Global distribution of incoming shortwave solar radiation averaged over the years 1981–2010 from the CHELSA-BIOCLIM+ data setThe
shield effect of Earth's atmosphere on solar irradiation. The top image
is the annual mean solar irradiation (or insolation) at the top of Earth's atmosphere
(TOA); the bottom image shows the annual insolation reaching the
Earth's surface after passing through the atmosphere. The two images use
the same color scale.
Solar irradiance is often integrated over a given time period in order to report the radiant energy emitted into the surrounding environment (joule per square metre, J/m2) during that time period. This integrated solar irradiance is called solar irradiation, solar radiation, solar exposure, solar insolation, or insolation.
Irradiance may be measured in space or at the Earth's surface after atmospheric absorption and scattering. Irradiance in space is a function of distance from the Sun, the solar cycle, and cross-cycle changes. Irradiance on the Earth's surface additionally depends on the tilt of
the measuring surface, the height of the Sun above the horizon, and
atmospheric conditions. Solar irradiance affects plant metabolism and animal behavior.
The study and measurement of solar irradiance has several
important applications, including the prediction of energy generation
from solar power plants, the heating and cooling loads of buildings, climate modeling and weather forecasting, passive daytime radiative cooling applications, and space travel.
Types
Global map of global horizontal radiationGlobal Map of Direct Normal Radiation
There are several measured types of solar irradiance.
Total solar irradiance (TSI) is a measure of the solar power over all wavelengths per unit area incident on the Earth's upper atmosphere. It is measured facing (pointing at / parallel to) the incoming sunlight (i.e. the flux through a surface perpendicular to the incoming sunlight; other angles would not be TSI). The solar constant is a conventional measure of mean TSI at a distance of one astronomical unit (AU).
Direct normal irradiance (DNI), or beam radiation, is measured perpendicularly to the Sun direction. It excludes diffuse solar radiation (radiation that is scattered or reflected by atmospheric components). Direct irradiance is equal to the extraterrestrial irradiance above the atmosphere minus the atmospheric losses due to absorption and scattering. Losses depend on time of day (length of light's path through the atmosphere depending on the solar elevation angle), cloud cover, moisture content and other contents.
The irradiance above the atmosphere also varies with time of year
(because the distance to the Sun varies), although this effect is
generally less significant compared to the effect of losses on DNI.
Solar monitoring system for GHI, DHI and DNI https://eko-instruments.com/product/ms-80sh-plus/Direct horizontal irradiance (DirHI), or beam horizontal irradiance (BHI),
is the direct component of irradiance received on a horizontal surface
as opposed to a surface perpendicular to the direct sunlight.
Diffuse horizontal irradiance (DHI), or diffuse sky radiation,
is the radiation at the Earth's surface from light scattered by the
atmosphere. It is measured on a horizontal surface with radiation coming
from all points in the sky excluding circumsolar radiation (radiation coming from the sun disk). There would be almost no DHI in the absence of atmosphere.
Global horizontal irradiance (GHI) is the
total irradiance from the Sun on a horizontal surface on Earth. For
instantaneous measurement, it is the sum of direct irradiance (after
accounting for the solar zenith angle, , of the Sun) and diffuse horizontal irradiance:
Global tilted irradiance (GTI) is the total radiation received on a surface with defined tilt and azimuth, fixed or Sun-tracking. GTI can be measured or modeled from GHI, DNI, DHI.It is often a reference for photovoltaic power plants, while photovoltaic modules are mounted on the fixed or tracking constructions.
Global normal irradiance (GNI)
is the total irradiance from the Sun at the surface of Earth at a given
location with a surface element perpendicular to the Sun.
Spectral versions of the above irradiances (e.g. spectral TSI, spectral DNI,
etc.) are any of the above with units divided either by meter or
nanometer (for a spectral graph as function of wavelength), or per-Hz (for a spectral function with an x-axis of frequency). When one plots such spectral distributions as a graph, the integral of
the function (area under the curve) will be the (non-spectral)
irradiance. e.g.: Say one had a solar cell on the surface of the earth
facing straight up, and had DNI in units of Wm−2nm−1, graphed as a function of wavelength (in nm). Then, the unit of the integral (Wm−2) is the product of those two units.
Units
The SI unit of irradiance is watts per square metre (W/m2 = Wm−2). The unit of insolation often used in the solar power industry is kilowatt hours per square metre (kWh/m2).
The langley is an alternative unit of insolation. One langley is one thermochemical calorie per square centimetre or 41,840J/m2.
At the top of Earth's atmosphere
Spherical triangle for application of the spherical law of cosines for calculating the solar zenith angle Θ of an observer at latitudeφ and longitudeλ, using the hour angle h and solar declination δ (where δ is latitude of subsolar point, and h is relative longitude of subsolar point)
The average annual solar radiation arriving at the top of the Earth's atmosphere is about 1361W/m2.
This represents the power per unit area of solar irradiance across the
spherical surface surrounding the Sun with a radius equal to the
distance to the Earth (1AU). This means that the approximately circular disc of the Earth, as viewed from the Sun, receives a roughly stable 1361W/m2 at all times. The area of this circular disc is πr2, in which r is the radius of the Earth. Because the Earth is approximately spherical, it has total area ,
meaning that the solar radiation arriving at the top of the atmosphere,
averaged over the entire surface of the Earth, is simply divided by
four to get 340W/m2. In other words, averaged over the year and the day, the Earth's atmosphere receives 340W/m2 from the Sun. This figure is important in radiative forcing.
Distribution is based on a fundamental identity from spherical trigonometry, the spherical law of cosines:
where a, b and c are arc lengths, in radians, of the sides of a spherical triangle. C is the angle in the vertex opposite the side which has arc length c. Applied to the calculation of solar zenith angleΘ, the following applies to the spherical law of cosines:
This equation can be also derived from a more general formula:
where β is an angle from the horizontal and γ is the solar azimuth angle.
The derivation of the cosine of solar zenith angle, , based on vector analysis instead of spherical trigonometry is also available in the article about solar azimuth angle.
, the theoretical daily-average irradiation at the top of the atmosphere, where θ is the polar angle of the Earth's orbit, and θ=0 at the March equinox, and θ=90° at the June solstice; φ is the latitude of the Earth. The calculation assumed conditions appropriate for 2000A.D.: a solar constant of S0=1367Wm−2, obliquity of ε=23.4398°, longitude of perihelion of ϖ=282.895°, eccentricity e=0.016704. Contour labels (green) are in units ofWm−2.
The separation of Earth from the Sun can be denoted RE and the mean distance can be denoted R0, approximately 1 astronomical unit (AU). The solar constant is denoted S0.
The solar flux density (insolation) onto a plane tangent to the sphere
of the Earth, but above the bulk of the atmosphere (elevation 100km or greater) is:
The average of Q over a day is the average of Q over one rotation, or the hour angle progressing from h = π to h = −π:
Let h0 be the hour angle when Q becomes positive. This could occur at sunrise when , or for h0 as a solution of
or
If tan(φ) tan(δ) > 1, then the sun does not set and the sun is already risen at h = π, so ho = π. If tan(φ) tan(δ) < −1, the sun does not rise and .
is nearly constant over the course of a day, and can be taken outside the integral
Therefore:
Let θ be the conventional polar angle describing a planetary orbit. Let θ=0 at the March equinox. The declinationδ as a function of orbital position is
where ε is the obliquity. (Note: The correct formula, valid for any axial tilt, is .) The conventional longitude of perihelion ϖ is defined relative to the March equinox, so for the elliptical orbit:
or
With knowledge of ϖ, ε and e from astrodynamical calculations and So from a consensus of observations or theory, can be calculated for any latitude φ and θ. Because of the elliptical orbit, and as a consequence of Kepler's second law, θ does not progress uniformly with time. Nevertheless, θ=0° is exactly the time of the March equinox, θ=90° is exactly the time of the June solstice, θ=180° is exactly the time of the September equinox and θ=270° is exactly the time of the December solstice.
A simplified equation for irradiance on a given day is:
where n is a number of a day of the year.
Variation
Total solar irradiance (TSI) changes slowly on decadal and longer timescales. The variation during solar cycle 21 was about 0.1% (peak-to-peak). In contrast to older reconstructions, most recent TSI reconstructions point to an increase of only about 0.05% to 0.1% between the 17th century Maunder Minimum and the present. However, current understanding based on various lines of evidence
suggests that the lower values for the secular trend are more probable. In particular, a secular trend greater than 2 Wm−2 is considered highly unlikely. Ultraviolet irradiance (EUV) varies by approximately 1.5 percent from solar maxima to minima, for 200 to 300nm wavelengths. However, a proxy study estimated that UV has increased by 3.0% since the Maunder Minimum.
Variations in Earth's orbit, resulting changes in solar energy flux at high latitude, and the observed glacial cycles
Some variations in insolation are not due to solar changes but rather due to the Earth moving between its perihelion and aphelion, or changes in the latitudinal distribution of radiation. These orbital changes or Milankovitch cycles
have caused radiance variations of as much as 25% (locally; global
average changes are much smaller) over long periods. The most recent
significant event was an axial tilt of 24° during boreal summer near the
Holocene climatic optimum.
Obtaining a time series for a
for a particular time of year, and particular latitude, is a useful
application in the theory of Milankovitch cycles. For example, at the
summer solstice, the declination δ is equal to the obliquityε. The distance from the Sun is
For this summer solstice calculation, the role of the elliptical orbit is entirely contained within the important product , the precession index, whose variation dominates the variations in insolation at 65°N
when eccentricity is large. For the next 100,000 years, with variations
in eccentricity being relatively small, variations in obliquity
dominate.
Measurement
The space-based TSI record comprises measurements from more than ten radiometers and spans three solar cycles.
All modern TSI satellite instruments employ active cavity electrical substitution radiometry.
This technique measures the electrical heating needed to maintain an
absorptive blackened cavity in thermal equilibrium with the incident
sunlight which passes through a precision aperture of calibrated area. The aperture is modulated via a shutter. Accuracy uncertainties of <0.01% are required to detect long term solar irradiance variations, because expected changes are in the range 0.05–0.15W/m2 per century.
Intertemporal calibration
In orbit, radiometric
calibrations drift for reasons including solar degradation of the
cavity, electronic degradation of the heater, surface degradation of the
precision aperture and varying surface emissions and temperatures that
alter thermal backgrounds. These calibrations require compensation to
preserve consistent measurements.
For various reasons, the sources do not always agree. The Solar Radiation and Climate Experiment/Total Irradiance Measurement (SORCE/TIM) TSI values are lower than prior measurements by the Earth Radiometer Budget Experiment (ERBE) on the Earth Radiation Budget Satellite (ERBS), VIRGO on the Solar Heliospheric Observatory (SoHO) and the ACRIM instruments on the Solar Maximum Mission (SMM), Upper Atmosphere Research Satellite (UARS) and ACRIMSAT.
Pre-launch ground calibrations relied on component rather than
system-level measurements since irradiance standards at the time lacked
sufficient absolute accuracies.
Measurement stability involves exposing different
radiometer cavities to different accumulations of solar radiation to
quantify exposure-dependent degradation effects. These effects are then
compensated for in the final data. Observation overlaps permits
corrections for both absolute offsets and validation of instrumental
drifts.
Uncertainties of individual observations exceed irradiance
variability (~0.1%). Thus, instrument stability and measurement
continuity are relied upon to compute real variations.
Long-term radiometer drifts can potentially be mistaken
for irradiance variations which can be misinterpreted as affecting
climate. Examples include the issue of the irradiance increase between
cycle minima in 1986 and 1996, evident only in the ACRIM composite (and
not the model) and the low irradiance levels in the PMOD composite
during the 2008 minimum.
Despite the fact that ACRIM I, ACRIM II, ACRIM III, VIRGO
and TIM all track degradation with redundant cavities, notable and
unexplained differences remain in irradiance and the modeled influences
of sunspots and faculae.
Persistent inconsistencies
Disagreement among overlapping observations indicates
unresolved drifts that suggest the TSI record is not sufficiently stable
to discern solar changes on decadal time scales. Only the ACRIM
composite shows irradiance increasing by ~1W/m2between
1986 and 1996. It is noteworthy that the most accurate TSI
reconstructions with empirical and physics-based semi-empirical models
using independent inputs consistently disfavor this increase during the
ACRIM-gap.
Recommendations to resolve the instrument discrepancies
include validating optical measurement accuracy by comparing
ground-based instruments to laboratory references, such as those at National Institute of Science and Technology (NIST); NIST validation of aperture area calibrations uses spares from each instrument; and applying diffraction corrections from the view-limiting aperture.
For ACRIM, NIST determined that diffraction from the
view-limiting aperture contributes a 0.13% signal not accounted for in
the three ACRIM instruments. This correction lowers the reported ACRIM
values, bringing ACRIM closer to TIM. In ACRIM and all other instruments
but TIM, the aperture is deep inside the instrument, with a larger
view-limiting aperture at the front. Depending on edge imperfections
this can directly scatter light into the cavity. This design admits into
the front part of the instrument two to three times the amount of light
intended to be measured; if not completely absorbed or scattered, this
additional light produces erroneously high signals. In contrast, TIM's
design places the precision aperture at the front so that only desired
light enters.
Variations from other sources likely include an annual
systematics in the ACRIM III data that is nearly in phase with the
Sun-Earth distance and 90-day spikes in the VIRGO data coincident with
SoHO spacecraft maneuvers that were most apparent during the 2008 solar
minimum.
TSI Radiometer Facility
TIM's high absolute accuracy creates new opportunities for
measuring climate variables. TSI Radiometer Facility (TRF) is a
cryogenic radiometer that operates in a vacuum
with controlled light sources. L-1 Standards and Technology (LASP)
designed and built the system, completed in 2008. It was calibrated for
optical power against the NIST Primary Optical Watt Radiometer, a
cryogenic radiometer that maintains the NIST radiant power scale to an
uncertainty of 0.02% (1σ). As of 2011 TRF was the
only facility that approached the desired <0.01% uncertainty for
pre-launch validation of solar radiometers measuring irradiance (rather
than merely optical power) at solar power levels and under vacuum
conditions.
TRF encloses both the reference radiometer and the
instrument under test in a common vacuum system that contains a
stationary, spatially uniform illuminating beam. A precision aperture
with an area calibrated to 0.0031% (1σ) determines the
beam's measured portion. The test instrument's precision aperture is
positioned in the same location, without optically altering the beam,
for direct comparison to the reference. Variable beam power provides
linearity diagnostics, and variable beam diameter diagnoses scattering
from different instrument components.
The Glory/TIM and PICARD/PREMOS flight instrument absolute
scales are now traceable to the TRF in both optical power and
irradiance. The resulting high accuracy reduces the consequences of any
future gap in the solar irradiance record.
Difference relative to TRF
Instrument
Irradiance, view-limiting aperture overfilled
Irradiance, precision aperture overfilled
Difference attributable to scatter error
Measured optical power error
Residual irradiance agreement
Uncertainty
SORCE/TIM ground
—N/a
−0.037%
—N/a
−0.037%
0.000%
0.032%
Glory/TIM flight
—N/a
−0.012%
—N/a
−0.029%
0.017%
0.020%
PREMOS-1 ground
−0.005%
−0.104%
0.098%
−0.049%
−0.104%
~0.038%
PREMOS-3 flight
0.642%
0.605%
0.037%
0.631%
−0.026%
~0.027%
VIRGO-2 ground
0.897%
0.743%
0.154%
0.730%
0.013%
~0.025%
2011 reassessment
The most probable value of TSI representative of solar minimum is 1360.9±0.5W/m2, lower than the earlier accepted value of 1365.4±1.3W/m2,
established in the 1990s. The new value came from SORCE/TIM and
radiometric laboratory tests. Scattered light is a primary cause of the
higher irradiance values measured by earlier satellites in which the
precision aperture is located behind a larger, view-limiting aperture.
The TIM uses a view-limiting aperture that is smaller than the precision
aperture that precludes this spurious signal. The new estimate is from
better measurement rather than a change in solar output.
A regression model-based split of the relative proportion
of sunspot and facular influences from SORCE/TIM data accounts for 92%
of observed variance and tracks the observed trends to within TIM's
stability band. This agreement provides further evidence that TSI
variations are primarily due to solar surface magnetic activity.
Instrument inaccuracies add a significant uncertainty in determining Earth's energy balance. The energy imbalance has been variously measured (during a deep solar minimum of 2005–2010) to be +0.58±0.15W/m2, +0.60±0.17W/m2 and +0.85W/m2. Estimates from space-based measurements range +3–7W/m2. SORCE/TIM's lower TSI value reduces this discrepancy by 1W/m2. This difference between the new lower TIM value and earlier TSI measurements corresponds to a climate forcing of −0.8W/m2, which is comparable to the energy imbalance.
On Earth's surface
A pyranometer, used to measure global irradianceA pyrheliometer, mounted on a solar tracker, is used to measure Direct Normal Irradiance (or beam irradiance).
Average annual solar radiation arriving at the top of the Earth's atmosphere is roughly 1361W/m2. The Sun's rays are attenuated as they pass through the atmosphere, leaving maximum normal surface irradiance at approximately 1000W/m2 at sea level on a clear day. When 1361W/m2 is arriving above the atmosphere (when the Sun is at the zenith in a cloudless sky), direct sun is about 1050W/m2, and global radiation on a horizontal surface at ground level is about 1120W/m2. The latter figure includes radiation scattered or reemitted by the
atmosphere and surroundings. The actual figure varies with the Sun's
angle and atmospheric circumstances. Ignoring clouds, the daily average
insolation for the Earth is approximately 6 kWh/m2 = 21.6 MJ/m2.
The output of, for example, a photovoltaic panel, partly depends on the angle of the sun relative to the panel. One Sun is a unit of power flux, not a standard value for actual insolation. Sometimes this unit is referred to as a Sol, not to be confused with a sol, meaning one solar day.
Absorption and reflection
Solar irradiance spectrum above atmosphere and at surface
Part of the radiation reaching an object is absorbed and
the remainder reflected. Usually, the absorbed radiation is converted to
thermal energy,
increasing the object's temperature. Humanmade or natural systems,
however, can convert part of the absorbed radiation into another form
such as electricity or chemical bonds, as in the case of photovoltaic cells or plants. The proportion of reflected radiation is the object's reflectivity or albedo.
Projection effect
Projection effect: One sunbeam one mile wide shines on the ground at a 90° angle, and another at a 30° angle. The oblique sunbeam distributes its light energy over twice as much area.
Insolation onto a surface is largest when the surface
directly faces (is normal to) the sun. As the angle between the surface
and the Sun moves from normal, the insolation is reduced in proportion
to the angle's cosine; see effect of Sun angle on climate.
In the figure, the angle shown is between the ground and
the sunbeam rather than between the vertical direction and the sunbeam;
hence the sine rather than the cosine is appropriate. A sunbeam one
mile wide arrives from directly overhead, and another at a 30° angle to
the horizontal. The sine of a 30° angle is1/2, whereas the sine of a 90° angle is1.
Therefore, the angled sunbeam spreads the light over twice the area.
Consequently, half as much light falls on each square mile.
This projection effect is the main reason why Earth's polar regions are much colder than equatorial regions.
On an annual average, the poles receive less insolation than does the
equator, because the poles are always angled more away from the Sun than
the tropics, and moreover receive no insolation at all for the six
months of their respective winters.
Absorption effect
At a lower angle, the light must also travel through more
atmosphere. This attenuates it (by absorption and scattering) further
reducing insolation at the surface.
Attenuation is governed by the Beer-Lambert Law, namely that the transmittance or fraction of insolation reaching the surface decreases exponentially in the optical depth or absorbance (the two notions differing only by a constant factor of ln(10) = 2.303)
of the path of insolation through the atmosphere. For any given short
length of the path, the optical depth is proportional to the number of
absorbers and scatterers along that length, typically increasing with
decreasing altitude. The optical depth of the whole path is then the
integral (sum) of those optical depths along the path.
When the density of absorbers is layered, that is, depends
much more on vertical than horizontal position in the atmosphere, to a
good approximation the optical depth is inversely proportional to the
projection effect, that is, to the cosine of the zenith angle. Since
transmittance decreases exponentially with increasing optical depth, as
the sun approaches the horizon there comes a point when absorption
dominates projection for the rest of the day. With a relatively high
level of absorbers this can be a considerable portion of the late
afternoon, and likewise of the early morning. Conversely, in the
(hypothetical) total absence of absorption, the optical depth remains
zero at all altitudes of the sun, that is, transmittance remains1, and so only the projection effect applies.
Solar potential maps
Assessment and mapping of solar potential at the global,
regional and country levels have been the subject of significant
academic and commercial interest. One of the earliest attempts to carry
out comprehensive mapping of solar potential for individual countries
was the Solar & Wind Resource Assessment (SWERA) project, funded by the United Nations Environment Program and carried out by the US National Renewable Energy Laboratory (NREL). The National Aeronautics and Space Administration (NASA) provides data for global solar potential maps through the CERES experiment and the POWER
project. Global mapping by many other similar institutes are available
on the Global Atlas for Renewable Energy provided by the International Renewable Energy Agency.
A number of commercial firms now exist to provide solar resource data
to solar power developers, including 3E, Clean Power Research, SoDa
Solar Radiation Data, Solargis, Vaisala (previously 3Tier), and Vortex,
and these firms have often provided solar potential maps for free. The Global Solar Atlas was launched by the World Bank in January 2017, using data provided by Solargis, to provide a single source for high-quality solar data, maps, and GIS layers covering all countries.
Maps of GHI potential by region and country (Note: colors are not consistent across maps)
Sub-Saharan Africa
Latin America and Caribbean
China
India
Mexico
South Africa
Solar radiation maps are built using databases derived
from satellite imagery, as for example using visible images from
Meteosat Prime satellite. A method is applied to the images to determine
solar radiation. One well validated satellite-to-irradiance model is
the SUNY model. The accuracy of this model is well evaluated. In general, solar
irradiance maps are accurate, especially for Global Horizontal
Irradiance.
Applications
Solar power
Sunlight carries radiant energy in the wavelengths of visible light. Radiant energy may be developed for solar power generation.
Solar irradiation figures are used to plan the deployment of solar power systems. In many countries, the figures can be obtained from an insolation map or
from insolation tables that reflect data over the prior 30–50 years.
Different solar power technologies are able to use different components
of the total irradiation. While solar photovoltaics panels are able to convert to electricity both direct irradiation and diffuse irradiation, concentrated solar power
is only able to operate efficiently with direct irradiation, thus
making these systems suitable only in locations with relatively low
cloud cover.
Because solar collectors panels are almost always mounted
at an angle towards the Sun, insolation figures must be adjusted to find
the amount of sunlight falling on the panel. This will prevent
estimates that are inaccurately low for winter and inaccurately high for
summer. This also means that the amount of sunlight falling on a solar panel at
high latitude is not as low compared to one at the equator as would
appear from just considering insolation on a horizontal surface.
Horizontal insolation values range from 800 to 950kWh/(kWp·y) in Norway to up to 2,900kWh/(kWp·y) in Australia. But a properly tilted panel at 50° latitude receives 1860kWh/m2/y, compared to 2370 at the equator. In fact, under clear skies a solar panel placed horizontally at the
north or south pole at midsummer receives more sunlight over 24 hours
(cosine of angle of incidence equal to sin(23.5°) or about 0.40) than a
horizontal panel at the equator at the equinox (average cosine equal to
1/π or about 0.32).
Photovoltaic panels are rated under standard conditions to determine the Wp (peak watts) rating, which can then be used with insolation, adjusted by factors such as
tilt, tracking and shading, to determine the expected output.
Buildings
Insolation variation by month; 1984–1993 averages for January (top) and April (bottom)
In construction, insolation is an important consideration when designing a building for a particular site.
The projection effect can be used to design buildings that
are cool in summer and warm in winter, by providing vertical windows on
the equator-facing side of the building (the south face in the Northern Hemisphere, or the north face in the Southern Hemisphere):
this maximizes insolation in the winter months when the Sun is low in
the sky and minimizes it in the summer when the Sun is high. (The Sun's north–south path through the sky spans 47° through the year).
Civil engineering
In civil engineering and hydrology, numerical models of snowmelt runoff use observations of insolation.
This permits estimation of the rate at which water is released from a melting snowpack.
Field measurement is accomplished using a pyranometer.
Climate research
Irradiance plays a part in climate modeling and weather forecasting.
A non-zero average global net radiation at the top of the atmosphere is
indicative of Earth's thermal disequilibrium as imposed by climate forcing.
The impact of the lower 2014 TSI value on climate models
is unknown. A few tenths of a percent change in the absolute TSI level
is typically considered to be of minimal consequence for climate
simulations. The new measurements require climate model parameter
adjustments.
Experiments with GISS Model 3 investigated the sensitivity
of model performance to the TSI absolute value during the present and
pre-industrial epochs, and describe, for example, how the irradiance
reduction is partitioned between the atmosphere and surface and the
effects on outgoing radiation.
Assessing the impact of long-term irradiance changes on climate requires greater instrument stability combined with reliable global surface temperature
observations to quantify climate response processes to radiative
forcing on decadal time scales. The observed 0.1% irradiance increase
imparts 0.22W/m2 climate forcing, which suggests a transient climate response of 0.6°C per W/m2.
This response is larger by a factor of 2 or more than in the
IPCC-assessed 2008 models, possibly appearing in the models' heat uptake
by the ocean.
Global cooling
Measuring a surface's capacity to reflect solar irradiance is essential to passive daytime radiative cooling, which has been proposed as a method of reversing local and global temperature increases associated with global warming. In order to measure the cooling power of a passive radiative cooling
surface, both the absorbed powers of atmospheric and solar radiations
must be quantified. On a clear day, solar irradiance can reach 1000 W/m2 with a diffuse component between 50 and 100 W/m2. On average the cooling power of a passive daytime radiative cooling surface has been estimated at ~100-150 W/m2.
A food pyramid and a corresponding food web, demonstrating some of the simpler patterns in a food webA graphic representation of energy transfer between trophic layers in an ecosystem
Energy flow is the flow of energy through living things within an ecosystem. All living organisms can be organized into producers and consumers, and those producers and consumers can further be organized into a food chain. Each of the levels within the food chain is a trophic level. In order to more efficiently show the quantity of organisms at each trophic level, these food chains are then organized into trophic pyramids. The arrows in the food chain show that the energy flow is
unidirectional, with the head of an arrow indicating the direction of
energy flow; energy is lost as heat at each step along the way.
The unidirectional flow of energy and the successive loss of energy as it travels up the food web are patterns in energy flow that are governed by thermodynamics, which is the theory of energy exchange between systems. Trophic dynamics relates to thermodynamics because it deals with the
transfer and transformation of energy (originating externally from the
sun via solar radiation) to and among organisms.
Energetics and the carbon cycle
The carbon cycle of a terrestrial ecosystem. Beginning with photosynthesis, water (blue) and carbon dioxide (white) from the air are taken in with solar energy (yellow), and are converted into plant energy (green). 100×1015 grams of carbon/year fixed by photosynthetic organisms, which is equivalent to 4×1018 kJ/yr = 4×1021 J/yr of free energy.Cellular respiration
is the reverse reaction, wherein energy of plants is taken in and
carbon dioxide and water are given off. The carbon dioxide and water
produced can be recycled back into plants.
The first step in energetics is photosynthesis, where in water and carbon dioxide from the air are taken in with energy from the sun, and are converted into oxygen and glucose. Cellular respiration is the reverse reaction, wherein oxygen and sugar
are taken in and release energy as they are converted back into carbon
dioxide and water. The carbon dioxide and water produced by respiration
can be recycled back into plants.
Energy loss can be measured either by efficiency (how much energy makes it to the next level), or by biomass (how much living material exists at those levels at one point in time, measured by standing crop). Of all the net primary productivity
at the producer trophic level, in general only 10% goes to the next
level, the primary consumers, then only 10% of that 10% goes on to the
next trophic level, and so on up the food pyramid. Ecological efficiency may be anywhere from 5% to 20% depending on how efficient or inefficient that ecosystem is. This decrease in efficiency occurs because organisms need to perform
cellular respiration to survive, and energy is lost as heat when
cellular respiration is performed. That is also why there are fewer tertiary consumers than there are producers.
A producer is any organism that performs photosynthesis. Producers are important because they convert energy from the sun into a storable and usable chemical form of energy, glucose, as well as oxygen. The producers themselves can use the energy stored
in glucose to perform cellular respiration. Or, if the producer is
consumed by herbivores in the next trophic level, some of the energy is passed on up the pyramid. The glucose stored within producers serves as food for consumers, and so it is only through producers that consumers are able to access the sun's energy.Some examples of primary producers are algae, mosses, and other plants such as grasses, trees, and shrubs.
Chemosynthetic
bacteria perform a process similar to photosynthesis, but instead of
energy from the sun they use energy stored in chemicals like hydrogen sulfide. This process, referred to as chemosynthesis,
usually occurs deep in the ocean at hydrothermal vents that produce
heat and chemicals such as hydrogen, hydrogen sulfide and methane. Chemosynthetic bacteria can use the energy in the bonds of the hydrogen
sulfide and oxygen to convert carbon dioxide to glucose, releasing
water and sulfur in the process. Organisms that consume the chemosynthetic bacteria can take in the
glucose and use oxygen to perform cellular respiration, similar to
herbivores consuming producers.
One of the factors that controls primary production is the
amount of energy that enters the producer(s), which can be measured
using productivity. Only one percent of solar energy enters the producer, the rest bounces off or moves through. Gross primary productivity is the amount of energy the producer actually gets. Generally, 60% of the energy that enters the producer goes to the producer's own respiration. The net primary productivity is the amount that the plant retains after
the amount that it used for cellular respiration is subtracted. Another factor controlling primary production is organic/inorganic
nutrient levels in the water or soil that the producer is living in. An example of the nutrients that can impact the efficiency of primary plant production are nitrogen (N) and phosphorus (P).
Venus Flytrap
Carnivorous plants
When it comes to dealing with environments that have low
nutrient availability, some plants have developed unique ways to adapt
to be able to perform photosynthesis. In order to do so, these plants
have evolved to be able to obtain important nutrients such as nitrogen
from other organisms, just as heterotrophs would giving them the unique
title of carnivorous plants.With methods such as the pitfall trap (pitcher plant), the flypaper trap (Drosera capensis), or the snap trap (venus flytrap) these plants have learned to lure insects in and digest them.
Pitcher plants
lure insects in using a variety of attractive cues such as scent and
color. Once an insect or small organism falls into the bulb shaped body
of the plant, a variety of enzymes are secreted beginning the digestion
process of the organism and preventing it from escaping. Flypaper trap plants,
the most common of carnivorous plants, secret a special liquid that
allow an insect to land on its leaves but then prevents the insect from
escaping. The snap trap plant,
use similar methods to the pitcher plant in order to attract various
insects. However, these carnivorous plants are able to detect when an
insect is touching its leaves thus triggering the "mouth" of the plant
to close and encase the insect. In developing this method of nutrient acquisition, carnivorous plants
are able to survive in almost any environment around the world,
excluding Antarctica and the Arctic Circle.
Secondary production
Secondary production is the use of energy stored in plants
converted by consumers to their own biomass. Different ecosystems have
different levels of consumers, all end with one top consumer. Most
energy is stored in organic matter of plants, and as the consumers eat
these plants they take up this energy. This energy in the herbivores and
omnivores is then consumed by carnivores.
There is also a large amount of energy that is in primary production
and ends up being waste or litter, referred to as detritus. The detrital
food chain includes a large amount of microbes, macroinvertebrates, meiofauna,
fungi, and bacteria. These organisms are consumed by omnivores and
carnivores and account for a large amount of secondary production. Secondary consumers can vary widely in how efficient they are in consuming. The efficiency of energy being passed on to consumers is estimated to be around 10%. Energy flow through consumers differs in aquatic and terrestrial environments.
In aquatic environments
Heterotrophs contribute to secondary production and it is dependent on primary productivity and the net primary products. Secondary production is the energy that herbivores and decomposers use and thus depends on primary productivity. Primarily herbivores and decomposers consume all the carbon from two main organic sources in aquatic ecosystems, autochthonous and allochthonous. Autochthonous carbon comes from within the ecosystem and includes
aquatic plants, algae and phytoplankton. Allochthonous carbon from
outside the ecosystem is mostly dead organic matter from the terrestrial
ecosystem entering the water. In stream ecosystems, approximately 66% of annual energy input can be
washed downstream. The remaining amount is consumed and lost as heat.
In terrestrial environments
Secondary production is often described in terms of trophic levels,
and while this can be useful in explaining relationships it
overemphasizes the rarer interactions. Consumers often feed at multiple
trophic levels. Energy transferred above the third trophic level is relatively unimportant. The assimilation efficiency can be expressed by the amount of food the
consumer has eaten, how much the consumer assimilates and what is
expelled as feces or urine. While a portion of the energy is used for respiration, another portion of the energy goes towards biomass in the consumer. There are two major food chains: The primary food chain is the energy
coming from autotrophs and passed on to the consumers; and the second
major food chain is when carnivores eat the herbivores or decomposers
that consume the autotrophic energy. Consumers are broken down into primary consumers, secondary consumers
and tertiary consumers. Carnivores have a much higher assimilation of
energy, about 80% and herbivores have a much lower efficiency of
approximately 20 to 50%. Energy in a system can be affected by animal emigration/immigration.
The movements of organisms are significant in terrestrial ecosystems. Energetic consumption by herbivores in terrestrial ecosystems has a low range of ~3-7%. The flow of energy is similar in many terrestrial environments. The
fluctuation in the amount of net primary product consumed by herbivores
is generally low. This is in large contrast to aquatic environments of
lakes and ponds where grazers have a much higher consumption of around
~33%. Ectotherms and endotherms have very different assimilation efficiencies.
Detritivores
Detritivores consume organic material that is decomposing and are in turn consumed by carnivores. Predator
productivity is correlated with prey productivity. This confirms that
the primary productivity in ecosystems affects all productivity
following.
Detritus
is a large portion of organic material in ecosystems. Organic material
in temperate forests is mostly made up of dead plants, approximately
62%.
In an aquatic ecosystem, leaf matter that falls into
streams gets wet and begins to leech organic material. This happens
rather quickly and will attract microbes and invertebrates. The leaves
can be broken down into large pieces called coarse particulate organic matter (CPOM). The CPOM is rapidly colonized by microbes. Meiofauna is extremely important to secondary production in stream ecosystems. Microbes breaking down and colonizing this leaf matter are very
important to the detritovores. The detritovores make the leaf matter
more edible by releasing compounds from the tissues; it ultimately helps
soften them. As leaves decay nitrogen will decrease since cellulose and lignin
in the leaves is difficult to break down. Thus the colonizing microbes
bring in nitrogen in order to aid in the decomposition. Leaf breakdown
can depend on initial nitrogen content, season, and species of trees.
The species of trees can have variation when their leaves fall. Thus the
breakdown of leaves is happening at different times, which is called a
mosaic of microbial populations.
Species effect and diversity in an ecosystem can be analyzed through their performance and efficiency. In addition, secondary production in streams can be influenced heavily
by detritus that falls into the streams; production of benthic fauna
biomass and abundance decreased an additional 47–50% during a study of
litter removal and exclusion.
Energy flow across ecosystems
Research has demonstrated that primary producers fix carbon at similar rates across ecosystems. Once carbon has been introduced into a system as a viable source of
energy, the mechanisms that govern the flow of energy to higher trophic
levels vary across ecosystems. Among aquatic and terrestrial ecosystems,
patterns have been identified that can account for this variation and
have been divided into two main pathways of control: top-down and
bottom-up. The acting mechanisms within each pathway ultimately regulate community
and trophic level structure within an ecosystem to varying degrees. Bottom-up controls involve mechanisms that are based on resource
quality and availability, which control primary productivity and the
subsequent flow of energy and biomass to higher trophic levels. Top-down controls involve mechanisms that are based on consumption by consumers. These mechanisms control the rate of energy transfer from one trophic
level to another as herbivores or predators feed on lower trophic
levels.
Aquatic vs terrestrial ecosystems
Much variation in the flow of energy is found within each
type of ecosystem, creating a challenge in identifying variation between
ecosystem types. In a general sense, the flow of energy is a function
of primary productivity with temperature, water availability, and light availability. For example, among aquatic ecosystems, higher rates of production are
usually found in large rivers and shallow lakes than in deep lakes and
clear headwater streams. Among terrestrial ecosystems, marshes, swamps, and tropical rainforests have the highest primary production rates, whereas tundra and alpine ecosystems have the lowest. The relationships between primary production and environmental
conditions have helped account for variation within ecosystem types,
allowing ecologists to demonstrate that energy flows more efficiently
through aquatic ecosystems than terrestrial ecosystems due to the
various bottom-up and top-down controls in play.
Bottom-up
The strength of bottom-up controls on energy flow are determined by the nutritional quality, size, and growth rates of primary producers in an ecosystem. Photosynthetic material is typically rich in nitrogen (N) and phosphorus (P) and supplements the high herbivore demand for N and P across all ecosystems. Aquatic primary production is dominated by small, single-celled phytoplankton that are mostly composed of photosynthetic material, providing an efficient source of these nutrients for herbivores. In contrast, multi-cellular terrestrial plants contain many large supporting cellulose structures of high carbon but low nutrient value. Because of this structural difference, aquatic primary producers have
less biomass per photosynthetic tissue stored within the aquatic
ecosystem than in the forests and grasslands of terrestrial ecosystems. This low biomass relative to photosynthetic material in aquatic
ecosystems allows for a more efficient turnover rate compared to
terrestrial ecosystems. As phytoplankton are consumed by herbivores, their enhanced growth and
reproduction rates sufficiently replace lost biomass and, in conjunction
with their nutrient dense quality, support greater secondary
production.
Additional factors impacting primary production includes
inputs of N and P, which occurs at a greater magnitude in aquatic
ecosystems. These nutrients are important in stimulating plant growth and, when passed to higher trophic levels, stimulate consumer biomass and growth rate. If either of these nutrients are in short supply, they can limit overall primary production. Within lakes, P tends to be the greater limiting nutrient while both N and P limit primary production in rivers. Due to these limiting effects, nutrient inputs can potentially
alleviate the limitations on net primary production of an aquatic
ecosystem. Allochthonous material washed into an aquatic ecosystem introduces N
and P as well as energy in the form of carbon molecules that are readily
taken up by primary producers. Greater inputs and increased nutrient concentrations support greater
net primary production rates, which in turn supports greater secondary
production.
Top-down
Top-down mechanisms exert greater control on aquatic primary producers due to the roll of consumers within an aquatic food web. Among consumers, herbivores can mediate the impacts of trophic cascades
by bridging the flow of energy from primary producers to predators in
higher trophic levels. Across ecosystems, there is a consistent association between herbivore growth and producer nutritional quality. However, in aquatic ecosystems, primary producers are consumed by
herbivores at a rate four times greater than in terrestrial ecosystems. Although this topic is highly debated, researchers have attributed the
distinction in herbivore control to several theories, including producer
to consumer size ratios and herbivore selectivity.
A
freshwater food web demonstrating the size differences between each
trophic level. Primary producers tend to be small algal cells.
Herbivores tend to be small macro-invertebrates. Predators tend to be
larger fish.
Modeling
of top-down controls on primary producers suggests that the greatest
control on the flow of energy occurs when the size ratio of consumer to
primary producer is the highest. The size distribution of organisms found within a single trophic level
in aquatic systems is much narrower than that of terrestrial systems. On land, the consumer size ranges from smaller than the plant it
consumes, such as an insect, to significantly larger, such as an ungulate,
while in aquatic systems, consumer body size within a trophic level
varies much less and is strongly correlated with trophic position. As a result, the size difference between producers and consumers is
consistently larger in aquatic environments than on land, resulting in
stronger herbivore control over aquatic primary producers.
Herbivores can potentially control the fate of organic matter as it is cycled through the food web. Herbivores tend to select nutritious plants while avoiding plants with structural defense mechanisms. Like support structures, defense structures are composed of nutrient poor, high carbon cellulose. Access to nutritious food sources enhances herbivore metabolism and
energy demands, leading to greater removal of primary producers. In aquatic ecosystems, phytoplankton are highly nutritious and generally lack defense mechanisms. This results in greater top-down control because consumed plant matter is quickly released back into the system as labile organic waste. In terrestrial ecosystems, primary producers are less nutritionally dense and are more likely to contain defense structures. Because herbivores prefer nutritionally dense plants and avoid plants
or plant parts with defense structures, a greater amount of plant matter
is left unconsumed within the ecosystem. Herbivore avoidance of low-quality plant matter may be why terrestrial
systems exhibit weaker top-down control on the flow of energy.