Search This Blog

Saturday, January 1, 2022

Troy

From Wikipedia, the free encyclopedia

Troy
Walls of Troy (2).jpg
Walls of Troy VI.
 
LocationHisarlik, Çanakkale Province, Turkey
RegionTroad
Coordinates39°57′27″N 26°14′20″ECoordinates: 39°57′27″N 26°14′20″E
TypeAncient city
Part ofHistorical National Park of Troia

Troy (Greek: Τροία) or Ilium (Greek: Ίλιον) was an ancient city located at Hisarlik in present-day Turkey, 30 kilometres (19 mi) south-west of Çanakkale. It is known as the setting for the Greek myth of the Trojan War.

In Ancient Greek literature, Troy is portrayed as a powerful kingdom of the Heroic Age, a mythic era when monsters roamed the earth and gods interacted directly with humans. The city was said to have ruled the Troad until the Trojan War led to its complete destruction at the hands of the Greeks. The story of its destruction was one of the cornerstones of Greek mythology and literature, featuring prominently in the Iliad and the Odyssey, as well as numerous other poems and plays. Its legacy played a large role in Greek society, with many prominent families claiming descent from those who had fought there. In the Archaic era, a new city was built at the site where legendary Troy was believed to have stood. In the Classical era, this city became a tourist destination, where visitors would leave offerings to the legendary heroes.

Until the late 19th century, scholars regarded the Trojan War as entirely legendary. However, starting in 1871, Heinrich Schliemann and Frank Calvert excavated the site of the classical era city, under whose ruins they found the remains of numerous earlier settlements. Several of these layers resemble literary depictions of Troy, leading some scholars to conclude that there is a kernel of truth to the legends. Subsequent excavations by others have added to our understanding of the site, though the exact relationship between myth and reality remains unclear.

The archaeological site of Troy consists of nine major layers, the earliest dating from the Early Bronze Age, the latest from the Byzantine era. The mythic city is typically identified with one of the Late Bronze Age layers, such as Troy VI, Troy VIIa, or Troy VIIb. The archaeological site is open to the public as a tourist destination, and was added to the UNESCO World Heritage list in 1998.

Name

In Classical Greek, the city was referred to as both Troia (Τροία) and Ilion ( Ἴλιον) or Ilios ( Ἴλιος). Metrical evidence from the Iliad and the Odyssey suggests that the latter was originally pronounced Wilios. These names may date back to the Bronze Age, as suggested by Hittite records which reference a city in northwest Anatolia called 𒌷𒃾𒇻𒊭 Wilusa or 𒋫𒊒𒄿𒊭 Truwisa; in Greek myth, these names were held to originate from the names of the kingdom's founders, Tros and his son Ilus.

In Latin, the city was referred to as Troia or Ilium. In Turkish, it is known as Truva or Troya.

Legendary Troy

An 18th century depiction of the legendary sack of Troy.

The main literary work set at Troy is the Iliad, an Archaic-era epic poem which tells the story of the final year of the Trojan War. The Iliad portrays Troy as the capital of a rich and powerful kingdom. In the poem, the city appears to be a major regional power capable of summoning numerous allies to defend it.

The city itself is built on a steep hill, protected by enormous sloping stone walls, rectangular towers, and massive gates whose wooden doors can be bolted shut. The city's streets are broad and well-planned. At the top of the hill is the Temple of Athena as well as King Priam's palace, an enormous structure with numerous rooms around an inner courtyard.

In the Iliad, the Achaeans set up their camp near the mouth of the Scamander river, where they beached their ships. The city itself stood on a hill across the plain of Scamander, where much of the fighting takes place.

Besides the Iliad, there are references to Troy in the other major work attributed to Homer, the Odyssey, as well as in other ancient Greek literature (such as Aeschylus's Oresteia). The Homeric legend of Troy was elaborated by the Roman poet Virgil in his Aeneid. The fall of Troy with the story of the Trojan Horse and the sacrifice of Polyxena, Priam's youngest daughter, is the subject of a later Greek epic by Quintus Smyrnaeus ("Quintus of Smyrna").

The Greeks and Romans took for a fact the historicity of the Trojan War and the identity of Homeric Troy with a site in Anatolia on a peninsula called the Troad (Biga Peninsula). Alexander the Great, for example, visited the site in 334 BC and there made sacrifices at tombs associated with the Homeric heroes Achilles and Patroclus.

Archaeological layers

The archaeological site of Troy consists of the hill of Hisarlik and the fields below it to the south. The hill is a tell, composed of strata containing the remains left behind by more than three millennia of human occupation. The primary divisions among layers are designated with Roman numerals, Troy I representing the oldest layer and Troy IX representing the most recent. Sublayers are distinguished with lowercase latters (e.g. VIIa and VIIb) and further subdivisions with numbers (e.g. VIIb1 and VIIb2). An additional major layer known as Troy 0 predates those which were initially given Roman numeral designations.

The layers have been given relative dates by comparing artifacts found in them to those found at other sites. However, precise absolute dates are not always possible due to limitations in the accuracy of C14 dating.


Layer Start End Period
Troy 0 c. 3600-3500 BC 3000 BC Western Anatolian LSA and EB 1 early
Troy I 3000 BC 2550 BC Western Anatolian EB 1 late
Troy II 2500 BC 2300 BC Western Anatolian EB 2
Troy III 2300 BC 2200 BC Western Anatolian EB 3 early
Troy IV 2200 BC 2000 BC Western Anatolian EB 3 middle
Troy V 2000 BC 1750 BC Western Anatolian EB 3 late
Troy VI 1750 BC 1300 BC West. Anat. MBA (Troy VI early)
West. Anat. LBA (Troy VI middle and late)
Troy VIIa 1300 BC 1180 BC Western Anatolian LBA
Troy VIIb 1180 BC 950 BC Western Anatolian LBA - Dark Age
Troy VIII 950 BC 85 BC Classical and Hellenistic Troy
Troy IX 85 BC 500 AD Roman Troy

Troy 0

Troy 0 is a recently discovered layer predating what had previously been the earliest at the site. Remains of the layer, first identified in 2019, are not very substantial and its exact dating remains unclear, although Troy 0 was likely no older than c. 3600-3500 BC. Traces of burns, pottery and wooden beams were found in a layer below the Troy 1 layer, confirming the existence of the Troy 0 layer.

Troy I

Troy I wall

Troy I was a small village founded around 3000 BC. In this era, the site was adjacent to a shallow bay which gradually silted up over the subsequent millenia. The village consisted of stone and mudbrick houses which were attached to one another and surrounded by stone walls. Finds from this layer include dark colored handmade pottery and artifacts made of copper. It had cultural similarities to Aegean sites such as Poliochni and Thermi, as well as to Anatolian sites such as Bademağacı.

Troy II

Troy II was built around 2550 BC. It was twice the size of the preceding city, featuring both a citadel and a lower town. The citadel contained large megaron-style buildings around a courtyard which was likely used for public events such as audiences or religious ceremonies. It was protected by massive stone walls which were topped with mudbrick superstructures. Houses in the lower town were protected by a wooden palisade. Finds from this layer include wheel-made pottery, and numerous items made from precious metals which attest to economic and cultural connections with regions as far as the Balkans and Afghanistan. Troy II was destroyed twice. After the first destruction, the citadel was rebuilt with a dense cluster of small houses. The second destruction took place around 2300 BC, as part of a crisis that affected other sites in the Eastern Mediterranean and the Middle East.

Troy II is notable for having been misidentified as Homeric Troy during initial excavations because of its massive architecture, treasure hordes, and catastrophic destruction. In particular Schliemann saw Homer's description of Troy's Scaean Gate reflected in Troy II's imposing western gate. However, later excavations demonstrated that the site was a thousand years too old to have coexisted with Mycenaean Greeks.

Troy III-V

Schliemann's trench. Layers are marked with Roman numerals.

Troy continued to be occupied between 2300 BC and 1750 BC. However, little is known about these several layers due to Schliemann's reckless excavation practices. In order to fully excavate the citadel of Troy II, he destroyed most remains from this period without first documenting them. These settlements appear to have been smaller and poorer than previous ones, though this interpretation could be merely the result of gaps in the surviving evidence. The settlements included a dense residential neighborhood in the citadel. Walls from Troy II may have been reused as part of Troy III. By the period of Troy V, the city had once again expanded outside the citadel to the west. Troy IV sees the introduction of domed ovens. In Troy V, artifacts include Anatolian-style "red-cross bowls" as well as imported Minoan objects.

Troy VI-VII

Troy VI-VII was a major Late Bronze Age city consisting of a steep fortified citadel and a sprawling lower town below it. It was a thriving coastal city with a considerable population, equal in size to second-tier Hittite settlements. It had a distinct Northwest Anatolian culture and extensive foreign contacts, including with Mycenaean Greece, and its position at the mouth of the Dardanelles has been argued to have given it the function of regional capital, its status protected by treaties.

Aspects of its architecture are consistent with the Iliad's description of mythic Troy, and several of its sublayers (VIh and VIIa) show potential signs of violent destruction. Thus, these sublayers are among the candidates for a potential historical setting of those myths.

Troy VI and VII were given separate labels by early excavators, but current research has shown that the first several sublayers of Troy VII were in fact continuations of the earlier city. Although some scholars have proposed revising the nomenclature to reflect this consensus, the original terms are typically used to avoid confusion.

Troy VI

Troy VI existed from around 1750 BC to 1300 BC. Its citadel was divided into a series of rising terraces, of which only the outermost is reasonably well-preserved. On this terrace, archaeologists have found the remains of freestanding multistory houses where Trojan elites would have lived. These houses lacked ground-floor windows, and their stone exterior walls mirrored the architecture of the citadel fortifications. However, they otherwise display an eclectic mix of architectural styles, some following the classic megaron design, others even having irregular floorplans. Some of these houses show potential Aegean influence, one in particular resembling the megaron at Midea in the Argolid. Archaeologists believe there may have been a royal palace on the highest terrace, but most Bronze Age remains from the top of the hill were cleared away by classical era building projects.

Artist's representation of House VI M, part of the palatial complex

The citadel was enclosed by massive walls. Present-day visitors can see the limestone base of these walls, which are five metres (16 ft) thick and eight metres (26 ft) tall. However, during the Bronze Age they would have been overlaid with wood and mudbrick superstructures, reaching a height over nine metres (30 ft). The walls were built in a "sawtooth" style commonly found at Mycenaean citadels, divided into seven metres (23 ft)-ten metres (33 ft) segments which joined with one another at an angle. The walls also have a notable slope, similar to those at other sites including Hattusa. These walls were watched over by several rectangular watchtowers, which would also have provided a clear view of Trojan plain and the sea beyond it. The citadel was accessed by five gates, which led into paved and drained cobblestone streets. Some of these gates featured enormous pillars which serve no structural purpose and have been interpreted as religious symbols.

The lower town was built to the south of the citadel, covering an area of roughly 30 hectares. Remains of a dense neighborhood have been found just outside the citadel walls, and traces of other buildings and Late Bronze Age pottery have been found further away. Little of it has been excavated, and few remains are likely to exist; buildings in the lower city are likely to have been made of wood and other perishable materials, and much of the area was built over in the classical and Roman era. The extent of the lower town is evidenced by a defensive ditch cut down to the bedrock and postholes which attest to wooden ramparts or walls which would have once been the outer defense of the city.

The lower city was only discovered in the late 1980s, earlier excavators having assumed that Troy VI occupied only the hill of Hisarlik. Its discovery led to a dramatic reassessment of Troy VI, showing that it was over 16 times larger than had been assumed and thus a major city with a large population rather than a mere aristocratic residence.

The material culture of Troy VI appears to belong to a distinct Northwest Anatolian cultural group, with influences from the Anatolia, the Aegean, and the Balkans. The primary local pottery styles were wheel-made West Anatolian Gray Ware and Tan ware, local offshoots of an earlier Middle Helladic tradition. Foreign pottery found at the site includes Minoan, Mycenaean, Cypriot, and Levantine items. Local potters also made their own imitations of foreign styles, including Gray Ware and Tan Ware pots made in Mycenaean-style shapes. Although the city appears to have been within the Hittite sphere of influence, no Hittite artifacts have been found in Troy VI. Also notably absent are sculptures and wall paintings, otherwise common features of Bronze Age cities. Troy VI is also notable for its architectural innovations as well as its cultural developments, which included the first evidence of horses at the site. The language spoken in Troy VI is unknown. The main candidate is Luwian, an Anatolian language which was spoken in many nearby states and which appears in the only piece of writing found at Troy. However, available evidence is not sufficient to establish that Luwian was the primary language of the city's population, and a number of alternatives have been proposed.

Troy VI was destroyed around 1250 BC, corresponding with the sublayer known as Troy VIh. Evidence of Troy VIh's destruction includes collapsed masonry, and subsidence in the southeast of the citadel, which led its initial excavators to conclude that it was destroyed by an earthquake. However, alternative hypotheses include an internal uprising as well as a foreign attack.

Troy VIIa

Troy VIIa was the final layer of the Late Bronze Age city. It was built soon after the destruction of Troy VI, seemingly by its previous inhabitants. The builders reused many of the earlier city's surviving structures, notably its citadel wall, which they renovated with additional stone towers and mudbrick breastworks. Numerous small houses were added inside the citadel, filling in formerly open areas. New houses were also built in the lower city, whose area appears to have been greater in Troy VIIa than in Troy VI. In many of these houses, archaeologists found enormous storage jars called pithoi buried in the ground. Troy VIIa seems to have been built by survivors of Troy VI's destruction, as evidenced by continuity in material culture. However, the character of the city appears to have changed, the citadel growing crowded and foreign imports declining.

The city was destroyed around 1180 BC, roughly contemporary with the Late Bronze Age collapse but subsequent to the destructions of the Mycenaean palaces. The destruction layer shows evidence of enemy attack, including scorch marks.

Troy VIIb

Anatolian Grey Ware

After the destruction of Troy VIIa around 1180 BC, the city was rebuilt as Troy VIIb. Older structures were again reused, including Troy VI's citadel walls. Its first phase, Troy VIIb1, is largely a continuation of Troy VIIa. Residents continued using wheel-made Grey Ware pottery alongside a new handmade style sometimes known as "barbarian ware". Imported Mycenaean-style pottery attests to some continuing foreign trade.

One of the most striking finds from Troy VIIb1 is a hieroglyphic Luwian seal giving the names of a woman and a man who worked as a scribe. The seal is important since it is the only example of preclassical writing found at the site, and provides potential evidence that Troy VIIb1 had a Luwian-speaking population. However, the find is puzzling since palace bureaucracies had largely disappeared by this era. Proposed explanations include the possibility that it belonged to an itinerant freelance scribe and alternatively that it dates from an earlier era than its find context would suggest.

Troy VIIb2 is marked by cultural changes including walls made of upright stones and a handmade knobbed pottery style known as Buckelkeramik. These practices, which existed alongside older local traditions, have been argued to reflect immigrant populations arriving from southwest Europe. Pottery finds from this layer also include imported Protogeometric pottery, showing that Troy was occupied continuously well into the Iron Age, contra later myths.

Troy VIIb was destroyed by fire around 950 BC. However, some houses in the citadel were left intact and the site continued to be occupied, if only sparsely.

Troy VIII-IX

Troy VIII was founded during the Dark Ages and lasted until the Roman era. Though the site had never been entirely abandoned, its redevelopment as a major city was spurred by Greek immigrants who began building around 700 BC. During the Archaic period, the city's defenses once again included the reused citadel wall of Troy VI. Later on, the walls became tourist attraction and sites of worship. Other remains of the Bronze Age city were destroyed by the Greeks' building projects, notably the peak of the citadel where the Troy VI palace is likely to have stood. By the classical era, the city had numerous temples, a theater, among other public buildings, and was once again expanding to the south of the citadel. Troy VIII was destroyed in 85 BC, and subsequently rebuilt as Troy IX. A series of earthquakes devastated the city around 500 AD, though finds from the Late Byzantine era attest to continued habitation at a small scale.

Excavation history

The search for Troy

Alexandria Troas

With the rise of critical history, Troy and the Trojan War were largely consigned to legend. Those who departed from this general view became the first archaeologists at Troy.

Early modern travellers in the 16th and 17th centuries, including Pierre Belon and Pietro Della Valle, had identified Troy with Alexandria Troas, a ruined Hellenistic town approximately 20 kilometres (12 mi) south of Hisarlik. In the late 18th century, Jean Baptiste LeChevalier identified a location near the village of Pınarbaşı, Ezine, a mound approximately 5 kilometres (3.1 mi) south of the currently accepted location. Published in his Voyage de la Troade, it was the most commonly proposed location for almost a century.

In 1822, the Scottish journalist Charles Maclaren was the first to identify with confidence the position of the city as it is now known. In the second half of the 19th century archaeological excavation of the site believed to have been Homeric Troy began.

Frank Calvert

The first excavations at Hisarlik were conducted by Frank Calvert, a Turkish Levantine man of English descent who owned a farm nearby. Calvert made extensive surveys of the site, identifying it with classical era Troy. This identification helped him convince Heinrich Schliemann that Troy was there, and to partner with him in its further excavation.

Heinrich Schliemann

Heinrich Schliemann

In 1868, German businessman Heinrich Schliemann visited Calvert, and secured permission to excavate Hisarlik. Schliemann believed that the literary events of the works of Homer could be verified archaeologically, and he decided to use his wealth to locate it.

Together with Calvert and others, Schliemann began by excavating a trench across the mound of Hisarlik to the depth of the settlements, today called "Schliemann's Trench." In 1871–73 and 1878–79, he discovered the ruins of a series of ancient cities dating from the Bronze Age to the Roman period. He proposed that the second layer, Troy II, corresponded to the city of legend, though later research has shown that it predated the Mycenaean era by several hundred years. Some of the most notable artifacts found by Schliemann are known as Priam's Treasure, after the legendary Trojan king. Schliemann's legacy remains controversial due to his excavation methods, which included removing features he considered insignificant without first studying and documenting them.

Artifacts which Schliemann dubbed Priam's Treasure.
 
Hisarlik, pictured in 1880. The notch at the top is "Schliemann's Trench".

Modern excavations

Wilhelm Dörpfeld

Wilhelm Dörpfeld (1893–94) began excavating the site alongside Schliemann and later inherited management of the site and published his own independent work. His chief contributions were to the study of Troy VI and VII, which Schliemann had overlooked due to his fixation on Troy II. Dörpfeld's interest in these layers was triggered by the need to close a hole in the initial excavators' chronology known as "Calvert's Thousand Year Gap". During his excavation, Dörpfeld came across a section of the Troy VI wall which was weaker than the rest. Since the mythic city had likewise had a weak section of its walls, Dörpfeld became convinced that this layer corresponded to Homeric Troy. Schliemann himself privately agreed that Troy VI was more likely to be the Homeric city, but he never published anything stating so.

University of Cincinnati

Carl Blegen

Carl Blegen, professor at the University of Cincinnati, managed the site 1932–38. These archaeologists, though following Schliemann's lead, added a professional approach not available to Schliemann. He showed that there were at least nine cities. In his research, Blegen came to a conclusion that Troy's nine levels could be further divided into forty-six sublevels, which he published in his main report.

Korfmann

In 1988, excavations were resumed by a team from the University of Tübingen and the University of Cincinnati under the direction of Professor Manfred Korfmann, with Professor Brian Rose overseeing Post-Bronze Age (Greek, Roman, Byzantine) excavation along the coast of the Aegean Sea at the Bay of Troy. Possible evidence of a battle was found in the form of bronze arrowheads and fire-damaged human remains buried in layers dated to the early 12th century BC. The question of Troy's status in the Bronze-Age world has been the subject of a sometimes acerbic debate between Korfmann and the Tübingen historian Frank Kolb in 2001–2002.

Korfmann proposed that the location of the city indicated a commercially oriented economy that would have been at the center of a vibrant trade between the Black Sea, Aegean, Anatolian and Eastern Mediterranean regions. Kolb disputed this thesis, calling it "unfounded" in a 2004 paper. He argued that archaeological evidence shows that economic trade during the Late Bronze Age was quite limited in the Aegean region compared with later periods in antiquity. On the other hand, the Eastern Mediterranean economy was more active during this time, allowing for commercial cities to develop only in the Levant. Kolb also noted the lack of evidence for trade with the Hittite Empire.

One of the major discoveries of these excavations was the Troy VI/VII lower city. This discovery led to a major reinterpretation of the site, which had previously been regarded as a small aristocratic residence rather than a major settlement.

Recent developments

In summer 2006, the excavations continued under the direction of Korfmann's colleague Ernst Pernicka, with a new digging permit.

In 2013, an international team made up of cross-disciplinary experts led by William Aylward, an archaeologist at the University of Wisconsin-Madison, was to carry out new excavations. This activity was to be conducted under the auspices of Çanakkale Onsekiz Mart University and was to use the new technique of "molecular archaeology". A few days before the Wisconsin team was to leave, Turkey cancelled about 100 excavation permits, including Wisconsin's.

In March 2014, it was announced that a new excavation would take place to be sponsored by a private company and carried out by Çanakkale Onsekiz Mart University. This will be the first Turkish team to excavate and is planned as a 12-month excavation led by associate professor Rüstem Aslan. The University's rector stated that "Pieces unearthed in Troy will contribute to Çanakkale’s culture and tourism. Maybe it will become one of Turkey’s most important frequented historical places."

Historical Troy

Troy I-V predate writing and thus study of them falls into the category of prehistoric archaeology. However, Troy emerges into protohistory in the Late Bronze Age, as records mentioning the city begin to appear at other sites. Troy VIII and Troy IX are dated to the historical period and thus are part of history proper.

Troy VI/VII in Hittite records

Troy VI/VII is thought to correspond to the placenames Wilusa and Taruisa known from Hittite records. These correspondences were first proposed in 1924 by Emil Forrer, who also suggested that the name Ahhiyawa corresponds to the Homeric term for the Greeks, Achaeans. These proposals were primarily motivated by linguistic similiarities, since "Taruisa" is a plausible match for the Greek name "Troia" and "Wilusa" likewise for the Greek "Wilios" (later "Ilios"). Subsequent research on Hittite geography has made these identifications more secure, though not all scholars regard them as firmly established.

Wilusa first appears in Hittite records around 1400 BC, when it was one of the twenty-two states of the Assuwa Confederation which unsuccessfully attempted to oppose the Hittite Empire. Circumstantial evidence raises the possibility that the rebellion was supported by the Ahhiyawa. By the late 1300s BC, Wilusa had become politically aligned with the Hittites. Texts from this period mention two kings named Kukkunni and Alaksandu who maintained peaceful relations with the Hittites even as other states in the area did not. Wilusan soldiers may have served in the Hittite army during the Battle of Kadesh. A bit later, Wilusa seems to have experienced the political turmoil suffered by many of its neighbors. References in the Manapa-Tarhunta letter and Tawagalawa letter suggest that a Wilusan king either rebelled or was deposed. This turmoil may have been related to the exploits of Piyamaradu, a Western Anatolian warlord who toppled other pro-Hittite rulers while acting on behalf the Ahhiyawa. However, Piyamaradu is never explicitly identified as the culprit and certain features of the text suggest that he was not. The final reference to Wilusa in the historical record appears in the Milawata letter, in which the Hittite king Tudhaliya IV expresses his intention to reinstall a deposed Wilusan king named Walmu.

In popular writing, these anecdotes have been interpreted as evidence for a historical kernel in myths of the Trojan War. However, scholars have not found historical evidence for any particular event from the legends, and the Hittite documents do not suggest that Wilusa-Troy was ever attacked by Greeks-Ahhiyawa themselves. Noted Hittiteologist Trevor Bryce cautions that our current understanding of Wilusa's history does not provide evidence for there having been an actual Trojan War since "the less material one has, the more easily it can be manipulated to fit whatever conclusion one wishes to come up with".

Classical and Hellenistic Troy (Troy VIII)

In 480 BC, the Persian king Xerxes sacrificed 1,000 cattle at the sanctuary of Athena Ilias while marching through the Hellespontine region towards Greece. Following the Persian defeat in 480–479, Ilion and its territory became part of the continental possessions of Mytilene and remained under Mytilenaean control until the unsuccessful Mytilenean revolt in 428–427. Athens liberated the so-called Actaean cities including Ilion and enrolled these communities in the Delian League. Athenian influence in the Hellespont waned following the oligarchic coup of 411, and in that year the Spartan general Mindaros emulated Xerxes by likewise sacrificing to Athena Ilias. From c. 410–399, Ilion was within the sphere of influence of the local dynasts at Lampsacus (Zenis, his wife Mania, and the usurper Meidias) who administered the region on behalf of the Persian satrap Pharnabazus.

In 399, the Spartan general Dercylidas expelled the Greek garrison at Ilion who were controlling the city on behalf of the Lampsacene dynasts during a campaign which rolled back Persian influence throughout the Troad. Ilion remained outside the control of the Persian satrapal administration at Dascylium until the Peace of Antalcidas in 387–386. In this period of renewed Persian control c. 387–367, a statue of Ariobarzanes, the satrap of Hellespontine Phrygia, was erected in front of the temple of Athena Ilias. In 360–359 the city was briefly controlled by Charidemus of Oreus, a Euboean mercenary leader who occasionally worked for the Athenians. In 359, he was expelled by the Athenian Menelaos son of Arrabaios, whom the Ilians honoured with a grant of proxeny—this is recorded in the earliest civic decree to survive from Ilion. In May 334 Alexander the Great crossed the Hellespont and came to the city, where he visited the temple of Athena Ilias, made sacrifices at the tombs of the Homeric heroes, and made the city free and exempt from taxes. According to the so-called 'Last Plans' of Alexander which became known after his death in June 323, he had planned to rebuild the temple of Athena Ilias on a scale that would have surpassed every other temple in the known world.

Antigonus Monophthalmus took control of the Troad in 311 and created the new city of Antigoneia Troas which was a synoikism of the cities of Skepsis, Kebren, Neandreia, Hamaxitos, Larisa, and Kolonai. In c. 311–306 the koinon of Athena Ilias was founded from the remaining cities in the Troad and along the Asian coast of the Dardanelles and soon after succeeded in securing a guarantee from Antigonus that he would respect their autonomy and freedom (he had not respected the autonomy of the cities which were synoikized to create Antigoneia). The koinon continued to function until at least the 1st century AD and primarily consisted of cities from the Troad, although for a time in the second half of the 3rd century it also included Myrlea and Chalcedon from the eastern Propontis. The governing body of the koinon was the synedrion on which each city was represented by two delegates. The day-to-day running of the synedrion, especially in relation to its finances, was left to a college of five agonothetai, on which no city ever had more than one representative. This system of equal (rather than proportional) representation ensured that no one city could politically dominate the koinon. The primary purpose of the koinon was to organize the annual Panathenaia festival which was held at the sanctuary of Athena Ilias. The festival brought huge numbers of pilgrims to Ilion for the duration of the festival as well as creating an enormous market (the panegyris) which attracted traders from across the region. In addition, the koinon financed new building projects at Ilion, for example a new theatre c. 306 and the expansion of the sanctuary and temple of Athena Ilias in the 3rd century, in order to make the city a suitable venue for such a large festival.

In the period 302–281, Ilion and the Troad were part of the kingdom of Lysimachus, who during this time helped Ilion synoikize several nearby communities, thus expanding the city's population and territory. Lysimachus was defeated at the Battle of Corupedium in February 281 by Seleucus I Nikator, thus handing the Seleucid kingdom control of Asia Minor, and in August or September 281 when Seleucus passed through the Troad on his way to Lysimachia in the nearby Thracian Chersonese Ilion passed a decree in honour of him, indicating the city's new loyalties. In September Seleucus was assassinated at Lysimachia by Ptolemy Keraunos, making his successor, Antiochus I Soter, the new king. In 280 or soon after Ilion passed a long decree lavishly honouring Antiochus in order to cement their relationship with him. During this period Ilion still lacked proper city walls except for the crumbling Troy VI fortifications around the citadel, and in 278 during the Gallic invasion the city was easily sacked. Ilion enjoyed a close relationship with Antiochus for the rest of his reign: for example, in 274 Antiochus granted land to his friend Aristodikides of Assos which for tax purposes was to be attached to the territory of Ilion, and c. 275–269 Ilion passed a decree in honour of Metrodoros of Amphipolis who had successfully treated the king for a wound he received in battle.

Roman Troy (Troy IX)

A new city called Ilium (from Greek Ilion) was founded on the site in the reign of the Roman emperor Augustus. It flourished until the establishment of Constantinople, which became a bishopric in the Roman province Hellespontus (civil Diocese of Asia), but declined gradually in the Byzantine era.

The city was destroyed by Sulla's rival, the Roman general Fimbria, in 85 BC following an eleven-day siege. Later that year when Sulla had defeated Fimbria, he bestowed benefactions on Ilion for its loyalty which helped rebuilding the city. Ilion reciprocated this act of generosity by instituting a new civic calendar which took 85 BC as its first year. However, the city remained in financial distress for several decades despite its favoured status with Rome. In the 80s BC, Roman publicani illegally levied taxes on the sacred estates of Athena Ilias, and the city was required to call on L. Julius Caesar for restitution; while in 80 BC, the city suffered an attack by pirates. In 77 BC the costs of running the annual festival of the koinon of Athena Ilias became too pressing for both Ilion and the other members of the koinon and L. Julius Caesar was once again required to arbitrate, this time reforming the festival so that it would be less of a financial burden. In 74 BC the Ilians once again demonstrated their loyalty to Rome by siding with the Roman general Lucullus against Mithridates VI. Following the final defeat of Mithridates in 63–62, Pompey rewarded the city's loyalty by becoming the benefactor of Ilion and patron of Athena Ilias. In 48 BC, Julius Caesar likewise bestowed benefactions on the city, recalling the city's loyalty during the Mithridatic Wars, the city's connection with his cousin L. Julius Caesar, and the family's claim that they were ultimately descended from Venus through the Trojan prince Aeneas and therefore shared kinship with the Ilians.

In 20 BC, the emperor Augustus visited Ilion and stayed in the house of a leading citizen, Melanippides son of Euthydikos. As a result of his visit, he also financed the restoration and rebuilding of the sanctuary of Athena Ilias, the bouleuterion (council house) and the theatre. Soon after work on the theatre was completed in 12–11 BC, Melanippides dedicated a statue Augustus in the theatre to record this benefaction.

Late Ilium in Church Records

From the 4th century AD until the Byzantine era, Ilium was a suffragan of the provincial capital's Metropolitan Archdiocese of Cyzicus. Several bishops of Troy are historically documented, including one named Orion who participated in the Council of Nicaea in 325 AD. Another named Leucadius was among the heretical bishops who embraced Arianism.

In modern times, Michel d'Herbigny was appointed titular bishop of Ilium. Several others subsequently held the office, though it has been vacant since 1968.

Site conservation

Troy Historical National Park

The west side of Troy Ridge. The road from Tevfikiye enters from the right.

The Turkish government created the Historical National Park at Troy on September 30, 1996. It contains 136 square kilometres (53 sq mi) to include Troy and its vicinity, centered on Troy. The purpose of the park is to protect the historical sites and monuments within it, as well as the environment of the region. In 1998 the park was accepted as a UNESCO World Heritage Site.

In 2015 a Term Development Revision Plan was applied to the park. Its intent was to develop the park into a major tourist site. Plans included marketing research to determine the features most of interest to the public, the training of park personnel in tourism management, and the construction of campsites and facilities for those making day trips. These latter were concentrated in the village of Tevfikiye, which shares Troy Ridge with Troy.

Wooden Trojan Horse monument in the plaza before the modern gate to the ancient city

Public access to the ancient site is along the road from the vicinity of the museum in Tevfikiye to the east side of Hisarlik. Some parking is available. Typically visitors come by bus, which disembarks its passengers into a large plaza ornamented with flowers and trees and some objects from the excavation. In its square is a large wooden horse monument, with a ladder and internal chambers for use of the public. Bordering the square is the gate to the site. The public passes through turnstiles. Admission is usually not free. Within the site, the visitors tour the features on dirt roads or for access to more precipitous features on railed boardwalks. There are many overlooks with multilingual boards explaining the feature. Most are outdoors, but a permanent canopy covers the site of an early megaron and wall.

UNESCO World Heritage Site

The archaeological site of Troy was added to the UNESCO World Heritage list in 1998 in recognition of its historical, cultural, and scientific significance.

Troy Museum

Troy Museum subterranean interior.
 
Troy Museum aboveground. Most of the entire field in which it sits roofs the underground galleries, work, and storage spaces. These are accessed via ramps not shown. There are also outdoor display spaces.

In 2018 the Troy Museum (Turkish Troya Müzesi) was opened at Tevfikiye village 800 metres (870 yd) east of the excavation. A design contest for the architecture had been won by Yalin Mimarlik in 2011. The cube-shaped building with extensive underground galleries holds more than 40,000 portable artifacts, 2000 of which are on display. Artifacts were moved here from a few other former museums in the region. The range is the entire prehistoric Troad. Displays are multi-lingual. In many cases the original contexts are reproduced.

Alternative location hypotheses

A small number of independent researchers doubt the site of Homeric Troy is Hisarlik and favour an alternative location. Smith (2022) has argued Homeric Troy was in fact the Bronze Age fortified mound Yenibademli Höyük on Gökçeada.

In 2008, Raoul Schrott proposed Troy is the ancient fortress Karatepe in Cilicia. His idea has been criticised by German archaeologists.

Mastodon

From Wikipedia, the free encyclopedia

Mastodon
Temporal range: Early PlioceneLate Pleistocene, 5.3–0.011 Ma
Mammut americanum.jpg
Mounted M. americanum skeleton (the "Warren mastodon"), AMNH
Scientific classification 
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: Mammutidae
Genus: Mammut
Blumenbach, 1799
Type species
Elephas americanum
Kerr, 1792
Species
  • M. americanum (Kerr, 1792)
  • M. cosoensis Schultz, 1937
  • M. matthewi Osborn, 1921
  • M. pacificus Dooley et al., 2019
  • M. raki Frick, 1933
Mammut distribution.svg
The inferred range of Mammut (Eurasian range includes that of Zygolophodon borsoni, whose genus assignment is uncertain, and M. matthewi)
Synonyms
  • Mastodon Cuvier, 1817
  • Tetracaulodon Godman, 1830
  • Missourium Koch, 1840
  • Leviathan Koch, 1841 (Emend. Koch, 1843)
  • Pliomastodon Osborn, 1926

A mastodon (mastós 'breast' + odoús 'tooth') is any proboscidean belonging to the extinct genus Mammut (family Mammutidae) that inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the Pleistocene 10,000 to 11,000 years ago. Mastodons lived in herds and were predominantly forest-dwelling animals. They survived on a mixed diet and obtained food by browsing and grazing, somewhat similar to modern elephants, but probably with greater emphasis on browsing

M. americanum, the American mastodon, and M. pacificus, the Pacific mastodon, are the youngest and best-known species of the genus. Mastodons disappeared from North America as part of a mass extinction of most of the Pleistocene megafauna, widely believed to have been caused by a combination of climate changes at the end of the Pleistocene combined with overexploitation by Clovis hunters.

History

Exhuming the First American Mastodon, 1806 painting by Charles Willson Peale

A Dutch tenant farmer found the first recorded remnant of Mammut, a tooth some 2.2 kg (5 lb) in weight, in the village of Claverack, New York, in 1705. The mystery animal became known as the "incognitum". In 1739 French soldiers at present-day Big Bone Lick State Park, Kentucky, found the first bones to be collected and studied scientifically. They carried them to the Mississippi River, from where they were transported to the National Museum of Natural History in Paris. Similar teeth were found in South Carolina, and some of the African slaves there supposedly recognized them as being similar to the teeth of African elephants. There soon followed discoveries of complete bones and tusks in Ohio. People started referring to the "incognitum" as a "mammoth", like the ones that were being dug out in Siberia – in 1796 the French anatomist Georges Cuvier proposed the radical idea that mammoths were not simply elephant bones that had been somehow transported north, but a species which no longer existed. Johann Friedrich Blumenbach assigned the scientific name Mammut to the American "incognitum" remains in 1799, under the assumption that they belonged to mammoths. Other anatomists noted that the teeth of mammoths and elephants differed from those of the "incognitum", which possessed rows of large conical cusps, indicating that they were dealing with a distinct species. In 1817 Cuvier named the "incognitum" Mastodon.

Cuvier assigned the name mastodon (or mastodont) - meaning "breast tooth" (Ancient Greek: μαστός "breast" and ὀδούς, "tooth"), - for the nipple-like projections on the crowns of the molars.

Taxonomy

Mastodon as a genus name is obsolete; the valid name is Mammut, as that name preceded Cuvier's description, making Mastodon a junior synonym. The change was met with resistance, and authors sometimes applied "Mastodon" as an informal name; consequently it became the common term for members of the genus.

Species include:

  • M. americanum, the American mastodon, is one of the best known and among the last species of Mammut. Its earliest occurrences date from the early-middle Pliocene (early Blancan stage). It was formerly regarded (see below) as having a continent-wide distribution, especially during the Pleistocene epoch, known from fossil sites ranging from present-day Alaska, Ontario and New England in the north, to Florida, southern California, and as far south as Honduras. The American mastodon has been widely thought to have resembled a woolly mammoth in appearance. However, consideration of the long tail (usually present in animals living in warm climates), size, body mass and environment implies the animal was not similarly hairy, and there is scant preserved evidence of body hair (what little has been recovered suggests a semiaquatic lifestyle). It had tusks that sometimes exceeded 5 m (16 ft) in length; they curved upwards, but less dramatically than those of the woolly mammoth. Its main habitat was cold spruce woodlands, and it is believed to have browsed in herds. It became extinct at the end of the Pleistocene approximately 11,000 years ago.
  • M. matthewi — found in the Snake Creek Formation of Nebraska, dating to the late Hemphillian. Some authors consider it practically indistinguishable from M. americanum. There is one report of it in China.
  • M. pacificus — based on a 2019 analysis, Pleistocene specimens from California and southern Idaho have been transferred from M. americanum to this new species. It differs from the eastern population in having narrower molars, six as opposed to five sacral vertebrae, a thicker femur, and a consistent absence of mandibular tusks.
  • M. raki — Its remains were found in the Palomas Formation, near Truth or Consequences, New Mexico, dating from the early-middle Pliocene, between 4.5 and 3.6 Ma. It coexisted with Equus simplicidens and Gigantocamelus and differs from M. americanum in having a relatively longer and narrower third molar, similar to the description of the defunct genus Pliomastodon, which supports its arrangement as an early species of Mammut. However, like M. matthewi, some authors do not consider it sufficiently distinct from M. americanum to warrant its own species.
  • M. cosoensis — found in the Coso Formation of California, dating to the Late Pliocene, originally a species of Pliomastodon, it was later assigned to Mammut.

Since a tentative 1977 report of M. matthewi in China, there have been no reports of currently recognized Mammut species outside of North America according to Paleobiology database (which does not recognize M. borsoni). However, the status of Mammut or Zygolophodon borsoni in the literature appears equivocal.

Evolution

Comparison of woolly mammoth (L) and American mastodon (R)
 
Excavation of a specimen in a golf course in Heath, Ohio, 1989

Mammut is a genus of the extinct family Mammutidae, closely related to the proboscidean family Elephantidae (mammoths and elephants), from which it originally diverged approximately twenty-seven million years ago. The following cladogram shows the placement of the American mastodon among other proboscideans, based on hyoid characteristics:

Mammut americanum (American mastodon)

Gomphotherium sp.

Stegodon zdanskyi

Loxodonta africana (African elephant)

Elephas maximus (Asian elephant)

Mammuthus columbi (Columbian mammoth)

Over the years, several fossils from localities in North America, Africa and Asia have been attributed to Mammut, but only the North American remains have been named and described, one of them being M. furlongi, named from remains found in the Juntura Formation of Oregon, dating from the late Miocene. However, it is no longer considered valid, leaving only five valid species.

A complete mitochondrial DNA (mtDNA) sequence has been obtained from the tooth of an M. americanum skeleton found in permafrost in northern Alaska. The remains are thought to be 50,000 to 130,000 years old. This sequence has been used as an outgroup to refine divergence dates in the evolution of the Elephantidae. The rate of mtDNA sequence change in proboscideans was found to be significantly lower than in primates.

A 2020 analysis of mtDNA from American mastodon remains collected in eastern Beringia indicated they belonged to two genetically divergent clades. The clades were dated to different interglacials, suggesting a repeating pattern of colonization during an interglacial followed by extirpation during the subsequent glacial advance. The Beringian clades had less genetic diversity than populations present south of the ice sheets, suggesting they were founded by relatively small migrating populations.

Description

Restoration of an American mastodon

Modern reconstructions based on partial and skeletal remains reveal that mastodons were very similar in appearance to elephants and, to a lesser degree, mammoths, though not closely related to either one. Compared to mammoths, mastodons had shorter legs and a longer body and were more heavily muscled, a build similar to that of the current Asian elephants. The average body size of the species M. americanum was around 2.3 m (7 ft 7 in) in height at the shoulders, corresponding to a large female or a small male; large males were up to 2.8 m (9 ft 2 in) in height. Among the largest male specimens, the 35-year-old AMNH 9950 was 2.89 m (9.5 ft) tall and weighed 7.8 tonnes (7.7 long tons; 8.6 short tons), while another was 3.25 m (10.7 ft) tall and weighed 11 tonnes (11 long tons; 12 short tons).

American mastodon molars at the State Museum of Pennsylvania

As in modern elephants, the females were smaller than the males. They had a low and long skull with long curved tusks, with those of the males being more massive and more strongly curved. Mastodons had cusp-shaped teeth, very different from mammoth and elephant teeth (which have a series of enamel plates), well-suited for chewing leaves and branches of trees and shrubs.

Mastodons are typically depicted with a thick woolly mammoth-like coat of hair, but there is no preserved evidence for this.

Paleobiology

Social behavior

M. pacificus female and calf at the George C. Page Museum

Based on the characteristics of mastodon bone sites, it can be inferred that, as in modern proboscideans, the mastodon social group consisted of adult females and young, living in bonded groups called mixed herds. The males abandoned the mixed herds once reaching sexual maturity and lived either alone or in male bond groupings. As in modern elephants, there probably was no seasonal synchrony of mating activity, with both males and females seeking out each other for mating when sexually active.

M. americanum male and female, University of Michigan

Diet

Mastodons have been characterized as predominantly browsing animals. Of New World proboscids, they appear to have been the most consistent in browsing rather than grazing, consuming C3 as opposed to C4 plants, and in occupying closed forests versus more open habitats. This dietary inflexibility may have prevented them from invading South America during the Great American Interchange, due to the need to cross areas of grassland to do so. Most accounts of gut contents have identified coniferous twigs as the dominant element in their diet. Other accounts (e.g., the Burning Tree mastodon) have reported no coniferous content and suggest selective feeding on low, herbaceous vegetation, implying a mixed browsing and grazing diet, with evidence provided by studies of isotopic bone chemistry indicating a seasonal preference for browsing. Study of mastodon teeth microwear patterns indicates that mastodons could adjust their diet according to the ecosystem, with regionally specific feeding patterns corresponding to boreal forest versus cypress swamps, while a population at a given location was sometimes able to maintain its dietary niche through changes in climate and browse species availability.

Distribution and habitat

Restoration of an American mastodon herd by Charles R. Knight

The range of most species of Mammut is unknown as their occurrences are restricted to few localities, the exception being the American mastodon (M. americanum), which is one of the most widely distributed Pleistocene proboscideans in North America. M. americanum fossil sites range in time from the Blancan to Rancholabrean faunal stages and in locations from as far north as Alaska, as far east as Florida, and as far south as the state of Puebla in central Mexico, with an isolated record from Honduras, probably reflecting the results of the maximum expansion achieved by the American mastodon during the Late Pleistocene. A few isolated reports tell of mastodons being found along the east coast up to the New England region, with high concentrations in the Mid-Atlantic region. There is strong evidence indicating that the members of Mammut were forest dwelling proboscideans, predominating in woodlands and forests, and browsed on trees and shrubs. They apparently did not disperse southward to South America, it being speculated that this was because of a dietary specialization on a particular type of vegetation.

Extinction

Fossil evidence indicates that mastodons probably disappeared from North America about 10,500 years ago as part of a mass extinction of most of the Pleistocene megafauna that is widely believed to have been a result of human hunting pressure. The latest Paleo-Indians entered the Americas and expanded to relatively large numbers 13,000 years ago, and their hunting may have caused a gradual attrition of the mastodon population. Analysis of tusks of mastodons from the American Great Lakes region over a span of several thousand years prior to their extinction in the area shows a trend of declining age at maturation; this is contrary to what one would expect if they were experiencing stresses from an unfavorable environment, but is consistent with a reduction in intraspecific competition that would result from a population being reduced by human hunting.

On the other hand, environmental DNA sequencing indicates that disappearance of megafaunal DNA in North America correlates in time with major changes in plant DNA, suggesting a key role of climate change. Modeling based on the whole of the proboscid fossil record also suggests climate was the more important factor, though with human hunting imposing a "double jeopardy" on mastodons and their kin.

 

Proboscidea

From Wikipedia, the free encyclopedia

Proboscidea
Temporal range: Middle Paleocene-Holocene 60.0–0 Ma
AfricanElephant.jpg
African bush elephant, Loxodonta africana
Scientific classification 
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Clade: Tethytheria
Order: Proboscidea
Illiger, 1811
Subclades

The Proboscidea (/prɒbəˈsɪdə/, from the Greek προβοσκίς and the Latin proboscis) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From the mid-Miocene onwards, most proboscideans were very large. The largest land mammal of all time may have been a proboscidean; Palaeoloxodon namadicus was up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t (24.3 short tons), almost double the weight of some sauropods like Diplodocus carnegii. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t (11.5 short tons). In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.

Elephants are the largest existing land animals. Three species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. Elephantidae is the only surviving family of the order Proboscidea; extinct members include the mastodons. The family Elephantidae also contains several extinct groups, including the mammoths and straight-tusked elephants. African elephants have larger ears and concave backs, whereas Asian elephants have smaller ears, and convex or level backs. Distinctive features of all elephants include a long proboscis called a trunk, tusks, large ear flaps, massive legs, and tough but sensitive skin. The trunk is used for breathing, bringing food and water to the mouth, and grasping objects. Tusks, which are derived from the incisor teeth, serve both as weapons and as tools for moving objects and digging. The large ear flaps assist in maintaining a constant body temperature as well as in communication. The pillar-like legs carry their great weight.

Evolution

The earliest known proboscidean is Eritherium, followed by Phosphatherium, a small animal about the size of a fox. Both date from late Paleocene deposits of Morocco.

Proboscideans evolved in Africa, where they increased in size and diversity during the Eocene and early Oligocene. Proboscideans have evolved greatly over time through three major forms of radiation: radiation of primitive Lophodont forms, radiation of gomphotheres and stegodons, and radiation of elepphantidae. These forms of radiation have illustrated that proboscideans characteristics such as trunk, large ears, tusks, flaps, and huge ears have evolved and were appearing late in the modern form. Several primitive families from these epochs have been described, including the Numidotheriidae, Moeritheriidae, and Barytheriidae, all found exclusively in Africa. The Anthracobunidae from the Indian subcontinent were also believed to be a family of proboscideans, but were excluded from the Proboscidea by Shoshani and Tassy (2005) and have more recently been assigned to the Perissodactyla. When Africa became connected to Europe and Asia after the shrinking of the Tethys Sea, proboscideans migrated into Eurasia, with some families eventually reaching the Americas. Proboscideans found in Eurasia as well as Africa include the Deinotheriidae, which thrived during the Miocene and into the early Quaternary, Stegolophodon, an early genus of the disputed family Stegodontidae; the highly diverse Gomphotheriidae and Amebelodontidae; and the Mammutidae, or mastodons.

Most proboscideans are now extinct, including all species endemic to the Americas, Europe, and northern Asia. Many of these extinctions occurred during or shortly after the last glacial period. Recently extinct species include the gomphotheres in the Americas, the American mastodon of family Mammutidae in North America, numerous stegodonts in Asia, the mammoths throughout the Northern Hemisphere, and several species of dwarf elephants found on various islands scattered around the world.

Classification

[[DJS -- see Wiki article above for an unranked taxonomy of proboscidean genera as of 2019 --]]

Elephantidae

From Wikipedia, the free encyclopedia
 
Elephantidae
Temporal range: Pliocene–Holocene
Elephas maximus (Bandipur).jpg
A male Asian elephant (Elephas maximus) in the wild at Bandipur National Park in India
Scientific classification 
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Superfamily: Elephantoidea
Family: Elephantidae
Gray, 1821
Type genus
Elephas
Genera
Synonyms
  • Elephasidae Lesson, 1842

Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are terrestrial large mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta (African elephants) and Elephas (Asiatic elephants), are living.

The family was first described by John Edward Gray in 1821, and later assigned to taxonomic ranks within the order Proboscidea. Elephantidae has been revised by various authors to include or exclude other extinct proboscidean genera.

Classification

"Man, and the elephant" plate from Hawkins' A comparative view of the human and animal frame, 1860
 

Scientific classification of Elephantidae taxa embraces an extensive record of fossil specimens, over millions of years, some of which existed until the end of the last ice age. Some species were extirpated more recently. The discovery of new specimens and proposed cladistics have resulted in systematic revisions of the family and related proboscideans.

Elephantids are classified informally as the elephant family, or in a paleobiological context as elephants and mammoths. The common name elephant primarily refers to the living taxa, the modern elephants, but may also refer to a variety of extinct species, both within this family and in others. Other members of the Elephantidae, especially members of the genus Mammuthus, are commonly called mammoths.

The family diverged from a common ancestor of the mastodons of Mammutidae. The classification of proboscideans is unstable and has been frequently revised.

The following cladogram shows the placement of the genus Mammuthus among other proboscideans, based on a 2007 study of hyoid characteristics:

Elephantidae
Elephantinae
Loxodontini

Loxodonta (2 species) Elephas africanus - 1700-1880 - Print - Iconographia Zoologica - (white background).jpg

  (African elephants)  
Elephantini
Palaeoloxodontina        

PalaeoloxodonElephas-antiquus.jpg

Elephantina
Elephas

(3~6 subspecies) Indian elephant white background.jpg

  (Asian elephants)  

Mammuthus trogontherii122DB.jpg

  (Mammoths)  
The most accurate phylogenetic tree of the elephants and mammoths as of 2010

However, a 2017 study of mitochondrial and nuclear DNA placed Palaeoloxodon as more closely related to Loxodonta (in particular, the African forest elephant) than to Elephantina.

The systematics of the living subspecies and species of the modern elephants has undergone several revisions. A list of the extant Elephantidae includes:

Elephantidae
Loxodonta (African)
L. africana African bush elephant
L. cyclotis African forest elephant

Elephas (Asiatic)
E. maximus Asian elephant
E. m. maximus Sri Lankan elephant
E. m. indicus Indian elephant
E. m. sumatranus Sumatran elephant
E. m. borneensis Borneo elephant

Evolutionary history

Evolution of elephants from the ancient Eocene (bottom) to the modern day (top)

Although the fossil evidence is uncertain, by comparing genes, scientists have discovered evidence that elephantids and other proboscideans share a distant ancestry with Sirenia (sea cows) and Hyracoidea (hyraxes). These have been assigned, along with the extinct demostylians and embrithopods, to the clade Paenungulata. In the distant past, members of the various hyrax families grew to large sizes, and the common ancestor of all three modern families is thought to have been some kind of amphibious hyracoid. One hypothesis is that these animals spent most of their time under water, using their trunks like snorkels for breathing. Modern elephants have this ability and are known to swim in that manner for up to six hours and 50 km (31 mi).

In the past, a much wider variety of genera and species was found, including the mammoths and stegodons.

Algorithmic information theory

From Wikipedia, the free encyclopedia https://en.wikipedia.org/wiki/Algorithmic_information_theory ...