method. Culture jamming is also a form of subvertising.

Culture jamming aims to highlight and challenge the political assumptions underlying commercial culture, and argues that culture jamming is a response to socially imposed conformity. Prominent examples of culture jamming include the adulteration of billboard advertising by the Billboard Liberation Front and contemporary artists such as Ron English. Culture jamming may involve street parties and protests. While culture jamming usually focuses on subverting or critiquing political and advertising messages, some proponents focus on a different form which brings together artists, designers, scholars, and activists to create works that transcend the status quo rather than merely criticize it. In addition to these political and cultural aims, researchers have noted that the pleasure involved in culture jamming plays an important role in sustaining participation. Analysts argue that fun can link individual acts of subversion to a broader, imagined protest community, helping disparate actions feel part of a shared project. This affective style both draws on and resists the dominant emotion regime of late capitalism, making emotional engagement not only a tactic but also an end in itself.

Origins of the term, etymology, and history

The term was coined by Don Joyce of American sound collage band Negativland, with the release of their album JamCon '84. The phrase "culture jamming" comes from the idea of radio jamming, where public frequencies can be pirated and subverted for independent communication, or to disrupt dominant frequencies used by governments. In one of the tracks of the album, Joyce stated:

As awareness of how the media environment we occupy affects and directs our inner life grows, some resist. The skillfully reworked billboard... directs the public viewer to a consideration of the original corporate strategy. The studio for the cultural jammer is the world at large.

According to Vince Carducci, although the term was coined by Negativland, the practice of culture jamming can be traced as far back as the 1950s. One particularly influential group that was active in Europe was the Situationist International and was led by Guy Debord. The SI asserted that in the past, humans dealt with life and the consumer market directly. They argued that this spontaneous way of life was slowly deteriorating as a direct result of the new "modern" way of life. Situationists saw everything from television to radio as a threat and argued that life in industrialized areas, driven by capitalist forces, had become monotonous, sterile, gloomy, linear, and productivity-driven. In particular, the SI argued humans had become passive recipients of the spectacle, a simulated reality that generates the desire to consume, and positions humans as obedient consumerist cogs within the efficient and exploitative productivity loop of capitalism. Through playful activity, individuals could create situations, the opposite of spectacles. For the SI, these situations took the form of the dérive, or the active drift of the body through space in ways that broke routine and overcame boundaries, creating situations by exiting habit and entering new interactive possibilities.

The cultural critic Mark Dery traces the origins of culture jamming to medieval carnival, which Mikhail Bakhtin interpreted, in Rabelais and His World and other writings, as an officially sanctioned inversion of the social hierarchy. Modern precursors might include: the media-savvy agit-prop of the anti-Nazi photomonteur John Heartfield, the sociopolitical street theater and staged media events of 1960s radicals such as Abbie Hoffman, Joey Skaggs, the German concept of Spaßguerilla, and in the Situationist International (SI) of the 1950s and 1960s. The SI first compared its own activities to radio jamming in 1968, when it proposed the use of guerrilla communication within mass media to sow confusion within the dominant culture. In 1985, the Guerrilla Girls formed to expose discrimination and corruption in the art world.

Dery's 1990 New York Times article on culture jamming, "The Merry Pranksters And the Art of the Hoax" was the first mainstream media mention of the phenomenon; Dery later expanded on this article in his 1993 Open Magazine pamphlet, Culture Jamming: Hacking, Slashing, and Sniping in the Empire of the Signs, a seminal essay that remains the most exhaustive historical, sociopolitical, and philosophical theorization of culture jamming to date. Adbusters, a Canadian publication espousing an environmentalist critique of consumerism and advertising, began promoting aspects of culture jamming after Dery introduced founder and editor Kalle Lasn to the term through a series of articles he wrote for the magazine. In her critique of consumerism, No Logo, the Canadian cultural commentator and political activist Naomi Klein examines culture jamming in a chapter that focuses on the work of Jorge Rodriguez-Gerada. Through an analysis of the Where the Hell is Matt viral videos, researchers Milstein and Pulos analyze how the power of the culture jam to disrupt the status quo is currently being threatened by increasing commercial incorporation. For example, T-Mobile utilized the Liverpool street underground station to host a flashmob to sell their mobile services.

Tactics

Graffitied text on billboard in Cambridge, UK

Culture jamming is a form of disruption that plays on the emotions of viewers and bystanders. Jammers want to disrupt the unconscious thought process that takes place when most consumers view a popular advertising and bring about a détournement. Activists that utilize this tactic are counting on their meme to pull on the emotional strings of people and evoke some type of reaction. The reactions that most cultural jammers are hoping to evoke are behavioral change and political action. There are four emotions that activists often want viewers to feel. These emotions – shock, shame, fear, and anger – are believed to be the catalysts for social change. Culture jamming also intersects with forms of legal transgression. Semiotic disobedience, for example, involves both authorial and proprietary disobedience, while techniques such as coercive disobedience comprise acts of culture jamming combined with a demonstration of the retaliatory actions (legal consequences) handed down by the ruling apparatus.

One of the earliest American culture jammers, Joey Skaggs began staging elaborate media hoaxes in the 1960s as a form of social critique and performance art. His satirical projects, including the Cathouse for Dogs (a fictitious brothel for canines) and the Comacocoon (a fake sensory-deprivation vacation therapy), targeted institutional authority and exposed the media's susceptibility to sensationalism. Skaggs' interventions are regarded as foundational to the development of culture jamming as a strategy for media activism and public discourse.

The basic unit in which a message is transmitted in culture jamming is the meme. Memes are condensed images that stimulate visual, verbal, musical, or behavioral associations that people can easily imitate and transmit to others. The term meme was coined and first popularized by geneticist Richard Dawkins, but later used by cultural critics such as Douglas Rushkoff, who claimed memes were a type of media virus. Memes are seen as genes that can jump from outlet to outlet and replicate themselves or mutate upon transmission, just like a virus.

Culture jammers will often use common symbols such as the McDonald's golden arches or Nike swoosh to engage people and force them to think about their eating habits or fashion sense. In one example, jammer Jonah Peretti used the Nike symbol to stir debate on sweatshop child labor and consumer freedom. Peretti made public exchanges between himself and Nike over a disagreement. Peretti had requested custom Nikes with the word "sweatshop" placed in the Nike symbol. Nike refused. Once this story was made public, it spread worldwide and contributed to the already robust conversation about Nike's use of sweatshops, which had been ongoing for a decade prior to Peretti's 2001 stunt.

Jammers can also organize and participate in mass campaigns. Examples of cultural jamming like Peretti's are more along the lines of tactics that radical consumer social movements would use. These movements push people to question the taken-for-granted assumption that consuming is natural and good and aim to disrupt the naturalization of consumer culture; they also seek to create systems of production and consumption that are more humane and less dominated by global corporate late capitalism.

Past mass events and ideas have included Buy Nothing Day, virtual sit-ins and protests over the Internet, producing ‘subvertisements' and placing them in public spaces, and creating and enacting ‘place jamming' projects where public spaces are reclaimed and nature is re-introduced into urban places.

The most effective form of jamming is to use an already widely recognizable meme to transmit the message. Once viewers are forced to take a second look at the mimicked popular meme they are forced out of their comfort zone. Viewers are presented with another way to view the meme and are forced to think about the implications presented by the jammer. More often than not, when this is used as a tactic the jammer is going for shock value. For example, to make consumers aware of the negative body image that big-name fashion brands are frequently accused of causing, a subvertisement of Calvin Klein's 'Obsession' was created and played worldwide. It depicted a young woman with an eating disorder throwing up into a toilet.

Another way that social consumer movements hope to utilize culture jamming effectively is by employing a metameme. A metameme is a two-level message that punctures a specific commercial image but does so in a way that challenges some larger aspect of the political culture of corporate domination. An example would be the "true cost" campaign set in motion by Adbusters. "True cost" forced consumers to compare the human labor cost and conditions and environmental drawbacks of products to the sales costs. Another example would be the "Truth" campaigns that exposed the deception tobacco companies used to sell their products.

Following critical scholars like Paulo Freire, culture jams are also being integrated into the university classroom setting, "in which students and teachers gain the opportunity not only to learn methods of informed public critique but also to collaboratively use participatory communication techniques to actively create new locations of meaning." For example, students disrupt public space to bring attention to community concerns or utilize subvertisements to engage with media literacy projects.

Examples


Groups

Criticism

Some scholars and activists, such as Amory Starr and Joseph D. Rumbo, have argued that culture jamming is futile because it is easily co-opted and commodified by the market, which tends to "defuse" its potential for consumer resistance. A newer understanding of the term has been called for that would encourage artists, scholars and activists to come together and create innovative, flexible, and practical mobile art pieces that communicate intellectual and political concepts and new strategies and actions.

Cladistics

From Wikipedia, the free encyclopedia

Cladistics (/kləˈdɪstɪks/ klÉ™-DIST-iks; from Ancient Greek κλάδος kládos 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies) that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms worms or fishes were used within a strict cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. Radiation results in the generation of new subclades by bifurcation, but in practice sexual hybridization may blur very closely related groupings.

As a hypothesis, a clade can be rejected only if some groupings were explicitly excluded. It may then be found that the excluded group did actually descend from the last common ancestor of the group, and thus emerged within the group. ("Evolved from" is misleading, because in cladistics all descendants stay in the ancestral group). To keep only valid clades, upon finding that the group is paraphyletic this way, either such excluded groups should be granted to the clade, or the group should be abolished.

Branches down to the divergence to the next significant (e.g. extant) sister are considered stem-groupings of the clade, but in principle each level stands on its own, to be assigned a unique name. For a fully bifurcated tree, adding a group to a tree also adds an additional (named) clade, and a new level on that branch. Specifically, also extinct groups are always put on a side-branch, not distinguishing whether an actual ancestor of other groupings was found.

The techniques and nomenclature of cladistics have been applied to disciplines other than biology. (See phylogenetic nomenclature.)

Cladistics findings are posing a difficulty for taxonomy, where the rank and (genus-)naming of established groupings may turn out to be inconsistent.

Cladistics is now the most commonly used method to classify organisms.

History

Willi Hennig 1972
Peter Chalmers Mitchell in 1920
Robert John Tillyard

The original methods used in cladistic analysis and the school of taxonomy derived from the work of the German entomologist Willi Hennig, who referred to it as phylogenetic systematics (also the title of his 1966 book); but the terms "cladistics" and "clade" were popularized by other researchers. Cladistics in the original sense refers to a particular set of methods used in phylogenetic analysis, although it is now sometimes used to refer to the whole field.

What is now called the cladistic method appeared as early as 1901 with a work by Peter Chalmers Mitchell for birds and subsequently by Robert John Tillyard (for insects) in 1921, and W. Zimmermann (for plants) in 1943. The term "clade" was introduced in 1958 by Julian Huxley after having been coined by Lucien Cuénot in 1940, "cladogenesis" in 1958, "cladistic" by Arthur Cain and Harrison in 1960, "cladist" (for an adherent of Hennig's school) by Ernst Mayr in 1965, and "cladistics" in 1966. Hennig referred to his own approach as "phylogenetic systematics". From the time of his original formulation until the end of the 1970s, cladistics competed as an analytical and philosophical approach to systematics with phenetics and so-called evolutionary taxonomy. Phenetics was championed at this time by the numerical taxonomists Peter Sneath and Robert Sokal, and evolutionary taxonomy by Ernst Mayr.

Originally conceived, if only in essence, by Willi Hennig in a book published in 1950, cladistics did not flourish until its translation into English in 1966 (Lewin 1997). Today, cladistics is the most popular method for inferring phylogenetic trees from morphological data.

In the 1990s, the development of effective polymerase chain reaction techniques allowed the application of cladistic methods to biochemical and molecular genetic traits of organisms, vastly expanding the amount of data available for phylogenetics. At the same time, cladistics rapidly became popular in evolutionary biology, because computers made it possible to process large quantities of data about organisms and their characteristics.

Methodology

The cladistic method interprets each shared character state transformation as a potential piece of evidence for grouping. Synapomorphies (shared, derived character states) are viewed as evidence of grouping, while symplesiomorphies (shared ancestral character states) are not. The outcome of a cladistic analysis is a cladogram – a tree-shaped diagram (dendrogram) that is interpreted to represent the best hypothesis of phylogenetic relationships. Although traditionally such cladograms were generated largely on the basis of morphological characters and originally calculated by hand, genetic sequencing data and computational phylogenetics are now commonly used in phylogenetic analyses, and the parsimony criterion has been abandoned by many phylogeneticists in favor of more "sophisticated" but less parsimonious evolutionary models of character state transformation. Cladists contend that these models are unjustified because there is no evidence that they recover more "true" or "correct" results from actual empirical data sets

Every cladogram is based on a particular dataset analyzed with a particular method. Datasets are tables consisting of molecular, morphological, ethological and/or other characters and a list of operational taxonomic units (OTUs), which may be genes, individuals, populations, species, or larger taxa that are presumed to be monophyletic and therefore to form, all together, one large clade; phylogenetic analysis infers the branching pattern within that clade. Different datasets and different methods, not to mention violations of the mentioned assumptions, often result in different cladograms. Only scientific investigation can show which is more likely to be correct.

Until recently, for example, cladograms like the following have generally been accepted as accurate representations of the ancestral relations among turtles, lizards, crocodilians, and birds:

If this phylogenetic hypothesis is correct, then the last common ancestor of turtles and birds, at the branch near the lived earlier than the last common ancestor of lizards and birds, near the . Most molecular evidence, however, produces cladograms more like this:

If this is accurate, then the last common ancestor of turtles and birds lived later than the last common ancestor of lizards and birds. Since the cladograms show two mutually exclusive hypotheses to describe the evolutionary history, at most one of them is correct.

Cladogram of the primates, showing a monophyletic taxon (a clade: the simians or Anthropoidea, in yellow), a paraphyletic taxon (the prosimians, in blue, including the red patch), and a polyphyletic taxon (the nocturnal primates – the lorises and the tarsiers – in red)

The cladogram to the right represents the current universally accepted hypothesis that all primates, including strepsirrhines like the lemurs and lorises, had a common ancestor all of whose descendants are or were primates, and so form a clade; the name Primates is therefore recognized for this clade. Within the primates, all anthropoids (monkeys, apes, and humans) are hypothesized to have had a common ancestor all of whose descendants are or were anthropoids, so they form the clade called Anthropoidea. The "prosimians", on the other hand, form a paraphyletic taxon. The name Prosimii is not used in phylogenetic nomenclature, which names only clades; the "prosimians" are instead divided between the clades Strepsirhini and Haplorhini, where the latter contains Tarsiiformes and Anthropoidea.

Lemurs and tarsiers may have looked closely related to humans, in the sense of being close on the evolutionary tree to humans. However, from the perspective of a tarsier, humans and lemurs would have looked close, in the exact same sense. Cladistics forces a neutral perspective, treating all branches (extant or extinct) in the same manner. It also forces one to try to make statements, and honestly take into account findings, about the exact historic relationships between the groups.

Terminology for character states

The following terms, coined by Hennig, are used to identify shared or distinct character states among groups:

  • A plesiomorphy ("close form") or ancestral state is a character state that a taxon has retained from its ancestors. When two or more taxa that are not nested within each other share a plesiomorphy, it is a symplesiomorphy (from syn-, "together"). Symplesiomorphies do not mean that the taxa that exhibit that character state are necessarily closely related. For example, Reptilia is traditionally characterized by (among other things) being cold-blooded (i.e., not maintaining a constant high body temperature), whereas birds are warm-blooded. Since cold-bloodedness is a plesiomorphy, inherited from the common ancestor of traditional reptiles and birds, and thus a symplesiomorphy of turtles, snakes and crocodiles (among others), it does not mean that turtles, snakes and crocodiles form a clade that excludes the birds.
  • An apomorphy ("separate form") or derived state is an innovation. It can thus be used to diagnose a clade – or even to help define a clade name in phylogenetic nomenclature. Features that are derived in individual taxa (a single species or a group that is represented by a single terminal in a given phylogenetic analysis) are called autapomorphies (from auto-, "self"). Autapomorphies express nothing about relationships among groups; clades are identified (or defined) by synapomorphies (from syn-, "together"). For example, the possession of digits that are homologous with those of Homo sapiens is a synapomorphy within the vertebrates. The tetrapods can be singled out as consisting of the first vertebrate with such digits homologous to those of Homo sapiens together with all descendants of this vertebrate (an apomorphy-based phylogenetic definition). Importantly, snakes and other tetrapods that do not have digits are nonetheless tetrapods: other characters, such as amniotic eggs and diapsid skulls, indicate that they descended from ancestors that possessed digits which are homologous with ours.
  • A character state is homoplastic or "an instance of homoplasy" if it is shared by two or more organisms but is absent from their common ancestor or from a later ancestor in the lineage leading to one of the organisms. It is therefore inferred to have evolved by convergence or reversal. Both mammals and birds are able to maintain a high constant body temperature (i.e., they are warm-blooded). However, the accepted cladogram explaining their significant features indicates that their common ancestor is in a group lacking this character state, so the state must have evolved independently in the two clades. Warm-bloodedness is separately a synapomorphy of mammals (or a larger clade) and of birds (or a larger clade), but it is not a synapomorphy of any group including both these clades. Hennig's Auxiliary Principle states that shared character states should be considered evidence of grouping unless they are contradicted by the weight of other evidence; thus, homoplasy of some feature among members of a group may only be inferred after a phylogenetic hypothesis for that group has been established.

The terms plesiomorphy and apomorphy are relative; their application depends on the position of a group within a tree. For example, when trying to decide whether the tetrapods form a clade, an important question is whether having four limbs is a synapomorphy of the earliest taxa to be included within Tetrapoda: did all the earliest members of the Tetrapoda inherit four limbs from a common ancestor, whereas all other vertebrates did not, or at least not homologously? By contrast, for a group within the tetrapods, such as birds, having four limbs is a plesiomorphy. Using these two terms allows a greater precision in the discussion of homology, in particular allowing clear expression of the hierarchical relationships among different homologous features.

It can be difficult to decide whether a character state is in fact the same and thus can be classified as a synapomorphy, which may identify a monophyletic group, or whether it only appears to be the same and is thus a homoplasy, which cannot identify such a group. There is a danger of circular reasoning: assumptions about the shape of a phylogenetic tree are used to justify decisions about character states, which are then used as evidence for the shape of the tree. Phylogenetics uses various forms of parsimony to decide such questions; the conclusions reached often depend on the dataset and the methods. Such is the nature of empirical science, and for this reason, most cladists refer to their cladograms as hypotheses of relationship. Cladograms that are supported by a large number and variety of different kinds of characters are viewed as more robust than those based on more limited evidence.

Terminology for taxa

Mono-, para- and polyphyletic taxa can be understood based on the shape of the tree (as done above), as well as based on their character states. These are compared in the table below.


Term Node-based definition Character-based definition
Holophyly, Monophyly A clade, a monophyletic taxon, is a taxon that consists of the last common ancestor and all its descendants. A clade is characterized by one or more apomorphies: derived character states present in the first member of the taxon, inherited by its descendants (unless secondarily lost), and not inherited by any other taxa.
Paraphyly A paraphyletic assemblage is one that is constructed by taking a clade and removing one or more smaller clades. (Removing one clade produces a singly paraphyletic assemblage, removing two produces a doubly paraphyletic assemblage, and so on.) A paraphyletic assemblage is characterized by one or more plesiomorphies: character states inherited from ancestors but not present in all of their descendants. As a consequence, a paraphyletic assemblage is truncated, in that it excludes one or more clades from an otherwise monophyletic taxon. An alternative name is evolutionary grade, referring to an ancestral character state within the group. While paraphyletic assemblages are popular among paleontologists and evolutionary taxonomists, cladists do not recognize paraphyletic assemblages as having any formal information content – they are merely parts of clades.
Polyphyly A polyphyletic assemblage is one which is neither monophyletic nor paraphyletic. A polyphyletic assemblage is characterized by one or more homoplasies: character states which have converged or reverted so as to be the same but which have not been inherited from a common ancestor. No systematist recognizes polyphyletic assemblages as taxonomically meaningful entities, although ecologists sometimes consider them meaningful labels for functional participants in ecological communities (e. g., primary producers, detritivores, etc.).

Criticism

Cladistics, either generally or in specific applications, has been criticized from its beginnings. Decisions as to whether particular character states are homologous, a precondition of their being synapomorphies, have been challenged as involving circular reasoning and subjective judgements. Of course, the potential unreliability of evidence is a problem for any systematic method, or for that matter, for any empirical scientific endeavor at all.

Transformed cladistics arose in the late 1970s  in an attempt to resolve some of these problems by removing a priori assumptions about phylogeny from cladistic analysis, but it has remained unpopular.

Issues

Ancestors

The cladistic method does not identify fossil species as actual ancestors of a clade. Instead, fossil taxa are identified as belonging to separate extinct branches. While a fossil species could be the actual ancestor of a clade, there is no way to know that. Therefore, a more conservative hypothesis is that the fossil taxon is related to other fossil and extant taxa, as implied by the pattern of shared apomorphic features.

Extinction status

An otherwise extinct group with any extant descendants, is not considered (literally) extinct, and for instance does not have a date of extinction.

Hybridization, interbreeding

Due to hybridization, it can be difficult to delineate the two last common ancestors of a given species Many species are capable of interbreeding for millions of years, during which many branches may have radiated and bred with species from earlier branches of the same grouping. The process of true cladistic bifurcation can thus take extended effort as branchings become more complex. In practice, for recent radiations, cladistically guided findings only give a coarse impression of the complexity. A more detailed account will give details about fractions of introgressions between groupings, and even geographic variations thereof. This has been used as an argument for the use of paraphyletic groupings,.

Horizontal gene transfer

Horizontal gene transfer is the mobility of genetic info between different organisms that can have immediate or delayed effects for the reciprocal host. There are several processes in nature which can cause horizontal gene transfer. This does typically not directly interfere with ancestry of the organism, but can complicate the determination of that ancestry. On another level, one can map the horizontal gene transfer processes, by determining the phylogeny of the individual genes using cladistics.

Naming stability

If there is unclarity in mutual relationships, there are a lot of possible trees. Assigning names to each possible clade may not be prudent. Furthermore, established names are discarded in cladistics, or alternatively carry connotations which may no longer hold, such as when additional groups are found to have emerged in them. Naming changes are the direct result of changes in the recognition of mutual relationships, which often is still in flux, especially for extinct species. Hanging on to older naming and/or connotations is counter-productive, as they typically do not reflect actual mutual relationships precisely at all. E.g. Archaea, Asgard archaea, protists, slime molds, worms, invertebrata, fishes, reptilia, monkeys, Ardipithecus, Australopithecus, Homo erectus all contain Homo sapiens cladistically, in their sensu lato meaning. For originally extinct stem groups, sensu lato generally means generously keeping previously included groups, which then may come to include even living species. A pruned sensu stricto meaning is often adopted instead, but the group would need to be restricted to a single branch on the stem. Other branches then get their own name and level. This is commensurate to the fact that more senior stem branches are in fact closer related to the resulting group than the more basal stem branches; that those stem branches only may have lived for a short time does not affect that assessment in cladistics.

In disciplines other than biology

The comparisons used to acquire data on which cladograms can be based are not limited to the field of biology. Any group of individuals or classes that are hypothesized to have a common ancestor, and to which a set of common characteristics may or may not apply, can be compared pairwise. Cladograms can be used to depict the hypothetical descent relationships within groups of items in many different academic realms. The only requirement is that the items have characteristics that can be identified and measured.

Anthropology and archaeology: Cladistic methods have been used to reconstruct the development of cultures or artifacts using groups of cultural traits or artifact features.

Comparative mythology and folktale use cladistic methods to reconstruct the protoversion of many myths. Mythological phylogenies constructed with mythemes clearly support low horizontal transmissions (borrowings), historical (sometimes Palaeolithic) diffusions and punctuated evolution. They also are a powerful way to test hypotheses about cross-cultural relationships among folktales.

Literature: Cladistic methods have been used in the classification of the surviving manuscripts of the Canterbury Tales, and the manuscripts of the Sanskrit Charaka Samhita.

Historical linguistics: Cladistic methods have been used to reconstruct the phylogeny of languages using linguistic features. This is similar to the traditional comparative method of historical linguistics, but is more explicit in its use of parsimony and allows much faster analysis of large datasets (computational phylogenetics).

Textual criticism or stemmatics: Cladistic methods have been used to reconstruct the phylogeny of manuscripts of the same work (and reconstruct the lost original) using distinctive copying errors as apomorphies. This differs from traditional historical-comparative linguistics in enabling the editor to evaluate and place in genetic relationship large groups of manuscripts with large numbers of variants that would be impossible to handle manually. It also enables parsimony analysis of contaminated traditions of transmission that would be impossible to evaluate manually in a reasonable period of time.

Astrophysics infers the history of relationships between galaxies to create branching diagram hypotheses of galaxy diversification.

Human mission to Mars

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