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Wednesday, September 20, 2023

Neocortex

From Wikipedia, the free encyclopedia
Neocortex
A representative column of neocortex. Cell body layers are labeled on the left, and fiber layers are labeled on the right.

The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, spatial reasoning and language. The neocortex is further subdivided into the true isocortex and the proisocortex.

In the human brain, the cerebral cortex consists of the larger neocortex and the smaller allocortex. The neocortex is made up of six layers, labelled from the outermost inwards, I to VI.

Etymology

The term is from cortex, Latin, "bark" or "rind", combined with neo-, Greek, "new". Neopallium is a similar hybrid, from Latin pallium, "cloak". Isocortex and allocortex are hybrids with Greek isos, "same", and allos, "other".

Anatomy

The neocortex is the most developed in its organisation and number of layers, of the cerebral tissues. The neocortex consists of the grey matter, or neuronal cell bodies and unmyelinated fibers, surrounding the deeper white matter (myelinated axons) in the cerebrum. This is a very thin layer though, about 2–4 mm thick. There are two types of cortex in the neocortex, the proisocortex and the true isocortex. The pro-isocortex is a transitional area between the true isocortex and the periallocortex (part of the allocortex). It is found in the cingulate cortex (part of the limbic system), in Brodmann's areas 24, 25, 30 and 32, the insula and the parahippocampal gyrus.

Of all the mammals studied to date (including humans), a species of oceanic dolphin known as the long-finned pilot whale has been found to have the most neocortical neurons.

Geometry

The neocortex is smooth in rodents and other small mammals, whereas in elephants, dolphins and primates and other larger mammals it has deep grooves (sulci) and ridges (gyri). These folds allow the surface area of the neocortex to be greatly increased. All human brains have the same overall pattern of main gyri and sulci, although they differ in detail from one person to another. The mechanism by which the gyri form during embryogenesis is not entirely clear, and there are several competing hypotheses that explain gyrification, such as axonal tension, cortical buckling or differences in cellular proliferation rates in different areas of the cortex.

Layers

The neocortex contains both excitatory (~80%) and inhibitory (~20%) neurons, named for their effect on other neurons. The human neocortex consists of hundreds of different types of cells. The structure of the neocortex is relatively uniform (hence the alternative names "iso-" and "homotypic" cortex), consisting of six horizontal layers segregated principally by cell type and neuronal connections. However, there are many exceptions to this uniformity; for example, layer IV is small or missing in the primary motor cortex. There is some canonical circuitry within the cortex; for example, pyramidal neurons in the upper layers II and III project their axons to other areas of neocortex, while those in the deeper layers V and VI often project out of the cortex, e.g. to the thalamus, brainstem, and spinal cord. Neurons in layer IV receive the majority of the synaptic connections from outside the cortex (mostly from thalamus), and themselves make short-range, local connections to other cortical layers. Thus, layer IV is the main recipient of incoming sensory information and distributes it to the other layers for further processing.

Cortical columns

The neocortex is often described as being arranged in vertical structures called cortical columns, patches of neocortex with a diameter of roughly 0.5 mm (and a depth of 2 mm, i.e., spanning all six layers). These columns are often thought of as the basic repeating functional units of the neocortex, but their many definitions, in terms of anatomy, size, or function, are generally not consistent with each other, leading to a lack of consensus regarding their structure or function or even whether it makes sense to try to understand neocortex in terms of columns.

Function

The neocortex is derived embryonically from the dorsal telencephalon, which is the rostral part of the forebrain. The neocortex is divided, into regions demarcated by the cranial sutures in the skull above, into frontal, parietal, occipital, and temporal lobes, which perform different functions. For example, the occipital lobe contains the primary visual cortex, and the temporal lobe contains the primary auditory cortex. Further subdivisions or areas of neocortex are responsible for more specific cognitive processes. In humans, the frontal lobe contains areas devoted to abilities that are enhanced in or unique to our species, such as complex language processing localized to the ventrolateral prefrontal cortex (Broca's area). In humans and other primates, social and emotional processing is localized to the orbitofrontal cortex.

The neocortex has also been shown to play an influential role in sleep, memory and learning processes. Semantic memories appear to be stored in the neocortex, specifically the anterolateral temporal lobe of the neocortex. It is also involved in instrumental conditioning; responsible for transmitting sensory information and information about plans for movement to the basal ganglia. The firing rate of neurons in the neocortex also has an effect on slow-wave sleep. When the neurons are at rest and are hyperpolarizing, a period of inhibition occurs during a slow oscillation, called the down state. When the neurons of the neocortex are in the excitatory depolarizing phase and are firing briefly at a high rate, a period of excitation occurs during a slow oscillation, called the up state.

Clinical significance

Lesions that develop in neurodegenerative disorders, such as Alzheimer's disease, interrupt the transfer of information from the sensory neocortex to the prefrontal neocortex. This disruption of sensory information contributes to the progressive symptoms seen in neurodegenerative disorders such as changes in personality, decline in cognitive abilities, and dementia. Damage to the neocortex of the anterolateral temporal lobe results in semantic dementia, which is the loss of memory of factual information (semantic memories). These symptoms can also be replicated by transcranial magnetic stimulation of this area. If damage is sustained to this area, patients do not develop anterograde amnesia and are able to recall episodic information.

Evolution

The neocortex is the newest part of the cerebral cortex to evolve (hence the prefix neo meaning new); the other part of the cerebral cortex is the allocortex. The cellular organization of the allocortex is different from the six-layered neocortex. In humans, 90% of the cerebral cortex and 76% of the entire brain is neocortex.

For a species to develop a larger neocortex, the brain must evolve in size so that it is large enough to support the region. Body size, basal metabolic rate and life history are factors affecting brain evolution and the coevolution of neocortex size and group size. The neocortex increased in size in response to pressures for greater cooperation and competition in early ancestors. With the size increase, there was greater voluntary inhibitory control of social behaviors resulting in increased social harmony.

The six-layer cortex appears to be a distinguishing feature of mammals; it has been found in the brains of all mammals, but not in any other animals. There is some debate, however, as to the cross-species nomenclature for neocortex. In avians, for instance, there are clear examples of cognitive processes that are thought to be neocortical in nature, despite the lack of the distinctive six-layer neocortical structure. In a similar manner, reptiles, such as turtles, have primary sensory cortices. A consistent, alternative name has yet to be agreed upon.

Neocortex ratio

The neocortex ratio of a species is the ratio of the size of the neocortex to the rest of the brain. A high neocortex ratio is thought to correlate with a number of social variables such as group size and the complexity of social mating behaviors. Humans have a large neocortex as a percentage of total brain matter when compared with other mammals. For example, there is only a 30:1 ratio of neocortical gray matter to the size of the medulla oblongata in the brainstem of chimpanzees, while the ratio is 60:1 in humans.

Dominion

From Wikipedia, the free encyclopedia
https://en.wikipedia.org/wiki/Dominion

A Dominion was any of several self-governing nations of the British Empire. With the evolution of the British Empire into the Commonwealth of Nations, the dominions became independent states.

"Dominion status" was first accorded to Canada, Australia, New Zealand, Newfoundland, South Africa, and the Irish Free State at the 1926 Imperial Conference through the Balfour Declaration of 1926, recognising Great Britain and the Dominions as "autonomous communities within the British Empire, equal in status, in no way subordinate one to another in any aspect of their domestic or external affairs, though united by a common allegiance to the Crown and freely associated as members of the British Commonwealth of Nations". Their full legislative independence was subsequently confirmed in the 1931 Statute of Westminster. Later India, Pakistan, and Ceylon (now Sri Lanka) also became dominions, for short periods of time.

With the transition of the British Empire into the Commonwealth of Nations after World War II, it was decided that the term Commonwealth country should formally replace dominion for official Commonwealth usage. This decision was made during the 1949 Commonwealth Prime Ministers' Conference when India was intending to become a republic, so that both types of governments could become and remain full members of the Commonwealth, and this term hence refers to the autonomous dominions and republics.

After this, the term dominion without its legal dimension stayed in use for thirty more years for Commonwealth countries which had the crown as head of state, before gradually, particularly after 1953, being replaced by the term realm, as equal realms of the crown of the Commonwealth.

Definition

Map of the British Empire under Queen Victoria at the end of the nineteenth century. "Dominions" refers to all territories belonging to the Crown.

The term dominion means "that which is mastered or ruled". It was used by the British to describe their colonies or territorial possessions.

Use of dominion to refer to a particular territory within the British Empire dates back to the 16th century and was sometimes used to describe Wales from 1535 to around 1800: for instance, the Laws in Wales Act 1535 applies to "the Dominion, Principality and Country of Wales". Dominion, as an official title, was conferred on the Colony of Virginia about 1660 and on the Dominion of New England in 1686.

Under the British North America Act 1867, the partially self-governing colonies of British North America were united into the Dominion of Canada. The new federal and provincial governments split considerable local powers, but Britain retained overall legislative supremacy. At the Colonial Conference of 1907, the self-governing polities of Canada and the Commonwealth of Australia were referred to collectively as Dominions for the first time. Two other self-governing colonies—New Zealand and Newfoundland—were granted the status of Dominion in the same year. These were followed by the Union of South Africa in 1910. The Order in Council annexing the island of Cyprus in 1914 declared that, from 5 November 1914, the island "shall be annexed to and form part of His Majesty's dominions".

King George V (front, centre) and his Prime Ministers at the 1926 Imperial Conference.

Dominion status was formally accorded to Canada, Australia, New Zealand, Newfoundland, South Africa, and the Irish Free State at the 1926 Imperial Conference to designate "autonomous communities within the British Empire, equal in status, in no way subordinate one to another in any aspect of their domestic or external affairs, though united by a common allegiance to the Crown and freely associated as members of the British Commonwealth of Nations". The British Government of Lloyd George had emphasised the use of the capital "D" when referring to the Irish Free State in the Anglo-Irish Treaty to assure it the same constitutional status in order to avoid confusion with the wider term "His Majesty's dominions", which referred to the British Empire as a whole. At the time of the founding of the League of Nations in 1924, the League Covenant made provision for the admission of any "fully self-governing state, Dominion, or Colony", the implication being that "Dominion status was something between that of a colony and a state".

With the adoption of the Statute of Westminster 1931, Britain and the Dominions (except Newfoundland) formed the British Commonwealth of Nations. Dominions asserted full legislative independence, with direct access to the Monarch as Head of State previously reserved only for British governments. It also recognised autonomy in foreign affairs, including participation as autonomous nations in the League of Nations with full power over appointing ambassadors to other countries.

Following the Second World War, the changes in the constitutional relationship between the countries that continued to share a common sovereign with the United Kingdom led to the upper case term 'Dominion' falling out of use.

"His/Her Majesty's dominions"

The status of "Dominion" established by the Statute of Westminster in 1931 was capitalised to distinguish it from the more general sense of "within the crown's dominions".

The phrase the crown's dominions or His/Her Majesty's dominions is a legal and constitutional phrase that refers to all the realms and territories of the British sovereign, whether independent or not. These territories include the United Kingdom, and its colonies, including those that had become Dominions. Dependent territories that had never been annexed and were not colonies of the Crown, were notionally foreign territory and not "within the crown's dominions". When these territories—including protectorates and protected states (a status with greater powers of self-government), as well as League of Nations mandates, which then became United Nations Trust Territories—were granted independence, and at the same time recognised the British monarch as head of state, the United Kingdom act granting independence declared that such and such a territory "shall form part of Her Majesty's dominions", and so become part of the territory in which the Queen exercises sovereignty, not merely suzerainty.

The legal status of "Dominion" under British nationality law ceased to exist from 1 January 1949, when it was determined each Dominion would legislate for its own citizenship. However, "Dominion status" itself never ceased to exist within the greater scope of British law, because acts pertaining to "Dominion status", such as the Statute of Westminster 1931, have not been repealed in both the United Kingdom and historic Dominions such as Canada. The term "within the crown's dominions" continues to apply in British law to those territories in which the British monarch remains head of state, and the term "self-governing dominion" is used in some legislation. When a territory ceases to recognise the monarch as head of state, this status is changed by statute. Thus, for example, the British Ireland Act 1949, recognised that the Republic of Ireland had "ceased to be part of His Majesty's dominions".

Historical development

Responsible government: precursor to Dominion status

The foundation of "Dominion" status followed the achievement of internal self-rule in British Colonies, in the specific form of full responsible government (as distinct from "representative government"). Colonial responsible government began to emerge during the mid-19th century. The legislatures of Colonies with responsible government were able to make laws in all matters other than foreign affairs, defence and international trade, these being powers which remained with the Parliament of the United Kingdom.

Nova Scotia soon followed by the Province of Canada (which included modern southern Ontario and southern Quebec) were the first Colonies to achieve responsible government, in 1848. Prince Edward Island followed in 1851, and New Brunswick and Newfoundland in 1855. All except for Newfoundland and Prince Edward Island agreed to form a new federation named Canada from 1867. This was instituted by the British Parliament in the British North America Act, 1867. (See also: Canadian Confederation). Section 3 of the Act referred to the new entity as a "Dominion", the first such entity to be created. From 1870 the Dominion included two vast neighbouring British territories that did not have any form of self-government: Rupert's Land and the North-Western Territory, parts of which later became the Provinces of Manitoba, Saskatchewan, Alberta, and the separate territories, the Northwest Territories, Yukon and Nunavut. In 1871, the Crown Colony of British Columbia became a Canadian province, Prince Edward Island joined in 1873 and Newfoundland in 1949.

The conditions under which the four separate Australian colonies—New South Wales, Tasmania, Western Australia, South Australia—and New Zealand could gain full responsible government were set out by the British government in the Australian Constitutions Act 1850. The Act also separated the Colony of Victoria (in 1851) from New South Wales. During 1856, responsible government was achieved by New South Wales, Victoria, South Australia, and Tasmania, and New Zealand. The remainder of New South Wales was divided in three in 1859, a change that established most of the present borders of NSW; the Colony of Queensland, with its own responsible self-government, and the Northern Territory (which was not granted self-government prior to federation of the Australian Colonies). Western Australia did not receive self-government until 1891, mainly because of its continuing financial dependence on the UK Government. After protracted negotiations (that initially included New Zealand), six Australian colonies with responsible government (and their dependent territories) agreed to federate, along Canadian lines, becoming the Commonwealth of Australia, in 1901.

In South Africa, the Cape Colony became the first British self-governing Colony, in 1872. (Until 1893, the Cape Colony also controlled the separate Colony of Natal.) Following the Second Boer War (1899–1902), the British Empire assumed direct control of the Boer Republics, but transferred limited self-government to Transvaal in 1906, and the Orange River Colony in 1907.

The New Zealand Observer (1907) shows New Zealand Prime Minister Joseph Ward as a pretentious dwarf beneath a massive "Dominion" top hat. The caption reads: The Surprise Packet:
Canada: "Rather large for him, is it not?"
Australia: "Oh, his head is swelling rapidly. The hat will soon fit."

The Commonwealth of Australia was recognised as a Dominion in 1901, and the Dominion of New Zealand and the Dominion of Newfoundland were officially given Dominion status in 1907, followed by the Union of South Africa in 1910.

Canadian Confederation and evolution of the term Dominion

In connection with proposals for the future government of British North America, use of the term "Dominion" was suggested by Samuel Leonard Tilley at the London Conference of 1866 discussing the confederation of the Province of Canada (subsequently becoming the provinces of Ontario and Quebec), Nova Scotia and New Brunswick into "One Dominion under the Name of Canada", the first federation internal to the British Empire.[25] Tilley's suggestion was taken from the 72nd Psalm, verse eight, "He shall have dominion also from sea to sea, and from the river unto the ends of the earth", which is echoed in the national motto, "A Mari Usque Ad Mare".[26] The new government of Canada under the British North America Act, 1867 began to use the phrase "Dominion of Canada" to designate the new, larger nation. However, neither the Confederation nor the adoption of the title of "Dominion" granted extra autonomy or new powers to this new federal level of government.[27][28] Senator Eugene Forsey wrote that the powers acquired since the 1840s that established the system of responsible government in Canada would simply be transferred to the new Dominion government:

By the time of Confederation in 1867, this system had been operating in most of what is now central and eastern Canada for almost 20 years. The Fathers of Confederation simply continued the system they knew, the system that was already working, and working well.

The constitutional scholar Andrew Heard argues that Confederation did not legally change Canada's colonial status to anything approaching its later status of a Dominion.

At its inception in 1867, Canada's colonial status was marked by political and legal subjugation to British Imperial supremacy in all aspects of government—legislative, judicial, and executive. The Imperial Parliament at Westminster could legislate on any matter to do with Canada and could override any local legislation, the final court of appeal for Canadian litigation lay with the Judicial Committee of the Privy Council in London, the Governor General had a substantive role as a representative of the British government, and ultimate executive power was vested in the British Monarch—who was advised only by British ministers in its exercise. Canada's independence came about as each of these sub-ordinations was eventually removed.

When the Dominion of Canada was created in 1867, it was granted powers of self-government to deal with all internal matters, but Britain still retained overall legislative supremacy. This Imperial supremacy could be exercised through several statutory measures. In the first place, the British North America Act of 1867 provided in Section 55 that the Governor General may reserve any legislation passed by the two Houses of Parliament for "the signification of Her Majesty's pleasure", which is determined according to Section 57 by the British Monarch in Council. Secondly, Section 56 provides that the Governor General must forward to "one of Her Majesty's Principal Secretaries of State" in London a copy of any Federal legislation that has been assented to. Then, within two years after the receipt of this copy, the (British) Monarch in Council could disallow an Act. Thirdly, at least four pieces of Imperial legislation constrained the Canadian legislatures. The Colonial Laws Validity Act of 1865 provided that no colonial law could validly conflict with, amend, or repeal Imperial legislation that either explicitly, or by necessary implication, applied directly to that colony. The Merchant Shipping Act of 1894, as well as the Colonial Courts of Admiralty Act of 1890 required reservation of Dominion legislation on those topics for approval by the British Government. Also, the Colonial Stock Act of 1900 provided for the disallowance of any Dominion legislation the British government felt would harm British stockholders of Dominion trustee securities. Most importantly, however, the British Parliament could exercise the legal right of supremacy that it possessed over common law to pass any legislation on any matter affecting the colonies.

For decades, the Dominions did not have their own embassies or consulates in foreign countries. International travel and commerce were transacted through British embassies and consulates. For example, matters concerning visas and lost or stolen passports of Dominion citizens were carried out at British diplomatic offices. In the late 1930s and early 1940s, Dominion governments established their own embassies, the first two of which were established by Australia and Canada in Washington, D.C., in the United States.

As Heard later explained, the British government seldom invoked its powers over Canadian legislation. British legislative powers over Canadian domestic policy were largely theoretical and their exercise was increasingly unacceptable in the 1870s and 1880s. The rise to the status of a Dominion and then full independence for Canada and other possessions of the British Empire did not occur by the granting of titles or similar recognition by the British Parliament but by initiatives taken by the new governments of certain former British dependencies to assert their independence and to establish constitutional precedents.

What is remarkable about this whole process is that it was achieved with a minimum of legislative amendments. Much of Canada's independence arose from the development of new political arrangements, many of which have been absorbed into judicial decisions interpreting the constitution—with or without explicit recognition. Canada's passage from being an integral part of the British Empire to being an independent member of the Commonwealth richly illustrates the way in which fundamental constitutional rules have evolved through the interaction of constitutional convention, international law, and municipal statute and case law.

What was significant about the creation of the Canadian and Australian federations was not that they were instantly granted wide new powers by the Imperial centre at the time of their creation; but that they, because of their greater size and prestige, were better able to exercise their existing powers and lobby for new ones than the various colonies they incorporated could have done separately. They provided a new model which politicians in New Zealand, Newfoundland, South Africa, Ireland, India, Malaysia could point to for their own relationship with Britain. Ultimately, "[Canada's] example of a peaceful accession to independence with a Westminster system of government came to be followed by 50 countries with a combined population of more than 2-billion people."

Colonial Conference of 1907

Issues of colonial self-government spilled into foreign affairs with The Second Boer War (1899–1902). The self-governing colonies contributed significantly to British efforts to stem the insurrection, but ensured that they set the conditions for participation in these wars. Colonial governments repeatedly acted to ensure that they determined the extent of their peoples' participation in imperial wars in the military build-up to the First World War.

The assertiveness of the self-governing nations was recognised in the Colonial Conference of 1907, which implicitly introduced the idea of the Dominion as a self-governing nation by referring to Canada and Australia as Dominions. It also retired the name "Colonial Conference" and mandated that meetings take place regularly to consult Dominions in running the foreign affairs of the empire.

The Colony of New Zealand, which chose not to take part in Australian federation, became the Dominion of New Zealand on 26 September 1907; Newfoundland became a Dominion on the same day. The Union of South Africa was referred to as a Dominion upon its creation in 1910.

First World War and Treaty of Versailles

The Parliamentary Recruiting Committee produced this First World War poster. Designed by Arthur Wardle, the poster urges men from the Dominions of the British Empire to enlist in the war effort.

The initiatives and contributions of British colonies to the British war effort in the First World War were recognised by Britain with the creation of the Imperial War Cabinet in 1917, which gave them a say in the running of the war. Dominion status as self-governing states, as opposed to symbolic titles granted various British colonies, waited until 1919, when the self-governing Dominions signed the Treaty of Versailles independently of the British government and became individual members of the League of Nations. This ended the purely colonial status of the Dominions.

The First World War ended the purely colonial period in the history of the Dominions. Their military contribution to the Allied war effort gave them claim to equal recognition with other small states and a voice in the formation of policy. This claim was recognised within the Empire by the creation of the Imperial War Cabinet in 1917, and within the community of nations by Dominion signatures to the Treaty of Versailles and by separate Dominion representation in the League of Nations. In this way the "self-governing Dominions", as they were called, emerged as junior members of the international community. Their status defied exact analysis by both international and constitutional lawyers, but it was clear that they were no longer regarded simply as colonies of Britain.

Irish Free State

The Irish Free State, set up in 1922 after the Anglo-Irish War, was the third Dominion to appoint a non-UK born, non-aristocratic Governor-General when Timothy Michael Healy, following the tenures of Sir Gordon Drummond in Canada and of Sir Walter Davidson and Sir William Allardyce in Newfoundland, took the position in 1922. Dominion status was never popular in the Irish Free State where people saw it as a face-saving measure for a British government unable to countenance a republic in what had previously been the United Kingdom of Great Britain and Ireland. Successive Irish governments undermined the constitutional links with the United Kingdom. In 1937 Ireland, as it renamed itself, adopted a new republican constitution that included powers for a president of Ireland. At the same time, a law delegating functions to the King, not as King in Ireland but as the symbol of the co-operation amongst Commonwealth countries with which Ireland associated itself, continued to apply in external relations. The last statutory functions of the King with respect to Ireland were abolished in 1949.

Balfour Declaration of 1926 and Statute of Westminster

The Balfour Declaration of 1926, and the subsequent Statute of Westminster, 1931, restricted Britain's ability to pass or affect laws outside of its own jurisdiction.

Until 1931, Newfoundland was referred to as a colony of the United Kingdom, as for example, in the 1927 reference to the Judicial Committee of the Privy Council to delineate the Quebec-Labrador boundary. Full autonomy was granted by the United Kingdom parliament with the Statute of Westminster in December 1931. However, the government of Newfoundland "requested the United Kingdom not to have sections 2 to 6[—]confirming Dominion status[—]apply automatically to it[,] until the Newfoundland Legislature first approved the Statute, approval which the Legislature subsequently never gave".

In 1934, following Newfoundland's economic collapse, and with the approval of its own legislature, Britain suspended self-government in Newfoundland and instituted a "Commission of Government", which continued until Newfoundland became a province of Canada in 1949.

White Dominions

Canada, Australia, New Zealand, the Irish Free State, Newfoundland and South Africa (prior to becoming a republic and leaving the Commonwealth in 1961), with their large populations of European descent, were sometimes collectively referred to as the "White Dominions".

Dominions

List of Dominions

Country From To Status
 Canada 1867

Continues as a Commonwealth realm and member of the Commonwealth of Nations. 'Dominion' was conferred as the country's title in the 1867 constitution, as a substitute for the title of "kingdom".

 Australia 1901

Continues as a Commonwealth realm and member of the Commonwealth of Nations.

 New Zealand 1907

Continues as a Commonwealth realm and member of the Commonwealth of Nations.

 Dominion of Newfoundland 1907 1934 The colony of Newfoundland enjoyed responsible government from 1855 to 1907 when it became a Dominion. Following the recommendations of a Royal Commission, parliamentary government was suspended in 1934 due to severe financial difficulties resulting from the depression and a series of riots against the Dominion government in 1932. In 1949, it joined Canada and the legislature was restored.
 South Africa 1910 1961 Continued as a monarchy until it became a republic in 1961 under the Republic of South Africa Constitution Act 1961, passed by the Parliament of South Africa, long title "To constitute the Republic of South Africa and to provide for matters incidental thereto", assented to 24 April 1961 to come into operation on 31 May 1961.
 Irish Free State (1922–37)
Éire (1937–49)
1922 1949 The link with the monarchy ceased with the passage of the Republic of Ireland Act 1948, which came into force on 18 April 1949 and declared that the state was a republic.
 Dominion of India 1947 1950 The Union of India (with the addition of Sikkim from 1975) became a federal republic after its constitution came into effect on 26 January 1950.
 Dominion of Pakistan 1947 1956 Continued as a monarchy until 1956 when it became a republic under the name "The Islamic Republic of Pakistan": Constitution of 1956.
 Dominion of Ceylon 1948 1972 Continued as a monarchy until 1972 when it became a republic under the name of Sri Lanka.
  • There was no single constitutional or legislative change that abolished the status of "dominions". The accession proclamation of 1952 referred to "realms", and the Royal Style and Titles Acts of 1953 changed references to "dominions" in the monarch's titles in the various Dominions to "realms", after which the term dominion generally fell into disuse, and the countries sharing the same monarch as the United Kingdom came to be referred to as realms (with the possible exception of Canada; see also Name of Canada).

  • The Irish Free State was renamed Éire in Irish or Ireland in English in 1937. In 1937–1949, the Dominion was referred to as "Eire" by the British government. See also Names of the Irish state.

    Australia

    Four colonies of Australia had enjoyed responsible government since 1856: New South Wales, Victoria, Tasmania and South Australia. Queensland had responsible government soon after its founding in 1859. Because of ongoing financial dependence on Britain, Western Australia became the last Australian colony to attain self-government in 1890. During the 1890s, the colonies voted to unite and in 1901 they were federated under the British Crown as the Commonwealth of Australia by the Commonwealth of Australia Constitution Act. The Constitution of Australia had been drafted in Australia and approved by popular consent. Thus Australia is one of the few countries established by a popular vote. Under the Balfour Declaration of 1926, the federal government was regarded as coequal with (and not subordinate to) the British and other Dominion governments, and this was given formal legal recognition in 1942 (when the Statute of Westminster was adopted retroactively to the commencement of the Second World War in 1939). In 1930, the Australian prime minister, James Scullin, reinforced the right of the overseas Dominions to appoint native-born governors-general, when he advised King George V to appoint Sir Isaac Isaacs as his representative in Australia, against the wishes of the opposition and officials in London. The governments of the states (colonies before 1901) remained under the Commonwealth but retained links to the UK until the passage of the Australia Act 1986.

    Canada

    Dominion of Canada Postage Stamp, 1898

    The term Dominion is employed in the Constitution Act, 1867 (originally the British North America Act, 1867), and describes the resulting political union. Specifically, the preamble of the act states: "Whereas the Provinces of Canada, Nova Scotia, and New Brunswick have expressed their Desire to be federally united into One Dominion under the Crown of the United Kingdom of Great Britain and Ireland, with a Constitution similar in Principle to that of the United Kingdom ..." Furthermore, Sections 3 and 4 indicate that the provinces "shall form and be One Dominion under the Name of Canada; and on and after that Day those Three Provinces shall form and be One Dominion under that Name accordingly".

    According to the Canadian Encyclopedia (1999), "The word came to be applied to the federal government and Parliament, and under the Constitution Act, 1982, 'Dominion' remains Canada's official title."

    Usage of the phrase Dominion of Canada was employed as the country's name after 1867, predating the general use of the term Dominion as applied to the other autonomous regions of the British Empire after 1907. The phrase Dominion of Canada does not appear in the 1867 act nor in the Constitution Act, 1982, but does appear in the Constitution Act, 1871, other contemporaneous texts, and subsequent bills. References to the Dominion of Canada in later acts, such as the Statute of Westminster, do not clarify the point because all nouns were formally capitalised in British legislative style. Indeed, in the original text of the Constitution Act, 1867, "One" and "Name" were also capitalised.

    Frank Scott theorised that Canada's status as a Dominion ended when Canadian parliament declared war on Germany on 9 September 1939, separately and distinctly from the United Kingdom's declaration of war six days earlier. By the 1950s, the term Dominion of Canada was no longer used by the United Kingdom, which considered Canada a "Realm of the Commonwealth". The government of Louis St. Laurent ended the practice of using Dominion in the statutes of Canada in 1951. This began the phasing out of the use of Dominion, which had been used largely as a synonym of "federal" or "national" such as "Dominion building" for a post office, "Dominion-provincial relations", and so on. The last major change was renaming the national holiday from Dominion Day to Canada Day in 1982. Official bilingualism laws also contributed to the disuse of Dominion, as it has no acceptable equivalent in French.

    While the term may be found in older official documents, and the Dominion Carillonneur still tolls at Parliament Hill, it is now hardly used to distinguish the federal government from the provinces or (historically) Canada before and after 1867. Nonetheless, the federal government continues to produce publications and educational materials that specify the currency of these official titles. The Constitution Act, 1982 does not mention and does not remove the title, and therefore a constitutional amendment may be required to change it.

    The word Dominion has been used with other agencies, laws, and roles:

    Notable Canadian corporations and organisations (not affiliated with government) that have used Dominion as a part of their name have included:

    Ceylon

    Ceylon, which, as a Crown colony, was originally promised "fully responsible status within the British Commonwealth of Nations", was formally granted independence as a Dominion in 1948. In 1972 it adopted a republican constitution to become the Free, Sovereign and Independent Republic of Sri Lanka. By a new constitution in 1978, it became the Democratic Socialist Republic of Sri Lanka.

    India, Pakistan and Bangladesh

    British India acquired a partially representative government in 1909, and the first Parliament was introduced in 1919. Discussions on the further devolution of power, and granting of Dominion status, continued through the 1920s, with the Commonwealth of India Bill 1925, Simon Commission 1927–1930, and Nehru Report 1928 being often cited proposals. Further powers were eventually devolved, following the 1930–1932 Round Table Conferences, to the locally elected legislatures, via the Government of India Act 1935. The Cripps Mission of 1942 proposed the further devolution of powers, within Dominion status, to the political leadership of British India. Cripps's plan was rejected and full independence was sought. Pakistan (including Muslim-majority East Bengal forming East Pakistan) seceded from India at the point of Indian Independence with the passage of the Indian Independence Act 1947 and ensuing partition, resulting in two dominions. For India, dominion status was transitory until its new republican constitution was drafted and promulgated in 1950. Pakistan remained a dominion until it became an Islamic republic under its 1956 constitution. East Pakistan gained independence from Pakistan through Liberation War, as Bangladesh, in 1971.

    Irish Free State / Ireland

    The Irish Free State (Ireland from 1937) was a British Dominion between 1922 and 1949. As established by the Irish Free State Constitution Act of the United Kingdom Parliament on 6 December 1922 the new state—which had Dominion status in the likeness of that enjoyed by Canada within the British Commonwealth of Nations—comprised the whole of Ireland. However, provision was made in the Act for the Parliament of Northern Ireland to opt out of inclusion in the Irish Free State, which—as had been widely expected at the time—it duly did one day after the creation of the new state, on 7 December 1922.

    Following a plebiscite of the people of the Free State held on 1 July 1937, a new constitution came into force on 29 December of that year, establishing a successor state with the name of "Ireland" which ceased to participate in Commonwealth conferences and events. Nevertheless, the United Kingdom and other member states of the Commonwealth continued to regard Ireland as a Dominion owing to the unusual role accorded to the British Monarch under the Irish External Relations Act of 1936. Ultimately, however, Ireland's Oireachtas passed the Republic of Ireland Act 1948, which came into force on 18 April 1949 and unequivocally ended Ireland's links with the British Monarch and the Commonwealth.

    Newfoundland

    In 1934, after a series of financial difficulties (owing in part to Newfoundland's railway debt from the 1890s, and its debt from the First World War, both of which were exacerbated by the collapse of fish prices during the Great Depression) and a riot against the elected government, Newfoundland voluntarily relinquished its elected parliament and autonomy, becoming a dependent territory of the British Empire until 1949. During these 15 years, Newfoundland was ruled by the Newfoundland Commission of Government, an unelected body of civil servants who were directly subordinate to the British Government in London, under the authority of a British statute, the Newfoundland Act, 1933. Despite the suspension of its legislature, and its formal loss of Dominion status, Newfoundland continued to be regarded during these 15 years as a de facto Dominion - evidently shown by the fact that Newfoundland continued to be a responsibility of the Dominions Office in London - the intention of its commission of government not only to deal with Newfoundland's affairs, and dire economic situation, but to prepare the populace for the day that the legislature would be reconvened, and nationhood thus resumed. After two referendums in 1948, Newfoundlanders rejected both the continuance of the Newfoundland Commission of Government and return to responsible government, voting instead to join the Canada as its 10th province. This was achieved under the British North America Act, 1949 (now known as the Newfoundland Act), which was passed in the Parliament of the United Kingdom on 23 March 1949, prior to the London Declaration of 28 April 1949.

    New Zealand

    The New Zealand Constitution Act 1852 gave New Zealand its own Parliament (General Assembly) and home rule in 1852. In 1907 New Zealand was proclaimed the Dominion of New Zealand.[63] New Zealand, Canada, and Newfoundland used the word Dominion in the official title of the nation, whereas Australia used Commonwealth of Australia and South Africa Union of South Africa. New Zealand adopted the Statute of Westminster in 1947 and in the same year legislation passed in London gave New Zealand full powers to amend its own constitution. In 1986, the New Zealand parliament passed the Constitution Act 1986, which repealed the Constitution Act of 1852 and the last constitutional links with the United Kingdom, formally ending its Dominion status.

    South Africa

    The Union of South Africa was formed in 1910 from the four self-governing colonies of the Cape Colony, Natal, the Transvaal, and the Orange River Colony (the last two were former Boer republics). The South Africa Act 1909 provided for a Parliament consisting of a Senate and a House of Assembly. The provinces had their own legislatures. In 1961, the Union of South Africa adopted a new constitution, became a republic, left the Commonwealth (and re-joined following end of Apartheid rule in 1994), and became the present-day Republic of South Africa.

    Southern Rhodesia

    A map. See description
    Southern Rhodesia (Zimbabwe since 1980), coloured red on a map of Africa

    Southern Rhodesia (renamed Zimbabwe in 1980) was a special case in the British Empire. Although it was never a Dominion de jure, it was treated as a Dominion in many respects, and came to be regarded as a de facto Dominion. Southern Rhodesia was formed in 1923 out of territories of the British South Africa Company and established as a self-governing colony with substantial autonomy on the model of the Dominions. The imperial authorities in London retained direct powers over foreign affairs, constitutional alterations, native administration and bills regarding mining revenues, railways and the governor's salary.

    Southern Rhodesia was not one of the territories that were mentioned in the 1931 Statute of Westminster although relations with Southern Rhodesia were administered in London through the Dominion Office, not the Colonial Office. When the Dominions were first treated as foreign countries by London for the purposes of diplomatic immunity in 1952, Southern Rhodesia was included in the list of territories concerned. This semi-Dominion status continued in Southern Rhodesia between 1953 and 1963, when it joined Northern Rhodesia and Nyasaland in the Central African Federation, with the latter two territories continuing to be British protectorates. When Northern Rhodesia was given independence in 1964 it adopted the new name of Zambia, prompting Southern Rhodesia to shorten its name to Rhodesia, but Britain did not recognise this latter change.

    Rhodesia unilaterally declared independence from Britain in 1965 as a result of the British government's insistence on no independence before majority rule (NIBMAR). London regarded this declaration as illegal, and applied sanctions and expelled Rhodesia from the sterling area. Rhodesia continued with its Dominion-style constitution until 1970, and continued to issue British passports to its citizens. The Rhodesian government continued to profess its loyalty to the Sovereign, despite being in a state of rebellion against Her Majesty's Government in London, until 1970, when it adopted a republican constitution following a referendum the previous year. This endured until the state's reconstitution as Zimbabwe Rhodesia in 1979 under the terms of the Internal Settlement; this lasted until the Lancaster House Agreement of December 1979, which put it under interim British rule while fresh elections were held. The country achieved independence deemed legal by the international community in April 1980, when Britain granted independence under the name Zimbabwe.

    From Dominions to Commonwealth realms

    The prime ministers of Britain and the four major Dominions at the 1944 Commonwealth Prime Ministers' Conference. Left to right: William Lyon Mackenzie King (Canada); Jan Smuts (South Africa); Winston Churchill (UK); Peter Fraser (New Zealand); John Curtin (Australia)

    Initially, the Dominions conducted their own trade policy, some limited foreign relations and had autonomous armed forces, although the British government claimed and exercised the exclusive power to declare wars. However, after the passage of the Statute of Westminster the language of dependency on the Crown of the United Kingdom ceased, where the Crown itself was no longer referred to as the Crown of any place in particular but simply as "the Crown". Arthur Berriedale Keith, in Speeches and Documents on the British Dominions 1918–1931, stated that "the Dominions are sovereign international States in the sense that the King in respect of each of His Dominions (Newfoundland excepted) is such a State in the eyes of international law". After then, those countries that were previously referred to as "Dominions" became Commonwealth realms where the sovereign reigns no longer as the British monarch, but as monarch of each nation in its own right, and are considered equal to the UK and one another.

    The Second World War, which fatally undermined Britain's already weakened commercial and financial leadership, further loosened the political ties between Britain and the Dominions. Australian Prime Minister John Curtin's unprecedented action (February 1942) in successfully countermanding an order from British Prime Minister Winston Churchill that Australian troops be diverted to defend British-held Burma (the 7th Division was then en route from the Middle East to Australia to defend against an expected Japanese invasion) demonstrated that Dominion governments might no longer subordinate their own national interests to British strategic perspectives. To ensure that Australia had full legal power to act independently, particularly in relation to foreign affairs, defence industry and military operations, and to validate its past independent action in these areas, Australia formally adopted the Statute of Westminster in October 1942 and backdated the adoption to the start of the war in September 1939.

    The Dominions Office merged with the India Office as the Commonwealth Relations Office upon the independence of India and Pakistan in August 1947. The last country officially made a Dominion was Ceylon in 1948.

    When the British Nationality Act 1948 entered into force on 1 January 1949, the former Dominions became fully independent, and adopted their own legislation governing nationality. In British nationality law, the Dominions were then referred to as "independent Commonwealth countries"; other former British dependencies that joined the Commonwealth were added to the list of "independent Commonwealth Countries" as they gained independence.

    Ireland ceased to be a member of the Commonwealth on 18 April 1949, upon the coming into force of the Republic of Ireland Act 1948. This formally signalled the end of the former dependencies' common constitutional connection to the British Crown. India also adopted a republican constitution in January 1950. Unlike many dependencies that became republics, Ireland never re-joined the Commonwealth, which agreed to accept the British monarch as head of that association of independent states (although most of the individual countries had become republics).

    The independence of the separate realms was emphasised after the accession of Queen Elizabeth II in 1952, when she was proclaimed not just as Queen of the United Kingdom, but also Queen of Canada, Queen of Australia, Queen of New Zealand, Queen of South Africa, and of all her other "realms and territories" etc. This also reflected the change from Dominion to realm; in the proclamation of Queen Elizabeth II's new titles in 1953, the phrase "of her other Realms and Territories" replaced "Dominion" with another mediaeval French word with the same connotation, "realm" (from royaume). Thus, recently, when referring to one of those fifteen countries within the Commonwealth of Nations that share the same monarch, the phrase Commonwealth realm has come into common usage instead of Dominion to differentiate the Commonwealth nations that continue to share the monarch as head of state (Australia, Canada, New Zealand, Jamaica, etc.) from those that do not (India, Pakistan, South Africa, etc.). The term "Dominion" is still found in the Canadian constitution where it appears numerous times, but it is largely a vestige of the past, as the Canadian government does not actively use it (see Canada section). The term "realm" does not appear in the Canadian constitution.

    The practice of designating a diplomatic representative named "High Commissioner" (instead of "ambassador") for communication between the government of a Dominion and the British government in London continues in respect of members of the Commonwealth, including those that were never Dominions and those that have become republics.

    Newly independent territories sometimes referred to as Dominions

    The term "Dominion" remained in informal use for some years when relating to newly independent territories and was sometimes used to refer to the status of former British territories during an immediate post-independence period while the British monarch remained head of state, and the form of government a Westminster-style parliamentary democracy. The legal status of Dominion in British nationality law had ceased to exist on 1 January 1949. However, leaders of the independence movements sometimes called for Dominion status as one stage in the negotiations for independence (for example, Kwame Nkrumah of Ghana). Moreover, while these independent states retained the British monarch as head of state, they remained "within the Crown's dominions" in British law, leading to the confusion of terminology. These constitutions were typically replaced by republican constitutions within a few years.

    After World War II, Britain attempted to repeat the Dominion model in decolonising the Caribbean. ... Though several colonies, such as Guyana and Trinidad and Tobago, maintained their formal allegiance to the British monarch, they soon revised their status to become republics. Britain also attempted to establish a Dominion model in decolonising Africa, but it, too, was unsuccessful. ... Ghana, the first former colony declared a Dominion in 1957, soon demanded recognition as a republic. Other African nations followed a similar pattern throughout the 1960s: Nigeria, Tanganyika, Uganda, Kenya, and Malawi. In fact, only Gambia, Sierra Leone, and Mauritius retained their Dominion status for more than three years.

    As the above quote indicates, the term Dominion was sometimes applied in Africa to Ghana (formerly the Gold Coast) during the period from 1957 until 1960, when it became the Republic of Ghana; Nigeria from 1960 until 1963, when it became the Federal Republic of Nigeria; Uganda from 1962 to 1963; Kenya, from 1963 to 1964; Tanganyika from 1961 to 1962, after which it became a republic and then merged with the former British protectorate of Zanzibar to become Tanzania; Gambia from 1965 until 1970; Sierra Leone from 1961 to 1971; and Mauritius from 1968 to 1992. Malta also retained the Queen as head of state from 1964 to 1974 under the name of State of Malta. The term was also applied to Fiji upon independence. Similar occasional references to Barbados (which retained the Queen as head of state from 1966 to 2021) as a "dominion" can be found in publications as late as the 1970s.

    Hydrogen-like atom

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